Daxishan, Linglongta: Late/Upper Bathonian - Early/Lower Kimmeridgian, China

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Bennettitales
Zamites cf. gigas (Lindley and Hutton 1831)
Peng et al. 2012
Polypodiopsida - Cyatheales - Cyatheaceae
Coniopteris margaretae
Peng et al. 2012
Coniopteris hymenophylloides (Brongniart 1828)
Peng et al. 2012
original and current combination Sphenopteris hymenophylloides
Polypodiopsida - Equisetales - Calamitaceae
Neocalamites carcinoides
Peng et al. 2012
Neocalamites nathorsti
Peng et al. 2012
Ginkgoopsida - Ginkgoales
Sphenobaiera kazachstanica
Peng et al. 2012
Ginkgoopsida - Czekanowskiales
Czekanowskia rifida
Peng et al. 2012
Czekanowskia ketova
Peng et al. 2012
Phoenicopsis speciosa Heer 1876
Peng et al. 2012
Reptilia - Troodontidae
Anchiornis huxleyi n. gen., n. sp. Xu et al. 2009
Xu et al. 2009 2 individuals
IVPP V14378, BMNHC PH828 (5 measurements)
Xiaotingia zhengi n. gen., n. sp. Xu et al. 2011
Xu et al. 2011 1 individual
(6 measurements)
Reptilia
Tianyulong confuciusi n. gen., n. sp. Zheng et al. 2009
Zheng et al. 2009 1 individual
STMN 26-3, incomplete skull and postcranium with filamentous integumentary structures
Reptilia - Scaphognathidae
Jianchangnathus robustus n. gen., n. sp. Cheng et al. 2012
Cheng et al. 2012 1 individual
IVPP V16866, partial skeleton with complete skull and lower jaw
cf. Jianchangnathus robustus Cheng et al. 2012
Li et al. 2014 1 individual
BMNHC PH000988
Fenghuangopterus lii n. gen., n. sp. Lü et al. 2010
Lü et al. 2010 1 individual
CYGB-0037 (holotype), incomplete skeleton with a partial skull and lower jaws
Reptilia - Wukongopteridae
Kunpengopterus sinensis n. gen., n. sp. Wang et al. 2010
Wang et al. 2010 1 individual
IVPP V16047 (holotype), nearly complete skeleton
Kunpengopterus antipollicatus n. sp. Zhou et al. 2021
Zhou et al. 2021 1 specimen
Holotype: BPMC 0042.
Reptilia
Wukongopterus lii n. gen., n. sp. Wang et al. 2009
1 individual
IVPP V15113 (holotype), nearly complete skeleton
Darwinopterus sp. Lü et al. 2010
Lü et al. 2011 1 individual
ZMNH M8802, complete skeleton with egg
Darwinopterus robustodens n. sp. Lü et al. 2011
Lü et al. 2011 1 individual
41HIII-0309A
Darwinopterus modularis n. gen., n. sp. Lü et al. 2010
Lü et al. 2010 2 individuals
ZMNH M8782 (holotype), skull and partial postcranium of a putative adult. YH-2000, almost complete skeleton of a putative female
Darwinopterus linglongtaensis n. sp. Wang et al. 2010
Wang et al. 2010 1 individual
IVPP V16049 (holotype), complete articulated specimen
Douzhanopterus zhengi n. gen., n. sp. Wang et al. 2017
Wang et al. 2017 1 specimen
STM 19–35A and B
Changchengopterus pani Lü 2009
Zhou and Schoch 2011 1 individual
PMOL-AP00010, articulated postcranial skeleton
Reptilia - Rhamphorhynchidae
Scaphognathus robustus n. sp. Cheng 2012
Bennett 2014 2 individuals
Holotype: IVPP V16866, incomplete articulated skeleton. Holotype: IVPP V16866, incomplete articulated skeleton.
Jianchangopterus zhaoianus n. gen., n. sp. Lü and Bo 2011
Lü and Bo 2011 1 individual
YHK-0931, complete skeleton
Reptilia - Istiodactylidae
Archaeoistiodactylus linglongtaensis n. gen., n. sp. Lü and Fucha 2010
Lü and Fucha 2010 1 individual
JPM04-0008, partial skeleton
Reptilia - Testudines
Testudines indet. Batsch 1788
Li et al. 2014 1 individual
PKUP V1070
Mammalia
Juramaia sp. Luo et al. 2011
Sullivan et al. 2014
Amphibia - Temnospondyli - Urodela
Urodela indet. (Duméril 1805)
Peng et al. 2012
Chunerpeton tianyiensis Gao and Shubin 2003
Sullivan et al. 2014
Ptycholepiformes - Ptycholepididae
? Ptycholepidae indet. (Brough 1939)
Peng et al. 2012
corrected as Ptycholepididae
Bivalvia - Cardiida - Shaanxiconchidae
Shaanxiconcha cliovata
Peng et al. 2012
Branchiopoda - Euestheriidae
Euestheria sp. Depéret and Mazeran 1912
Peng et al. 2012
Ostracoda - Podocopida - Limnocytheridae
Timiriasevia catenularia
Peng et al. 2012
Ostracoda - Podocopida - Darwinulidae
Darwinula sarytirmenensis
Peng et al. 2012
Darwinula impudica
Peng et al. 2012
Darwinula magna
Peng et al. 2012
Darwinula submagna
Peng et al. 2012
Darwinula changxinensis
Peng et al. 2012
Darwinula xiaofenzhangziensi
Peng et al. 2012
Insecta
Insecta indet. Linnaeus 1758
Peng et al. 2012
see common names

