Coffee Creek, 34 (Arroyo Formation): Kungurian, Texas
collected by J. W. Kitching, P. C. Miller, E. C. Olson, C. H. Sternberg, S. W. Williston, among others 1895, 1897, 1909, 1965, 1972

List of taxa
Where & when
Taphonomy & methods
Metadata & references
Taxonomic list
Zatrachys conchigerus n. sp. Cope 1896
Romer 1928
nomen dubium belonging to Temnospondyli
Authority is not Romer 1928.
Eryops sp. Cope 1877
Romer 1928 14 specimens
Anisodexis imbricarius n. gen., n. sp. Cope 1882
1 individual
synonym of Eryops megacephalus
Anisodexis sp. Cope 1882
Romer 1928 1 specimen
cf. Zatrachys sp. Cope 1878
Schoch and Milner 2014
SMU specimens (Murray & Johnson, 1987) from East Coffee Creek
Zatrachys microphthalmus n. sp. Cope 1869
Romer 1928
recombined as Dasyceps microphthalmus
Authority is not Romer 1928. 7 Zatryachys specimens total, see below Z. conchigerus.
Dissorophus multicinctus Cope 1895
Cope 1895 3 specimens
Alegeinosaurus aphthitos n. gen., n. sp. Case 1911
Schoch and Milner 2014 1 individual
recombined as Aspidosaurus aphthitos
AMNH 4756 (holotype)
Aspidosaurus chiton n. gen., n. sp. Broili 1904
Romer 1928 2 specimens
Authority is not Romer 1928.
Acheloma sp. Cope 1882
Romer 1928 1 specimen
Trimerorhachis mesops n. sp. Cope 1896
Romer 1928 8 specimens
Authority is not Romer 1928. entered as "Trimerorachis"
Isodectes obtusus (Cope 1868)
Schoch and Milner 2014
AMNH 4744 (Originally A. ? casei figured by Broili 1913, pl. 9, figs 1-2, presumed destroyed in 1944)
? Acheloma casei n. sp. Broili 1913
Broili 1913 1 individual
synonym of Isodectes obtusus
BSP unknown catalogue number; occurence is deducted from the original 2 specimens of Acheloma sp. reported in the primary reference
Brachydectes elongatus Wellstead 1991
Wellstead 1991
AMNH 4762a and 7558
Reptilia - Captorhinidae
Pariotichus hamatus n. sp. Cope 1896
Seltin 1959 1 individual
recombined as Labidosaurus hamatus
AMNH 4341 (type)
Labidosaurus hamatus (Cope 1896)
Romer 1928 32 specimens
AMNH 4406, 4414, 4417, 4421, 4427; CNHM ?*, CNHM UR 161, 273, UC ?*, UC 180, 634*, 643, 724-726, 728-730, 732, 1059, 1199, P 12758; CM 73370 (formerly UCLA VP 3167), CM 73371 (formerly UCLA VP 3200), CM 76876; UCLA VP 435, 436, 3115, 3491; USNM 17045; MCZ 8923; catalogue numbers are from Seltin (1959); specimens not reported by Seltin are compiled from Sumida (1987, 1989) and Modesto (2007); asterisks mark specimens of which the locality is not precisely known but come from the Arroyo Fm. of Baylor Co.
