Ukhaa Tolgod (AMNH): Late/Upper Campanian, Mongolia
collected 1992-1996
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia
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Estesia mongoliensis
Norell et al. 1992
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1 individual | |||||||||
IGM 3/169 (a skull with mandible, preserved three-dimensionally) and IGM 3/760 (partial axial skeletons including dorsal vertebrae with articulating ribs and an incomplete left forelimb) | ||||||||||
Gobiderma pulchrum
(Borsuk-Bialynicka 1984)
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Gao and Norell 2000 | 3 individuals | ||||||||
misspelling Gobiderma pulchra | ||||||||||
IGM 3/56-3/58 | ||||||||||
Varanoidea indet.
Gray 1827
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Gao and Norell 2000 | 1 individual | ||||||||
IGM 3/172 | ||||||||||
Reptilia
- Slavoiidae
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Slavoia darevskii
Sulimski 1984
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Gao and Norell 2000 | 9 individuals | ||||||||
IGM 3/145-3/153 | ||||||||||
Reptilia
- Polyglyphanodontidae
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Gobinatus arenosus
Alifanov 1993
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Gao and Norell 2000 | 1 individual | ||||||||
IGM 3/128 | ||||||||||
Adamisaurus magnidentatus
Sulimski 1972
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5 individuals | |||||||||
IGM 3/111-3/115 | ||||||||||
Macrocephalosaurus sp.
Gilmore 1943
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Gao and Norell 2000 | 2 individuals | ||||||||
replaced by Gilmoreteius | ||||||||||
IGM 3/132-3/133 | ||||||||||
Reptilia
- Carusiidae
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Shinisauroides latipalatum
Borsuk-Bialynicka 1985
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Reptilia
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Isodontosaurus gracilis
Gilmore 1943
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Gao and Norell 2000 | 4 individuals | ||||||||
IGM 3/85, 86, 88, 89 | ||||||||||
Saichangurvel davidsoni n. gen., n. sp.
Conrad and Norell 2007
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Conrad and Norell 2007 | 1 individual | ||||||||
IGM 3/858 (holotype skeleton) | ||||||||||
Temujinia ellisoni n. gen., n. sp.
Gao and Norell 2000
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Gao and Norell 2000 | 7 specimens | ||||||||
IGM 3/63-3/69 | ||||||||||
Reptilia
- Priscagamidae
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Mimeosaurus crassus
Gilmore 1943
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Gao and Norell 2000 | 2 individuals | ||||||||
IGM 3/74-3/75 | ||||||||||
Priscagama gobiensis
Borsuk-Bialynicka and Moody 1984
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2 individuals | |||||||||
IGM 3/77-3/78 | ||||||||||
? Priscagama gobiensis
Borsuk-Bialynicka and Moody 1984
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Gao and Norell 2000 | 1 individual | ||||||||
IGM 3/80 | ||||||||||
Phrynosomimus asper
Alifanov 1996
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Gao and Norell 2000 | 1 individual | ||||||||
IGM 3/83 | ||||||||||
Reptilia
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Hymenosaurus clarki n. gen., n. sp.
Gao and Norell 2000
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Gao and Norell 2000 | 1 individual | ||||||||
Myrmecodaptria microphagosa n. gen., n. sp.
Gao and Norell 2000
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Gao and Norell 2000 | 1 individual | ||||||||
Parmeosaurus scutatus n. gen., n. sp.
Gao and Norell 2000
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Gao and Norell 2000 | 3 individuals | ||||||||
IGM 3/138, 141, 143 | ||||||||||
Carusia intermedia
(Borsuk-Bialynicka 1985)
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Gao and Norell 1998 | 35 individuals | ||||||||
IGM 3/16-3/50 | ||||||||||
Globaura venusta
Borsuk-Bialynicka 1988
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Gao and Norell 2000 | 4 individuals | ||||||||
IGM 3/160-3/163 | ||||||||||
Reptilia
- Avetheropoda
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Carnosauria indet.
Huene 1920
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Reptilia
- Dromaeosauridae
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cf. Velociraptor sp.
