Stark Quarry, Schamhaupten: Late/Upper Kimmeridgian, Germany
collected by H. Balzer, G. Viohl, U. Gerhard, M. Röper 1988–1998

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
"Protoglobigerina" sp.
Viohl and Zapp 2007
Coeloidea indet.
Renesto and Viohl 1997
Coccolithophorida indet.
Viohl and Zapp 2007
Anthozoa - Scleractinia
Scleractinia indet. Bourne 1900
Renesto and Viohl 1997
unclassified
Muensteria vermicularis Sternberg 1833
Viohl and Zapp 2007
Coelacanthimorpha - Coelacanthiformes - Latimeriidae
Undina pencillata
Viohl and Zapp 2007
Reptilia
Eurysternum wagleri Meyer 1839
Karl et al. 2012 2 individuals
JME Scha 97, 70
Solnhofia parsonsi Gaffney 1975
Joyce 2000 1 individual
JME Scha 70
Reptilia - Eurysternidae
Eurysternidae indet. Dollo 1886
Joyce and Zelenitsky 2002 1 individual
JM SCHA 101, shell with eggs
Reptilia - Ichthyosauria
Ichthyosauria indet. de Blainville 1835
Viohl and Zapp 2007 1 specimen
sclerotic ring
Reptilia - Sphenodontidae
Leptosaurus pulchellus (Zittel 1887)
Renesto and Viohl 1997 1 individual
JME SCHA 40
Reptilia
Juravenator starki n. gen., n. sp. Göhlich and Chiappe 2006
Göhlich and Chiappe 2006 1 individual
JME SCh 200 (8 measurements)
Reptilia - Rhamphorhynchidae
Rhamphorhynchus sp. Meyer 1847
Viohl and Zapp 2007 2 specimens
Reptilia - Metriorhynchidae
Dakosaurus maximus (Plieninger 1846)
Andrade et al. 2010
Actinopteri - Lepisosteiformes
Lepidotes sp. Agassiz 1832
Viohl and Zapp 2007
JME Scha 80
    = Scheenstia zappi n. sp. López-Arbarello and Sfercoa 2009
López-Arbarello and Sfercoa 2009
Actinopteri - Macrosemiiformes
Voelklichthys comitatus n. gen., n. sp. Arratia and Schultze 2012
Arratia and Schultze 2012 1 individual
JM-E SCH 102 (found on same slab as Juravenator starki holotype)
Actinopteri - Pycnodontiformes - Gyrodontidae
Gyrodus sp. Agassiz 1833
Viohl and Zapp 2007
Actinopteri - Ichthyodectiformes
Allothrissops sp. Nybelin 1964
1 individual
BM.f.7286
Ascalabothrissops voelkli n. gen., n. sp. Arratia 2000
1 individual
JM SCh 30a, b
Actinopteri - Elopiformes - Elopidae
Anaethalion zapporum n. sp. Arratia 2000
1 individual
JM SCh 85
? Elopsomolos sp. Arratia 2000
= "Elops-like sp. 1"
Actinopteri - Leptolepidiformes - Leptolepididae
Tharsis sp. Giebel 1848
Viohl and Zapp 2007
synonym of Leptolepis
Chondrichthyes - Heterodontiformes - Heterodontidae
Paracestracion viohli n. sp. Kriwet 2008
Kriwet 2008 1 individual
JM Scha 728
Crinoidea - Millericrinida - Millericrinidae
Millericrinus sp. d'Orbigny 1840
Viohl and Zapp 2007
Crinoidea - Roveacrinida - Saccocomidae
Saccocoma tenella (Goldfuss 1831)
Viohl and Zapp 2007
original and current combination Comatula tenella
Echinoidea
Echinoidea indet. (Leske 1778)
Renesto and Viohl 1997
Echinoidea - Pygasteridae
Pygaster sp.
Viohl and Zapp 2007
Stenolaemata - Cyclostomata - Diastoporidae
"Berenicea" sp. Lamouroux 1821
Viohl and Zapp 2007
Brachiopoda
Brachiopoda indet. (Cuvier 1805)
Renesto and Viohl 1997
Rhynchonellata - Terebratulida - Terebratulidae
Terebratulidae indet. Gray 1840
Viohl and Zapp 2007
Rhynchonellata - Terebratulida - Zeilleriidae
Ornithella pentagonalis
Viohl and Zapp 2007
Rhynchonellata - Rhynchonellida - Cyclothyrididae
Torquirhynchia sp. Childs 1969
Viohl and Zapp 2007
Rhynchonellata - Rhynchonellida - Basiliolidae
Lacunosella sp. Wisniewska 1932
Viohl and Zapp 2007
Bivalvia
Bivalvia indet. Linnaeus 1758
Renesto and Viohl 1997
Bivalvia - Ostreida - Gryphaeidae
Liostrea socialis
Viohl and Zapp 2007
Bivalvia - Pectinida - Propeamussiidae
Propeamussium nonarium
Viohl and Zapp 2007
original and current combination Pecten (Propeamussium)
Cephalopoda - Belemnitida - Mesohibolitidae
Hibolithes semisulcatus
Viohl and Zapp 2007
Cephalopoda - Belemnitida - Belemnotheutidae
Acanthoteuthis sp. Wagner in Munster 1839
Viohl and Zapp 2007
Cephalopoda - Octobrachia - Muensterellidae
Engeseriteuthis arcuatus n. gen., n. sp. Fuchs et al. 2019
Fuchs et al. 2019 1 specimen
JME SOS Scha 709
Cephalopoda - Octobrachia
Vampyromorpha indet. Robson 1929
Viohl and Zapp 2007
corrected as Vampyromorphida
Cephalopoda - Ammonoidea
Ammonoidea indet. Zittel 1884
Renesto and Viohl 1997
35 spp.
Cephalopoda - Ammonitida - Oppeliidae
Oppeliidae indet. Bonarelli 1894
Viohl and Zapp 2007
Gastropoda - Pleurotomariidae
Bathrotomaria reticulata
Viohl and Zapp 2007
Gastropoda - Trochidae
Buckmanina sp. Cossmann 1920
Viohl and Zapp 2007
Insecta - Orthoptera
Orthoptera indet. Olivier 1789
Renesto and Viohl 1997
    = Grylloidea indet. Laicharting 1781
Viohl and Zapp 2007
Malacostraca - Isopoda
Isopoda indet. Latreille 1817
Viohl and Zapp 2007
on skeleton of Juravenator
Malacostraca - Decapoda
Palinuroidea indet. Latreille 1802
Viohl and Zapp 2007
Malacostraca - Decapoda - Axiidae
Etallonia hoellorum n. sp. Polz 1999
Viohl and Zapp 2007
Malacostraca - Decapoda - Glypheidae
Glyphea sp. Von Meyer 1835
Viohl and Zapp 2007
Malacostraca - Decapoda - Coleiidae
Coleia longipes (Fraas 1855)
Viohl and Zapp 2007
JME Scha 1531
Ostracoda
Ostracoda indet. Latreille 1802
Renesto and Viohl 1997
Phaeophyceae
Phaeophyceae indet.
Viohl and Zapp 2007
Sphenopteridae
Sphenopteris sp. Sternberg 1825
Viohl and Zapp 2007
Equisetopsida - Araucariaceae
Araucarites sp. Presl 1833
Viohl and Zapp 2007
Brachyphyllum sp. Brongniart 1828
Renesto and Viohl 1997
Gymnospermopsida - Corystospermales - Corystospermaceae
Cycadopteris jurensis
Viohl and Zapp 2007
unclassified
Goniolina sp. de Saporta 1891
Viohl and Zapp 2007
"Tubiphytes" morronensis Crescenti 1969
Viohl and Zapp 2007
recombined as Crescentiella morronensis
see common names