Labidosaurus broilii n. sp. Case 1911
Case 1911 1 individual
synonym of Labidosaurus hamatus
Ectocynodon aguti n. sp. Cope 1882
1 individual
recombined as Captorhinus aguti
AMNH 4333 (type)
Hypopnous squaliceps n. gen., n. sp. Cope 1896
Romer 1928 2 individuals
synonym of Captorhinus aguti
AMNH 4335 (type); referred specimen of Captorhinus aguti in Seltin (1959); = "indeterminate romeriid" of Fox & Bowman (1966)
Captorhinus angusticeps n. gen., n. sp. Cope 1896
Cope 1896 37 specimens
synonym of Captorhinus aguti
AMNH 4438 (type); 4334, 4340, 4407, 4408, 4410A, 4438, 4443-4445, 4455*, 4457, 4458, 4760, 6790, 6791; CNHM UC 651, 700, 1043, 1702, 1703, UR 274-276, 292, 327, 351, 371, 372, 1206*; MCZ 1059; USNM 17084*; catalogue numbers are from Seltin (1959) and from Fox & Bowman (1966); all are referred specimens of C. aguti in these references and former assignment to C. angusticeps is not confirmed; asterisks mark specimens of which the locality is unknown but come from the Arroyo Fm. of Baylor Co.; at least 5 specimens unnumbered or missing
Pariotichus isolomus n. sp. Cope 1896
Cope 1896 1 individual
synonym of Captorhinus aguti
AMNH 4338 (type)
Tetraceratops insignis n. gen., n. sp. Matthew 1908
Matthew 1908 1 individual
AMNH 4526 (type)
Varanosaurus acutirostris Broili 1904
Romer and Price 1940 1 individual
FMNH (WM) 34
Poecilospondylus francisi n. gen., n. sp. Case 1910
Case 1910 1 individual
synonym of Varanosaurus acutirostris
AMNH 4174 (type)
Edaphosaurus pogonias n. gen., n. sp. Cope 1882
1 individual
AMNH 4009 (type)
Naosaurus claviger n. gen., n. sp. Cope 1886
Cope 1886 1 individual
synonym of Edaphosaurus pogonias
AMNH 4002 (type)
Embolophorus dollovianus n. sp. Cope 1888
Cope 1888 1 individual
recombined as Dimetrodon dollovianus
AMNH 4064 (type)
Dimetrodon loomisi Romer 1937
Reisz 1986 13 specimens
FMNH 114 (formerly referred to D. dollovianus) + the rest of the total of 15 Dimetrodon specimens originally reported by Romer (1928)
Dimetrodon gigashomogenes n. sp. Case 1907
Case 1907 1 individual
FMNH 112 (type)
Bolbodon tenuitectes n. gen., n. sp. Cope 1896
Cope 1896 1 individual
recombined as Diadectes tenuitectes
AMNH 4375 (type)
Diadectes huenei n. sp. Broom 1914
Romer 1928 24 specimens
synonym of Diadectes tenuitectes
Authority is not Romer 1928.
Seymouria baylorensis n. gen., n. sp. Broili 1904
Broili 1904 2 individuals
AMNH 4342 (type); MCZ 1081, 1083, 1084, 1086; FMNH UR 458; specimen numbers are from Laurin (1996); MCZ 1081, 1083, 1084, 1086 are not counted here because they appear to be already included in collection 22715
Desmospondylus anomalus n. gen., n. sp. Williston 1910
Williston 1910 1 individual
synonym of Seymouria baylorensis
FMNH UC 664, 665 (both constitute the holotype)
Cardiocephalus sternbergi n. gen., n. sp. Broili 1904
Broili 1904 2 individuals
Gymnarthrus willoughbyi n. gen., n. sp. Case 1910
Case 1910 1 specimen
synonym of Cardiocephalus sternbergi
AMNH 4892 (type)
Pariotichus brachyops Cope 1878
Romer 1928 1 individual
AMNH 4760 ("homotype"); type skull of "Captorhinus gregoryi" Broom (see Gregory et al., 1956)
Pelodosotis elongatum n. gen., n. sp. Carroll and Gaskill 1978
Carroll and Gaskill 1978 1 individual
UMMP 11156 (type), formerly assigned to Ostodolepis brevispinatus by Case (1929)
Ostodolepis brevispinatus n. gen., n. sp. Williston 1913
Williston 1913 1 individual
FMNH UR 860 (type)
? Micraroter erythrogeios Daly 1973
Carroll and Gaskill 1978 1 individual
Quasicaecilia texana n. gen., n. sp. Carroll 1990
Carroll 1990
Exact location where this specimen was found is unknown. Museum data indicates it was collected by C. H. Sternberg in 1917 from "?Baylor County, Texas; Lower Permian, ?Arroyo Formation". USNM 22079 is the holotype (an isolated skull)
Diplocaulus sp. Cope 1877
Olson 1951 44 specimens
Romer (1928) reports 72 Diplocaulus specimens total; since Olson (1951) refers only 28 specimens from this locality to particular species the remainder (no catalogue numbers given) are listed here as Diplocaulus sp.
