Sokolki locality, PIN 2005: Late/Upper Severodvinian - Vyatkian, Russian Federation
collected by V. P. Amalitskii
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
|
Reptilia
- Pareiasauridae
|
||||||||||
|
Pareiosaurus karpinskii n. sp.
Amalitzky 1922
|
Ivakhnenko 2008 | 1 individual | ||||||||
| recombined as Scutosaurus karpinskii | ||||||||||
| PIN 2005/1532 (type) | ||||||||||
|
Scutosaurus karpinskii
(Amalitzky 1922)
|
Ivakhnenko 2008 | 9 individuals | ||||||||
| PIN 2005/1534 (a skull, listed by Lee, 1997, ?erroneously listed as 2005/1543 in Ivakhnenko, 2008), 2005/1536*, 37*, 2005/1573**, 2005/1883**, 2005/1885, 2005/2471, 72**, 2005/2546**; all are represented by skeletons or skulls; *mark specimens mentioned in Lee, 1997, but not in Ivakhnenko, 2008; **mark specimens not mentioned in Lee, 1997, but in Ivakhnenko, 2008; for additonal material listed by Lee, 1997, see comments on taxonomic list | ||||||||||
|
Pareiosaurus horridus n. sp.
Amalitzky 1922
|
Ivakhnenko 2008 | 1 individual | ||||||||
| synonym of Scutosaurus karpinskii | ||||||||||
| PIN 2005/1535 (type) | ||||||||||
|
Pareiosaurus tuberculatus n. sp.
Amalitzky 1922
|
Ivakhnenko 2008 | 1 individual | ||||||||
| synonym of Scutosaurus karpinskii | ||||||||||
| PIN 2005/1533 (type) | ||||||||||
|
Scutosaurus tuberculatus
(Amalitzky 1922)
|
Ivakhnenko 2008 | 1 individual | ||||||||
| synonym of Scutosaurus karpinskii | ||||||||||
| PIN 2005/1578 (a juvenile); see comments on taxonomic list | ||||||||||
|
Pareiosaurus elegans n. sp.
Amalitzky 1922
|
Ivakhnenko 2008 | 1 individual | ||||||||
| synonym of Scutosaurus karpinskii | ||||||||||
| PIN 2005/1538 (type) | ||||||||||
|
Therocephalia
- Annatherapsididae
|
||||||||||
|
Anna petri n. gen., n. sp.
Amalitzky 1922
|
Ivakhnenko 2011 | 10 specimens | ||||||||
| recombined as Annatherapsidus petri | ||||||||||
| PIN 2005/1993 (holotype), partial skull. PIN 2005/1562, 1991, 1992, 1994, 1995, 1996, 1997, 1998, 2155 | ||||||||||
|
Dviniidae
|
||||||||||
|
Dvinia prima n. gen., n. sp.
Amalitzky 1922
|
Battail and Surkov 2000 | 2 specimens | ||||||||
| PIN 2005/2465 (holotype), anterior half of skull. PIN 2245/1237, a right maxilla | ||||||||||
|
Permocynodon sushkini n. gen., n. sp.
Woodward 1932
|
Battail and Surkov 2000 | 1 specimen | ||||||||
| synonym of Dvinia prima | ||||||||||
| PIN 2005/2469 (holotype), almost complete skull | ||||||||||
|
Anomodontia
- Dicynodontidae
|
||||||||||
|
Elph borealis
Kurkin 1999
|
2 specimens | |||||||||
| PIN 2005/8, 2005/2643 | ||||||||||
|
Anomodontia
|
||||||||||
|
Dicynodon amalitzkii n. sp.
Seeley 1908
|
Sushkin 1926 | |||||||||
| recombined as Peramodon amalitzkii | ||||||||||
| PIN 2005/38, a well preserved complete skull and lower jaws, axial column (missing the caudal region), pectoral girdle and fragmentary pelvic girdle | ||||||||||
|
Dicynodon trautscholdi n. sp.
Amalitzky 1922
|
Amalitzky 1922 | |||||||||
| recombined as Vivaxosaurus trautscholdi | ||||||||||
| PIN 2005/1 a well preserved, complete skull and lower jaws | ||||||||||
|
Gordonia annae n. sp.
Amalitzky 1922
|
Amalitzky 1922 | |||||||||
| synonym of Vivaxosaurus trautscholdi | ||||||||||
| PIN 2005/5 a poorly preserved but nearly complete skull missing portions of the squamosals | ||||||||||
|
Gordonia rossica n. sp.
Amalitzky 1922
|
Amalitzky 1922 | |||||||||
| synonym of Vivaxosaurus trautscholdi | ||||||||||
| PIN 2005/3 a badly distorted skull | ||||||||||
|
Oudenodon venyukovi n. sp.
Amalitzky 1922
|
Amalitzky 1922 | |||||||||
| synonym of Vivaxosaurus trautscholdi | ||||||||||
| PIN 2005/6 a poorly preserved, laterally crushed skull | ||||||||||
|
Gorgonopidae
|
||||||||||
|
Inostrancevia parva n. sp.