Osborn 1924
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"neonate skulls" | ||||||||||
Velociraptor mongoliensis
Osborn 1924
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Reptilia
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cf. Gobipteryx sp.
Elzanowski 1974
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= Gobipteryx minuta
Elzanowski 1974
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Chiappe et al. 2001 | |||||||||
based on IGM-100/1011, collected in 1994 from "Ukhaa Tolgod" (no details given) | ||||||||||
Reptilia
- Oviraptoridae
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Oviraptoridae indet.
Barsbold 1976
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"taxon B" | ||||||||||
Reptilia
- Alvarezsauridae
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Mononykus olecranus
(Perle et al. 1993)
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1 specimen | |||||||||
IGM 100/975 (1 measurement) | ||||||||||
= Shuvuuia deserti n. gen., n. sp.
Chiappe et al. 1998
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Chiappe et al. 1998 | |||||||||
Reptilia
- Protoceratopsidae
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Bagaceratops sp.
Maryanska and Osmólska 1975
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Reptilia
- Ankylosauridae
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Ankylosauridae indet.
Brown 1908
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Reptilia
- Paralligatoridae
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Shamosuchus djadochtaensis
Mook 1924
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Pol et al. 2009 | |||||||||
IGM 100/1195 (nearly complete skull, lower jaws, and associated postcranial remains including; all eight cervical vertebrae with three right cervical ribs (c5–c7) and one left cervical rib (c8), dorsal vertebrae d1 through d3 with four right dorsal ribs (d1–d3) and three left dorsal ribs (d2?–d6?), five anterior caudal vertebrae and two haemal arches, a complete right humerus, the distal half of the left humerus, the proximal portion of the right radius, a complete left radius in association with the left humerus, a complete left ulna, a fragmentary right ischium, a partial right femur consisting of a damaged diaphysis and distal condyle, a complete right tibia, a nearly complete left tibia, a complete right fibula, a partial left fibula, and a single left metatarsal and phalanx) | ||||||||||
Reptilia
- Testudines
- Nanhsiungchelyidae
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? Basilemys sp.
Hay 1902
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Mammalia
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Theria indet.
Parker and Haswell 1897
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"new genus D" | ||||||||||
Mammalia
- Zalambdalestidae
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Zalambdalestes lechei
Gregory and Simpson 1926
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Mammalia
- Proteutheria
- Palaeoryctidae
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cf. Hyotheridium sp.
Gregory and Simpson 1926
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Mammalia
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Kennalestes cf. gobiensis
Kielan-Jaworowska 1968
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cf. Asioryctes sp.
Kielan-Jaworowska 1975
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= Ukhaatherium nessovi n. gen., n. sp.
Novacek et al. 1997
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Novacek et al. 1997 | |||||||||
PSS-MAE 102 - holotype; referred material: PSS-MAE 105 and 106; 3 individuals | ||||||||||
Mammalia
- Asioryctidae
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Asioryctes nemegtensis
Kielan-Jaworowska 1975
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Novacek et al. 2022 | |||||||||
PSS-MAE 366 | ||||||||||
Mammalia
- Deltatheroida
- Deltatheridiidae
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Deltatheridium pretrituberculare
Gregory and Simpson 1926
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Rougier et al. 1998 | 2 specimens | ||||||||
Mammalia
- Allotheria
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Allotheria indet.
Marsh 1880
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"new genus C" | ||||||||||
Mammalia
- Multituberculata
- Eucosmodontidae
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Eucosmodontidae indet.
Jepsen 1940
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"new genus B" | ||||||||||
Mammalia
- Multituberculata
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Bulganbaatar cf. nemegtbaataroides
Kielan-Jaworowska 1974
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"new genus A" | ||||||||||
Mammalia
- Multituberculata
- Djadochtatheriidae
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Tombaatar sabuli n. gen., n. sp.
Rougier et al. 1997
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Rougier et al. 1997 | 1 individual | ||||||||
PSS-MAE 122 (partial skull) | ||||||||||
Catopsbaatar sp.