Diplocaulus magnicornis n. sp. Cope 1882
23 specimens
AMNH 4466, 4467, 4469, 4484, 4491, 4494, 4498, 4504, 4511*, 4512*, 4514, 4539 (type); CNHM UC 229, 636, 637, 1650, 1654, 1656-1658, 1660, 1663, P 12689; catalogue numbers are from Olson (1951), asterisks mark specimens with uncertain locality data
Diplocaulus limbatus n. sp. Cope 1896
Cope 1896 1 individual
synonym of Diplocaulus magnicornis
AMNH 4471 (type); because D. limbatus was found to be an invalid species by Olson (1951) only the type specimen is listed here
Diplocaulus brevirostris n. sp. Olson 1951
Olson 1951 4 individuals
AMNH 4470 (type), 4544; CNHM UC 1655, 1661
Lysorophus sp. Cope 1877
Romer 1928 54 specimens
Pantylus coicodus n. sp. Cope 1896
Romer 1928 4 specimens
Authority is not Romer 1928.
see common names

Country:United States State/province:Texas County:Baylor/Willbarger
Coordinates: 33.8° North, 99.2° West (view map)
Paleocoordinates:2.3° North, 28.7° West
Basis of coordinate:based on nearby landmark
Geographic resolution:local area
Period:Permian Epoch:Cisuralian
Stage:Kungurian 10 m.y. bin:Permian 2
*Period:Early/Lower Permian - Early/Lower Permian *Epoch:Rotliegendes
Key time interval:Kungurian
Age range of interval:279.30000 - 272.30000 m.y. ago
* legacy (obsolete) database fields
Geological group:Clear Fork Formation:Arroyo
Stratigraphic resolution:group of beds
Stratigraphy comments: "probably not much over 100 feet above the Lueders limestone; the lowest portion may have lain below it.." Details for Faunal Zone 4 of Romer 1928: "The vertebrate remains are confined to approximately the lowest 250 feet or so of the Clear Fork.."
the base and top of the Clear Fork Group (i.e., Arroyo, Vale, and Choza formations) are both within the Kungurian: see Wardlaw 2005 (Permophiles) for the base, and DiMichele et al. 2001 (J Paleont) for the top
Lithology and environment
Primary lithology:red claystone
Secondary lithology: conglomeratic sandstone
Includes fossils?Y
Lithology description: "red beds... red clays, with small amounts of sandstone and conglomerates."
Environment:terrestrial indet.
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:field collection,survey of museum collection
Reason for describing collection:taxonomic analysis
Museum repositories:AMNH,FMNH,MCZ,USNM
Collectors:J. W. Kitching, P. C. Miller, E. C. Olson, C. H. Sternberg, S. W. Williston, among others Collection dates:1895, 1897, 1909, 1965, 1972
Collection method comments: Romer (1928) paper is a review of vertebrates from the Texas red-beds with previously unreported (or undetailed) localities.