Pravoslavlev 1927
|
Ivakhnenko 2003 | 1 individual | ||||||||
| recombined as Pravoslavlevia parva | ||||||||||
| PIN 2005/1859 (holotype), skull | ||||||||||
|
Inostranceviidae
|
||||||||||
|
Inostrancevia latifrons n. sp.
Pravoslavlev 1927
|
Ivakhnenko 2003 | 1 individual | ||||||||
| PIN 2005/1857 (type) | ||||||||||
|
Amalitzkia vladimiri n. gen., n. sp.
Pravoslavlev 1927
|
Ivakhnenko 2003 | 1 individual | ||||||||
| synonym of Inostrancevia latifrons | ||||||||||
| PIN 2005/1856 (type) | ||||||||||
|
Inostrancevia alexandri n. gen., n. sp.
Amalitski 1922
|
Ivakhnenko 2003 | 3 individuals | ||||||||
| PIN 2005/1587 (lectotype), skeleton. PIN 2005/1588, skeleton, PIN 2005/2099, skull roof | ||||||||||
|
Inostrancevia proclivis n. sp.
Pravoslavlev 1927
|
Ivakhnenko 2003 | 1 individual | ||||||||
| synonym of Inostrancevia alexandri | ||||||||||
| PIN 2005/1858 (type) | ||||||||||
|
Kotlassidae
|
||||||||||
|
Kotlassia prima n. gen., n. sp.
Amalitzky 1898
|
Bulanov 2003 | 3 specimens | ||||||||
| PIN 2005/74 (holotype). PIN 2005/75, 2689 | ||||||||||
|
Karpinskiosauridae
|
||||||||||
|
Kotlassia secunda n. sp.
Amaltzki 1898
|
Bulanov 2003 | 2 individuals | ||||||||
| recombined as Karpinskiosaurus secundus | ||||||||||
| PIN 2005/81 (holotype), 82 | ||||||||||
|
Chroniosuchidae
|
||||||||||
|
Chroniosuchus licharevi
(Riabinin 1962)
|
Golubev 1998 | 2 specimens | ||||||||
| PIN 2005/2578, 2579 | ||||||||||
|
Amphibia
- Temnospondyli
- Dvinosauridae
|
||||||||||
|
Dvinosaurus primus n. gen., n. sp.
Amalitzki 1921
|
Schoch and Milner 2014 | |||||||||
| PIN 2005/39 (holotype), skull and anterior postcranial skeleton | ||||||||||
|
Dvinosaurus secundus n. sp.
Amalitzki 1921
|
Schoch and Milner 2014 | |||||||||
| synonym of Dvinosaurus primus | ||||||||||
| PIN 2005/40 (holotype) | ||||||||||
|
Dvinosaurus tertius n. sp.
Amalitzki 1921
|
Schoch and Milner 2014 | |||||||||
| synonym of Dvinosaurus primus | ||||||||||
| PIN 2005/41 (holotype) | ||||||||||
| see common names | ||||||||||
Geography
| Country: | Russian Federation | State/province: | Arkhangelsk | County: | Kotlas |
| Coordinates: | 61.2° North, 46.6° East (view map) | ||||
| Paleocoordinates: | 33.5° North, 37.4° East | ||||
| Basis of coordinate: | based on nearby landmark | ||||
| Geographic resolution: | outcrop |
Time
| Period: | Permian | ||
| *Period: | Late/Upper Permian | *Epoch: | Zechstein |
| *International age/stage: | Tartarian | *Local age/stage: | Vyatka |
| Key time interval: | Late/Upper Severodvinian - Vyatkian | Other zone: | Scutosaurus karpinskii |
| Age range of interval: | 265.00000 - 252.30000 m.y. ago | ||
| * legacy (obsolete) database fields | |||
Stratigraphy
| Formation: | Salarevo | ||||
| Stratigraphic resolution: | group of beds | ||||
| Stratigraphy comments: Sokolki Subassemblage of the Sokolki Assemblage; the later part of the Late Tatarian (late Severodvinian–Vyatkian) (Golubev 2000; Benton 2012, Geology).
"Vyatka Regional Stage" (Kurkin, 1999). Chronostratigraphy originally entered as >>Zechstein<<. Lithostratigraphic data are from Ivakhnenko (2008); Salarevo Formation therein referred to as "Salarevskaya Svita" (TL). | |||||
Lithology and environment
| Primary lithology: | not reported | ||
| Environment: | terrestrial indet. | Tectonic setting: | cratonic basin |
Taphonomy
| Modes of preservation: | body |
| Size of fossils: | macrofossils |
| Articulated whole bodies: | many |
| Associated major elements: | many |
| Disassociated major elements: | many |
| Disassociated minor elements: | many |
| Fragmentation: | occasional |
Collection methods and comments
| Collection methods: | survey of museum collection |
| Reason for describing collection: | taxonomic analysis |
| Museum repositories: | PIN |
| Collectors: | V. P. Amalitskii |
| Collection method comments: It appears that Amalitzky collected at the Sokolki site whereas Edemski excavated at the Zavrazhe locality (end of 19th and beginning of 20th century). | |
| Taxonomic list comments:"Scutosaurus tuberculatus": Ivakhnenko (2008) lists PIN 2005/1578 among specimens of Scutosaurus karpinskii as well as under S. tuberculatus. Since Lee (1997) says that 2005/1578 is a juvenile and Ivakhnenko (2008) says the same about the specimen listed as S. tuberculatus is seems clear that 2005/1578 is meant to be referred to the latter taxon by Ivakhnenko (2008).