Geography
Country:Germany State/province:Bayern County:Eichstätt
Coordinates: 48.9° North, 11.6° East (view map)
Paleocoordinates:40.2° North, 19.7° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Jurassic Epoch:Late/Upper Jurassic
Stage:Kimmeridgian 10 m.y. bin:Jurassic 5
Key time interval:Late/Upper Kimmeridgian
Age range of interval:152.20000 - 149.20000 m.y. ago
Stratigraphy
Geological group:Weißjura Formation:Painten
Stratigraphic resolution:group of beds
Stratigraphy comments: Malm Epsilon 2 (setatum subzone), Weissjura epsilon; listed as both Solnhofen Formation, and as Painten Fm./Schamhaupten Subformation, "Profilschicht E3". Beckeri zone, Ulmense subzone.
Lithology and environment
Primary lithology: cherty/siliceous lime mudstone
Secondary lithology: bindstone
Includes fossils?Y
Includes fossils?Y
Lithology description: "silicified, laminated limestone" and bindstones
Environment:lagoonal/restricted shallow subtidal
Taphonomy
Modes of preservation:body,adpression,soft parts
Lagerst�tten type:conservation
Size of fossils:macrofossils
Spatial orientation:life position
Preservation of anatomical detail:excellent
Articulated whole bodies:all
Temporal resolution:snapshot
Spatial resolution:autochthonous
Collection methods and comments
Collection methods:selective quarrying,surface (in situ),mechanical,field collection
Reason for describing collection:taxonomic analysis
Collectors:H. Balzer, G. Viohl, U. Gerhard, M. Röper Collection dates:1988–1998
Collection method comments: collected by Jura Museum and University of Bonn. Taxa not separated into beds, but clearly there are differences in abundances between layers - could be separated into collections if future publications provide more info.
Metadata
Database number:59175
Authorizer:M. Carrano, P. Mannion, J. Alroy, M. Clapham Enterer:K. Maguire, M. Clapham, C. Feng, J. Tennant, M. Carrano, J. Alroy
Modifier:M. Carrano Research group:vertebrate
Created:2006-03-20 10:17:39 Last modified:2021-12-13 14:41:29
Access level:the public Released:2006-03-20 10:17:39
Creative Commons license:CC BY
Reference information