Geography
Country:China State/province:Liaoning County:Jianchang
Coordinates: 40.9° North, 120.0° East (view map)
Paleocoordinates:42.1° North, 123.2° East
Basis of coordinate:stated in text
Geographic resolution:local area
Time
Period:Jurassic
Key time interval:Late/Upper Bathonian - Early/Lower Kimmeridgian
Age range of interval:168.20000 - 152.20000 m.y. ago
Age estimate:minimum 160.7 ± 3.2 Ma (U/Pb)
Stratigraphy
Formation:Tiaojishan
Local section:Linglongta Local bed:13
Local order:bottom to top
Stratigraphic resolution:bed
Stratigraphy comments: Wang et al. (2009) referred to this locality as from the Daohugou Formation (?Late Jurassic-Early Cretaceous), while Lü et al. (2009) suggested that it is the Tiaojishan Formation (Middle Jurassic).

"The Tiaojishan Formation is generally regarded as Middle (Zhang & Zheng 1991; Liu et al. 2006; Cheng & Li 2007) to Upper Jurassic (Chang et al. 2009) (Late Bathonian-Early Kimmeridgian) in age, although it may date to the early Middle Jurassic (Aalenian) in Jianchang County (Davis et al. 2001). The unit has been dated using radiometric techniques at 169-152 Mya by Liu et al. (2006) and 165-156 Mya by Zhang et al. (2008)." (Lü et al. 2010: supplement)
Lithology and environment
Primary lithology:tuffaceous,gray sandstone
Secondary lithology:gray,green mudstone
Includes fossils?Y
Lithology description: "The muddy shales and mudstones, and the tuffaceous layers, have yielded an important fossil assemblage" (Lü et al. 2011: supplement) Bed 13: "Grayish-green and shallow grayish mudstone and muddy shale, occasionally tuffs and tuffaceous sandstone…fossils…were unearthed from tuffaceous sediments or tuff in the lower parts of this bed."
Environment:lacustrine indet.
Taphonomy
Modes of preservation:body,adpression,soft parts
Lagerst�tten type:conservation
Size of fossils:macrofossils
Preservation of anatomical detail:excellent
Abundance in sediment:common
Articulated whole bodies:all
Fragmentation:none
Temporal resolution:snapshot
Collection methods and comments
Collection excludes:some genera
Collection methods:selective quarrying,surface (in situ),mechanical,field collection,survey of museum collection
Reason for describing collection:taxonomic analysis
Museum repositories:IVPP
Collection method comments: ZMNH: Zhejiang Museum of Natural History. YH: Yizhou Museum, Yixian, Liaoning.
PMOL: Paleontological Museum of Liaoning, Shenyang Normal University.
Fossils collected primarily by local farmers
The heterodontosaurid ornithischian Tianyulong was originally reported to be from the Yixian Formation (Early Cretaceous) of Jianchang County, but is now believed to be from Linglongta (Lü et al. 2011: supplementary data)
Taxonomic list comments:It is now known where in this area the Xiaotingia holotype was found, and it could have come from the Yixian Fm., which also outcrops in the area.
Metadata
Also known as:Wukongopterus, Darwinopterus type locality, Linglengta
Database number:92421
Authorizer:R. Butler, P. Mannion, E. Dunne, M. Carrano, J. Alroy Enterer:R. Butler, M. Carrano, E. Dunne, J. Tennant, J. Alroy
Modifier:M. Carrano Research group:vertebrate
Created:2009-11-11 08:55:21 Last modified:2014-01-24 16:34:58
Access level:the public Released:2009-11-11 08:55:21
Creative Commons license:CC BY
Reference information

Primary reference:
31153. X. Wang, A. W. A. Kellner, S. Jiang and X. Meng. 2009. An unusual long-tailed pterosaur with elongated neck from western Liaoning of China. Anais da Academia Brasileira de Ciências 81(4):793-812 [R. Butler/R. Butler]

Secondary references:

50319 S. C. Bennett. 2014. A new speicmen of the pterosaur Scaphognathus crassirostris, with comments on constraint of cervical vertebrae number in pterosaurs. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 271(3):327-348 [P. Mannion/J. Tennant]
43454 X. Cheng, X. Wang, S. Jiang and A. W. A. Kellner. 2012. A new scaphognathid pterosaur from western Liaoning, China. Historical Biology 24(1):101-111 [R. Butler/R. Butler]
52405 Q. Li, J. A. Clarke, K.-Q. Gao, C.-F. Zhou, Q. Meng, D. Li, L. D'Alba and M. D. Shawkey. 2014. Melanosome evolution indicates a key physiological shift within feathered dinosaurs. Nature 507:350-353 [M. Carrano/M. Carrano]
38026 J. Lü and X. Bo. 2011. A new rhamphorhynchid pterosaur (Pterosauria) from the Middle Jurassic Tiaojishan Formation of western Liaoning, China. Acta Geologica Sinica 86(5):977-983 [R. Butler/R. Butler/R. Butler]
35167 J. Lü and X. Fucha. 2010. A new pterosaur (Pterosauria) from Middle Jurassic Tiaojishan Formation of western Liaoning, China. Global Geology 13:113-118 [R. Butler/R. Butler]
31222 J. Lü, D. M. Unwin, X. Jin, Y. Liu, and Q. Ji. 2010. Evidence for modular evolution in a long-tailed pterosaur with a pterodactyloid skull. Proceedings of the Royal Society B 277:383-389 [R. Butler/R. Butler/R. Butler]
34847 J. Lü, D. M. Unwin, D. C. Deeming, X. Jin, Y. Liu and Q. Ji. 2011. An egg-adult association, gender, and reproduction in pterosaurs. Science 331(6015):321-324 [R. Butler/R. Butler]
37985 J. Lü, L. Xu, H. Chang and X. Zhang. 2011. A new darwinopterid pterosaur from the Middle Jurassic of western Liaoning, northeastern China and its ecological implications. Acta Geologica Sinica 85(3):507-514 [M. Carrano/M. Carrano]
32590 J.-C. Lü, X.-H. Fucha, and J.-M. Chen. 2010. A new scaphognathine pterosaur from the Middle Jurassic of western Liaoning, China. Acta Geoscientica Sinica 31(2):263-266 [R. Butler/R. Butler]
49934 N. Peng, Y. Liu, H. Kuang, X. Jiang, and H. Xu. 2012. Stratigraphy and geochronology of vertebrate fossil-bearing Jurassic strata from Linglongta, Jianchang county, western Liaoning, northeastern China. Acta Geologica Sinica 86(6):1326-1339 [M. Carrano/M. Carrano]
50313 C. Sullivan, Y. Wang, D. W. E. Hone, Y. Wang, X. Xu and F. Zhang. 2014. The vertebrates of the Jurassic Daohugou biota of northeastern China. Journal of Vertebrate Paleontology 34(2):243-280 [P. Mannion/J. Tennant]
34511 X. Wang, A. W. A. Kellner, S. Jiang, X. Cheng, X. Meng and T. Rodrigues. 2010. New long-tailed pterosaurs (Wukongopteridae) from western Liaoning, China. Anais da Academia Brasileira de Ciências 82(4):1045-1062 [R. Butler/R. Butler/P. Wagner]
67473 X.-L. Wang, S.-X. Jiang, J.-Q. Zhang, X. Cheng, X.-F. Yu, Y.-M. Li, G.-J. Wei and X.-L. Wang. 2017. New evidence from China for the nature of the pterosaur evolutionary transition. Scientific Reports 7(42763) [G. Lloyd/G. Lloyd]
36800 X. Xu, H. You, K. Du and F. Han. 2011. An Archaeopteryx-like theropod from China and the origin of Avialae. Nature 475:465-470 [J. Alroy/J. Alroy]
35452ETE X. Xu, Q. Zhao, M. Norell, C. Sullivan, D. Hone, G. Erickson, X. Wang, F. Han, and Y. Guo. 2009. A new feathered maniraptoran dinosaur fossil that fills a morphological gap in avian origin. Chinese Science Bulletin 54(3):430-435 [M. Carrano/M. Carrano]
29485 X.-T. Zheng, H.-L. You, X. Xu and Z.-M. Dong. 2009. An Early Cretaceous heterodontosaurid dinosaur with filamentous integumentary structures. Nature 458:333-336 [J. Alroy/J. Alroy/M. Carrano]
35658 C.-F. Zhou and R. R. Schoch. 2011. New material of the non-pterodactyloid pterosaur Changchengopterus pani Lü, 2009 from the Late Jurassic Tiaojishan Formation of western Liaoning. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 260:265-275 [R. Butler/R. Butler/R. Butler]
76359 X. Zhou, R. V. Pegas, W. Ma, G. Han, X. Jin, M. E.C. Leal, N. Bonde, Y. Kobayashi, S. Lautenschlager, X. Wei, C. Shen and S. Ji. 2021. Current Biology. 31:1-8 [E. Dunne/E. Dunne/E. Dunne]