Kielan-Jaworowska 1994
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This is a typo? "Catopsbaatar Catopsaloides" | ||||||||||
Mammalia
- Multituberculata
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Nemegtbaatar gobiensis
Kielan-Jaworowska 1974
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Mammalia
- Multituberculata
- Sloanbaataridae
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Kamptobaatar cf. kuczynskii
Kielan-Jaworowska 1969
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Sloanbaatar cf. mirabilis
Kielan-Jaworowska 1969
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Mammalia
- Multituberculata
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Chulsanbaatar cf. vulgaris
Kielan-Jaworowska 1974
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see common names |
Geography
Country: | Mongolia | State/province: | Omnogov Aimak |
Coordinates: | 43.7° North, 100.9° East (view map) | ||
Paleocoordinates: | 40.9° North, 90.2° East | ||
Basis of coordinate: | based on nearby landmark | ||
Geographic resolution: | local area |
Time
Period: | Cretaceous | Epoch: | Late/Upper Cretaceous |
Stage: | Campanian | 10 m.y. bin: | Cretaceous 7 |
Key time interval: | Late/Upper Campanian | ||
Age range of interval: | 83.60000 - 72.10000 m.y. ago |
Stratigraphy
Formation: | Djadokhta | ||||
Stratigraphy comments: Age is not well constrained for the Djadokhta Formation (Clark et al. 2001). "Fossil vertebrate sites occur in the uppermost 65 m of a section..." see Fig. 1b, Dashzeveg et al. (1995).
Dashzeveg et al. 2005: The quick stratigraphic succession of normal and reversed magnetozones suggests, but does not clearly establish, that the sediments may have been deposited during the rapid sequence of polarity changes in the late part of the Campanian between about 75 to 71 Ma. |
Lithology and environment
Primary lithology: | "cross stratification",fine,coarse,red silty sandstone |
Secondary lithology: | red or brown mudstone |
Includes fossils? | Y |
Includes fossils? | Y |
Lithology description: "Fossil vertebrate sites occur in the uppermost 65 m of a section dominated by red-orange, slightly silty, fine-to-coarse-grained sandstones with occasional beds of red-brown mudstones and grey-brown conglomerates (Fig. 1b)...The most fossiliferous sites are stratigraphically situated in a prominent cross-bedded unit of sandstone near the middle of the section." --Dashzeveg et al. 1995 | |
Environment: | eolian indet. |
Taphonomy
Modes of preservation: | body |
Degree of concentration: | -multiple events |
Size of fossils: | macrofossils,mesofossils |
Spatial orientation: | life position |
Preservation of anatomical detail: | excellent |
Articulated whole bodies: | some |
Associated major elements: | some |
Disassociated major elements: | some |
Disassociated minor elements: | some |
Size sorting: | poor |
Fragmentation: | occasional |
Encrustation: | none |
Temporal resolution: | snapshot |
Spatial resolution: | autochthonous |
Collection methods and comments
Collection methods: | surface (float),surface (in situ),field collection |
Reason for describing collection: | taxonomic analysis | Collection dates: | 1992-1996 |
Collection method comments: Institute of Geology, Mongolia | |
Taxonomic list comments:Over 100 recovered and uncollected articulated skeletons of theropod, ankylosaurian, and protoceratopsian dinosaurs form the main 4 sq km collecting area. Also skulls (many with associated skeletons) of over 400 mammals and lizards. These are just the specimen numbers they include, there are certainly more in number. |
Metadata
Database number: | 37184 | ||
Authorizer: | J. Alroy, R. Benson, M. Carrano, P. Mannion | Enterer: | R. Whatley, J. Alroy, R. Benson, M. Carrano, P. Mannion |