Primary reference:
27128.ETE G. Arratia. 2000. Remarkable teleostean fishes from the Late Jurassic of southern Germany and their phylogenetic relationships. Mitteilungen aus dem Museum für Naturkunde in Berlin, Geowissenschaften Reihe 3:137-179 [M. Carrano/M. Carrano]

Secondary references:

43156 M. B. Andrade, M. T. Young, J. B. Desojo and S. L. Brusatte. 2010. The evolution of extreme hypercarnivory in Metriorhynchidae (Mesoeucrocodylia: Thalattosuchia) based on evidence from microscopic denticle morphology. Journal of Vertebrate Paleontology 30(5):1451-1465 [M. Young/M. Young]
71014 G. Arratia and H.-P. Schultze. 2012. The macrosemiiform fish companion of the Late Jurassic theropod Juravenator from Schamhaupten, Bavaria, Germany. Fossil Record 15(5):5-25 [M. Carrano/M. Carrano]
38017 L. M. Chiappe and U. B. Göhlich. 2010. Anatomy of Juravenator starki (Theropoda: Coelurosauria) from the Late Jurassic of Germany. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 258(3):257-296 [M. Carrano/M. Carrano]
69677 D. Fuchs, Y. Iba, A. Heyng, M. Iijima, C. Klug, N. L. Larson, and G. Schweigert. 2019. The Muensterelloidea: phylogeny and character evolution of Mesozoic stem octopods. Papers in Palaeontology 6:31-92 [R. Butler/E. Dunne/M. Clapham]
16916ETE U. B. Göhlich and L. M. Chiappe. 2006. A new carnivorous dinosaur from the Late Jurassic Solnhofen archipelago. Nature 440:329-332 [M. Carrano/K. Maguire/M. Carrano]
26842ETE U. B. Göhlich, H. Tischlinger, and L. M. Chiappe. 2006. Juravenator starki (Reptilia, Theropoda), ein neuer Raubdinosaurier aus dem Oberjura der Südlichen Frankenalb (Süddeutschland): Skelettanatomie und Weichteilbefunde [Juravenator starki (Reptilia, Theropoda), a new predatory dinosaur from the Upper Jurassic of the southern Franconian Alb (southern Germany): skeletal anatomy and soft tissues]. Archaeopteryx 24:1-26 [M. Carrano/M. Carrano]
27130ETE W. Joyce and D. K. Zelenitsky. 2002. Turtle egg pseudomorphs from the Late Jurassic of Schamhaupten, Germany. Archaeopteryx 20:57-62 [M. Carrano/M. Carrano]
44043 W. G. Joyce. 2000. The first complete skeleton of Solnhofia parsoni (Cryptodira, Eurysternidae) from the Upper Jurassic of Germany and its taxonomic implications. Journal of Paleontology 74(4):684-700 [R. Benson/R. Benson]
46818 H.-V. Karl, G. Tichy, and D. Valdisberri. 2012. Sontiochelys cretacea Stache, 1905 and new description of the families Eurysternidae Dollo, 1886 and Thalassemydidae Rutimeyer, 1873 (Testudines: Cryptodira). Studia Geologica Salmanticensia 48(1):45-76 [P. Mannion/J. Tennant/P. Wagner]
30504 J. Kriwet. 2008. Anew species of extinct bullhead sharks, Paracestracion viohli sp. nov. (Neoselachii, Heterodontiformes), from the Upper Jurassic of South Germany. Acta Geologica Polonica 58(2):235-241 [J. Alroy/J. Alroy]
37653 A. López-Arbarello and E. Sfercoa. 2009. New Semionotiform (Actinopterygii, Neopterygii) from the Late Jurassic of Southern Germany. Journal of Systematic Palaeontology 9:197-215 [M. Clapham/C. Feng/M. Clapham]
64581 O. W. M. Rauhut, C. Foth, and H. Tischlinger. 2018. The oldest Archaeopteryx (Theropoda: Avialiae): a new specimen from the Kimmeridgian/Tithonian boundary of Schamhaupten, Bavaria. PeerJ 6:e4191 [M. Uhen/M. Uhen]
78879 A. G. Reisdorf and M. Wuttke. 2012. Re-evaluating Moodie’s opisthotonic-posture hypothesis in fossil vertebrates part I: reptiles—the taphonomy of the bipedal dinosaurs Compsognathus longipes and Juravenator starki from the Solnhofen Archipelago (Jurassic, Germany). Palaeobiodiversity and Palaeoenvironments 92(1):119-168 [M. Carrano/M. Carrano]
27129ETE S. Renesto and G. Viohl. 1997. A sphenodontid (Reptilia, Diapsida) from the late Kimmeridgian of Schamhaupten (Southern Franconian Alb, Bavaria, Germany). Archaeopteryx 15:27-46 [M. Carrano/M. Carrano]
27315ETE G. Viohl and M. Zapp. 2007. Schamhaupten, an outstanding Fossil-Lagerstätte in a silicified Plattenkalk around the Kimmeridgian-Tithonian boundary (Southern Franconian Alb, Bavaria). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 245(1):127-142 [M. Carrano/M. Carrano]