Additional institutions storing material from Coffee Creek:
BP (Bernard Price Institute, University of the Witwatersrand, Johannesburg, South Africa)
BSP (Bayerische Staatssammlung für Paläontologie und Geologie, Munich, Germany)
UMMP (University of Michigan Museum of Paleontology, Ann Arbor, MI)
Also known as:Locality 34 (Romer 1928), faunal Zone 4, Lower Clear Fork Group, Texas red-beds, redbeds; Locality VIIa (Romer & Price 1940); "Labidosaurus pocket"; Coffee Creek Bonebed; West Coffee Creek; Middle Coffee Creek; East Coffee Creek
Database number:28256
Authorizer:J. Alroy, R. Butler, J. Mueller, M. Carrano Enterer:R. Whatley, R. Butler, T. Liebrecht, E. Dunne, M. Carrano
Modifier:E. Dunne Research group:vertebrate
Collections that are a subset of this one:13046
Created:2003-02-03 14:30:51 Last modified:2017-04-07 05:55:42
Access level:the public Released:2003-02-03 14:30:51
Creative Commons license:CC BY
Reference information

Primary reference:

17702.ETE E. D. Cope. 1882. Third contribution to the history of the Vertebrata of the Permian formation of Texas. Proceedings of the American Philosophical Society 20:447-461 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

28751 F. Broili. 1904. Permische Stegocephalen und Reptilien aus Texas. Palaeontographica 51:1-120 [J. Mueller/T. Liebrecht/T. Liebrecht]
31846 F. Broili. 1913. Über zwei Stegocephalenreste aus dem texanischen Perm. Neues Jahrbuch für Mineralogie, Geologie und Paläontologie 1913(1):96-100 [J. Mueller/T. Liebrecht]
58525 R. L. Carroll. 1990. A tiny microsaur from the lower Permian of Texas: size constraints in Palaeozoic tetrapods. Palaeontology 33:893-909 [R. Butler/E. Dunne]
28692 R. L. Carroll and P. Gaskill. 1978. The Order Microsauria. Memoirs of the American Philosophical Society 126:1-211 [J. Mueller/T. Liebrecht/T. Liebrecht]
73745% 52340E. C. Case. 1907. Revision of the Pelycosauria of North America. Carnegie Institution of Washington 55:3-176 [C. Sidor/C. Sidor/M. Carrano]
28818 E. C. Case. 1910. The skeleton of Poecilospondylus francisi, a new genus and species of Pelycosauria. Bulletin of the American Museum of Natural History 23:183-188 [J. Mueller/T. Liebrecht/M. Carrano]
28729 E. C. Case. 1910. New or little known reptiles and amphibians from the Permian (?) of Texas. Bulletin of the American Museum of Natural History 28:163-181 [J. Mueller/T. Liebrecht/M. Carrano]
28341 E. C. Case. 1911. A revision of the Cotylosauria of North America. Carnegie Institution of Washington Publication 145:1-121 [J. Mueller/T. Liebrecht/T. Liebrecht]
30432 E. C. Case. 1929. Description of a Nearly Complete Skeleton of Ostodolepis brevispinatus Williston. Contributions from the Museum of Paleontology, University of Michigan 3(5):81-107 [J. Mueller/T. Liebrecht/J. Alroy]
28380 E. D. Cope. 1886. The long-spined Theromorpha of the Permian Epoch. The American Naturalist 20(6):544-545 [J. Mueller/T. Liebrecht/M. Carrano]
28371 E. D. Cope. 1888. Systematic catalogue of the species of Vertebrata found in the beds of the Permian Epoch in North America with notes and descriptions. Transactions of the Philosophical Society of America, New Series 16(2):285-297 [J. Mueller/T. Liebrecht/M. Carrano]
54069 E. D. Cope. 1895. A batrachian armadillo. The American Naturalist 29:998 [M. Carrano/M. Carrano]
28342 E. D. Cope. 1896. The reptilian order Cotylosauria. Proceedings of the American Philosophical Society 34:436-457 [J. Mueller/T. Liebrecht/M. Carrano]