S. karpinskii: Lee (1997) lists 9 specimens of S. karpinskii predominantly represented by single postcranial elements (PIN 2005/123, 455, 698, 1543, 1877, 2070, 2473, 2485, 2489), which are not included in the taxonomic list of the present collection. | |
Metadata
| Also known as: | Northern Dvina; Little Northern Dvina; Malaya Severnaya Dvina; Kotlas | ||
| Database number: | 28065 | ||
| Authorizer: | J. Alroy, J. Mueller, R. Butler | Enterer: | R. Whatley, J. Benito Moreno, R. Butler, T. Liebrecht |
| Modifier: | R. Butler | Research group: | vertebrate |
| Created: | 2003-01-22 13:07:41 | Last modified: | 2017-12-04 08:35:22 |
| Access level: | the public | Released: | 2003-01-22 13:07:41 |
| Creative Commons license: | CC BY | ||
Reference information
Primary reference:
| 7660. | 5% 52920 | A. A. Kurkin. 1999. Novyy ditsinodont iz raskopok na reke Maloy Severnoy Dvine. [New dicynodont from Little Northern Dvina River excavations]. Paleontologicheskii Zhurnal 1999(3):87-92 [J. Alroy/R. Whatley/T. Liebrecht] |
Secondary references:
| 31389 | V. P. Amalitzky. 1922. Diagnoses of the new forms of Vertebrates and Plants from the Upper Permian on North Dvina. Bulletin de l’Académie des Sciences de Russie, VIème série [Izvestiya Rossijskoj Akademii Nauk, VI seriya] 16:329-340 [J. Mueller/T. Liebrecht/T. Liebrecht] | |
| 28966 | B. Battail and M. V. Surkov. 2000. Mammal-like reptiles from Russia. In M. J. Benton, M. A. Shishkin, D. M. Unwin & E. N. Kurochkin (eds.), The Age of Dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge 86-119 [J. Mueller/T. Liebrecht] | |
| 63657 | V. V. Bulanov. 2003. Evolution and systematics of seymouriamorph parareptiles. Paleontological Journal 37(Suppl. 1):S1-S105 [R. Butler/R. Butler] | |
| 63602 | V. K. Golubev. 1998. Revision of the Late Permian Chroniosuchians (Amphibia, Anthracosauromorpha) from Eastern Europe. Paleontological Journal 32(4):390-401 [R. Butler/R. Butler] | |
| 63997 | V. K. Golubev. 2000. The Faunal Assemblages of Permian Terrestrial Vertebrates from Eastern Europe. Paleontological Journal 34:S211-S224 [R. Butler/R. Butler] | |
| 63003 | M. F. Ivakhnenko. 2003. Eotherapsids from the East European Placket (Late Permian). Paleontological Journal 37(Suppl. 4):S339-S465 [R. Butler/R. Butler] | |
| 28657 | M. F. Ivakhnenko. 2008. Podklass Parareptilia. In M. F. Ivakhnenko and E. N. Kurotchkin (eds.), Iskopaemye pozvonotchnye Rossii i sopredel'nykh stran: Iskopaemye reptilii i ptitsy, Tchast' 1 [Fossil vertebrates of Russia and adjacent countries: Fossil reptiles and birds, Part 1], GEOS, Moscow 49-85 [J. Mueller/T. Liebrecht/T. Liebrecht] | |
| 46409 | M. F. Ivakhnenko. 2011. Permian and Triassic therocephals (Eutherapsida) of Eastern Europe. Paleontological Journal 45(9):981-1144 [R. Butler/R. Butler] | |
| 29616 | M. S. Y. Lee. 1997. A taxonomic revision of pareiasaurian reptiles: implications for Permian terrestrial paleoecology. Modern Geology 21:231-298 [J. Mueller/T. Liebrecht/M. Carrano] | |
| 57795 | R. R. Schoch and A. R. Milner. 2014. Handbook of Paleoherpetology Part 3A2 Temnospondyli I. [R. Butler/E. Dunne/E. Dunne] | |
| 51609 | P. P. Sushkin. 1926. Notes on the pre-Jurassic Tetrapoda from Russia. I. Dicynodon amalitzkii, n. sp. — II. Contributions to the morphology and ethology of the Anomodontia. — III. On Seymouriamorphae from the Upper Permian of North Dvina. Palaeontologia Hungarica 1:323-344 [M. Carrano/M. Carrano/J. Benito Moreno] |