Modifier: | G. Varnham | Research group: | vertebrate |
Collections that are a subset of this one: | |||
Created: | 2004-03-03 16:56:49 | Last modified: | 2022-01-28 08:04:32 |
Access level: | the public | Released: | 2004-03-03 16:56:49 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
5757. | ETE 1515 | D. Dashzeveg, M. J. Novacek, M.A. Norell, J.M. Clark, L.M. Chiappe, A. Davidson, M.C. McKenna, L. Dingus, C. Swisher and P. Altangerel. 1995. Extraordinary preservation in a new vertebrate assemblage from the Late Cretaceous of Mongolia. Nature 374:446-449 [A. Behrensmeyer/A. Behrensmeyer/R. Whatley] |
Secondary references:
13618 | ETE | L. M. Chiappe, M. A. Norell, and J. M. Clark. 1998. The skull of a relative of the stem-group bird Mononykus. Nature 392:275-278 [M. Carrano/M. Carrano/M. Carrano] |
28690 | L. M. Chiappe, M. Norell, and J. Clark. 2001. A new skull of Gobipteryx minuta (Aves, Enantiornithes) from the Cretaceous of the Gobi Desert. American Museum Novitates 3346:1-15 [J. Alroy/J. Alroy] | |
38116 | J. L. Conrad and M. A. Norell. 2007. A complete Late Cretaceous iguanian (Squamata, Reptilia) from the Gobi and identification of a new iguanian clade. American Museum Novitates 3584:1-47 [R. Benson/R. Benson] | |
15459 | ETE | D. Dashzeveg, L. Dingus, D.B. Loope, C.C. Swisher, III, T. Dulam and M.R. Sweeney. 2005. New stratigraphic subdivision, depositional environment, and age estimate for the Upper Cretaceous Djadokhta Formation, southern Ulan Nur Basin, Mongolia. American Museum Novitates (3498)1-31 [M. Carrano/K. Maguire/G. Varnham] |
19734 | 5% 54920 | K. Gao and M. A. Norell. 2000. Taxonomic composition and systematics of Late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert. Bulletin of the American Museum of Natural History 249:1-118 [J. Alroy/E. Leckey/R. Benson] |
38066 | K. -Q. Gao and M. A. Norell. 1998. Taxonomic revision of Carusia (Reptilia: Squamata) from the Late Cretaceous of the Gobi Desert and phylogenetic relationships of anguimorphan lizards. American Museum Novitates 3230:1-51 [R. Benson/R. Benson] | |
43987 | I. Horovitz. 2003. Postcranial skeleton of Ukhaatherium nessovi (Eutheria, Mammalia) from the Late Cretaceous of Mongolia. Journal of Vertebrate Paleontology 23(4):857-868 [P. Mannion/P. Mannion] | |
85503 | M. J. Novacek, E. A. Hoffman, and M. A. O'Leary. 2022. First occurrence of the eutherian mammal Asioryctes nemegtensis from the Upper Cretaceous Djadokhta Formation, Gobi Desert, Mongolia, and a revised alpha taxonomy based on the skull and dentition. Journal of Vertebrate Paleontology 42(4):e2196320 [P. Mannion/P. Mannion] | |
28809 | M. J. Novacek, G. W. Rougier, J. R. Wible, M. C. McKenna, D. Dashzeveg and I. Horovitz. 1997. Epipubic bones in eutherian mammals from the Late Cretaceous of Mongolia. Nature 389:483-486 [J. Alroy/J. Alroy] | |
38082 | D. Pol, A. H. Turner, and M. A. Norell. 2009. Morphology of the Late Cretaceous crocodylomorph Shamosuchus djadochtaensis and a discussion of neosuchian phylogeny as related to the origin of Eusuchia. Bulletin of the American Museum of Natural History 324:1-103 [P. Mannion/P. Mannion] | |
2986 | G. W. Rougier, M. J. Novacek, and D. Dashzeveg. 1997. A new multituberculate from the Late Cretaceous locality Ukhaa Tolgod, Mongolia. Considerations on multituberculate interrelationships. American Museum Novitates 3191:1-26 [J. Alroy/J. Alroy/R. Benson] | |
9959 | 5% 2220 | G. W. Rougier, J. R. Wible, and M. J. Novacek. 1998. Implications of Deltatheridium specimens for early marsupial history. Nature 396(6710):459-463 [J. Alroy/R. Whatley/R. Whatley] |
44530 | H.-Y. Yi and M. A. Norell. 2013. New materials of Estesia mongoliensis (Squamata: Anguimorpha) and the evolution of venom grooves in lizards. American Museum Novitates 3767:1-31 [P. Mannion/P. Mannion] |