28358 E. D. Cope. 1896. Second contribution to the history of the Cotylosauria. Proceedings of the American Philosophical Society 35(151):122-139 [J. Mueller/T. Liebrecht/T. Liebrecht]
28784 R. C. Fox and M. C. Bowman. 1966. Osteology and Relationships of Captorhinus aguti (Cope) (Reptilia: Captorhinomorpha). The University of Kansas Paleontological Contributions, Series Vertebrata 11:1-79 [J. Mueller/T. Liebrecht/J. Alroy]
28484 J. T. Gregory, F. E. Peabody, and L. I. Price. 1956. Revision of the Gymnarthridae - American Permian Microsaurs. Bulletin of the Peabody Museum of Natural History 10:1-77 [J. Mueller/T. Liebrecht]
28352ETE E. G. Kordikova, P. D. Polly, V. A. Alifanov, Z. Rocek, G. F. Gunnell and A. O. Averianov. 2001. Small vertebrates from the Late Cretaceous and Early Tertiary of the northeastern Aral Sea region, Kazakhstan. Journal of Paleontology 75(2):390-400 [M. Carrano/M. Carrano]
28754 M. Laurin. 1996. A Redescription of the Cranial Anatomy of Seymouria baylorensis, the best known Seymouriamorph (Vertebrata: Seymouriamorpha). PaleoBios 17(17):1-16 [J. Mueller/T. Liebrecht]
29679 W. D. Matthew. 1908. A four-horned pelycosaurian from the Permian of Texas. Bulletin of the American Museum of Natural History 24:183-185 [J. Mueller/T. Liebrecht]
28357 S. P. Modesto, D. M. Scott, D. S. Berman, J. Müller, and R. R. Reisz. 2007. The skull and the paleoecological significance of Labidosaurus hamatus, a captorhinid reptile from the Lower Permian of Texas. Zoological Journal of the Linnean Society 149(2):237-262 [J. Mueller/T. Liebrecht]
29284 E. C. Olson. 1951. Diplocaulus, a Study in Growth and Variation. Fieldiana: Geology 11(2):57-154 [J. Mueller/T. Liebrecht]
27105 R. R. Reisz. 1986. Pelycosauria. Handbuch der Paläoherpetologie / Encyclopedia of Paleoherpetology 17A:1-102 [J. Mueller/J. Dummasch/T. Liebrecht]
76935% 49780A. S. Romer. 1928. Vertebrate faunal horizons in the Texas Permo-Carboniferous red beds. University of Texas Bulletin 2801:67-108 [J. Alroy/R. Whatley/W. Clyde]
5692 A. S. Romer and L. I. Price. 1940. Review of the Pelycosauria. Geological Society of America Special Paper 28:1-538 [A. Behrensmeyer/A. Behrensmeyer/T. Liebrecht]
57795 R. R. Schoch and A. R. Milner. 2014. Handbook of Paleoherpetology Part 3A2 Temnospondyli I. [R. Butler/E. Dunne/E. Dunne]
28359 R. J. Seltin. 1959. A review of the family Captorhinidae. Fieldiana: Geology 10(34):461-509 [J. Mueller/T. Liebrecht/T. Liebrecht]
28824 S. S. Sumida. 1987. Two Different Vertebral Forms in the Axial Column of Labidosaurus (Captorhinomorpha: Captorhinidae). Journal of Paleontology 61(1):155-167 [J. Mueller/T. Liebrecht]
28817 S. S. Sumida. 1989. The Appendicular Skeleton of the Early Permian Genus Labidosaurus (Reptilia, Captorhinomorpha, Captorhinidae) and the Hind Limb Musculature of Captorhinid Reptiles. Journal of Vertebrate Paleontology 9(3):295-313 [J. Mueller/T. Liebrecht]
58492 C. F. Wellstead. 1991. Taxonomic revision of the Lysorophia, Permo-Carboniferous lepospondyl amphibians. Bulletin of the American Museum of Natural History 209:1-90 [R. Butler/E. Dunne]
9272 S. W. Williston. 1910. Cacops, Desmospondylus; new genera of Permian vertebrates. Geological Society America Bulletin 21:249-284 [J. Alroy/R. Whatley/M. Carrano]
30431 S. W. Williston. 1913. Ostodolepis brevispinatus, a new reptile from the Permian of Texas. Journal of Geology 21(4):363-366 [J. Mueller/T. Liebrecht/M. Carrano]