Sokolki locality, PIN 2005: Late/Upper Severodvinian - Vyatkian, Russian Federation
collected by V. P. Amalitskii

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli - Dvinosauridae
Dvinosaurus primus n. gen., n. sp. Amalitzki 1921
Schoch and Milner 2014
PIN 2005/39 (holotype), skull and anterior postcranial skeleton
Dvinosaurus tertius n. sp. Amalitzki 1921
Schoch and Milner 2014
synonym of Dvinosaurus primus
PIN 2005/41 (holotype)
Dvinosaurus secundus n. sp. Amalitzki 1921
Schoch and Milner 2014
synonym of Dvinosaurus primus
PIN 2005/40 (holotype)
Chroniosuchidae
Chroniosuchus licharevi (Riabinin 1962)
Golubev 1998 2 specimens
PIN 2005/2578, 2579
Reptilia - Pareiasauridae
Pareiosaurus karpinskii n. sp. Amalitzky 1922
Ivakhnenko 2008 1 individual
recombined as Scutosaurus karpinskii
PIN 2005/1532 (type)
Scutosaurus karpinskii (Amalitzky 1922)
Ivakhnenko 2008 9 individuals
PIN 2005/1534 (a skull, listed by Lee, 1997, ?erroneously listed as 2005/1543 in Ivakhnenko, 2008), 2005/1536*, 37*, 2005/1573**, 2005/1883**, 2005/1885, 2005/2471, 72**, 2005/2546**; all are represented by skeletons or skulls; *mark specimens mentioned in Lee, 1997, but not in Ivakhnenko, 2008; **mark specimens not mentioned in Lee, 1997, but in Ivakhnenko, 2008; for additonal material listed by Lee, 1997, see comments on taxonomic list
Pareiosaurus elegans n. sp. Amalitzky 1922
Ivakhnenko 2008 1 individual
synonym of Scutosaurus karpinskii
PIN 2005/1538 (type)
Pareiosaurus tuberculatus n. sp. Amalitzky 1922
Ivakhnenko 2008 1 individual
synonym of Scutosaurus karpinskii
PIN 2005/1533 (type)
Scutosaurus tuberculatus (Amalitzky 1922)
Ivakhnenko 2008 1 individual
synonym of Scutosaurus karpinskii
PIN 2005/1578 (a juvenile); see comments on taxonomic list
Pareiosaurus horridus n. sp. Amalitzky 1922
Ivakhnenko 2008 1 individual
synonym of Scutosaurus karpinskii
PIN 2005/1535 (type)
Anomodontia - Dicynodontidae
Elph borealis Kurkin 1999
2 specimens
PIN 2005/8, 2005/2643
Anomodontia
Dicynodon trautscholdi n. sp. Amalitzky 1922
Amalitzky 1922
recombined as Vivaxosaurus trautscholdi
PIN 2005/1 a well preserved, complete skull and lower jaws
Oudenodon venyukovi n. sp. Amalitzky 1922
Amalitzky 1922
synonym of Vivaxosaurus trautscholdi
PIN 2005/6 a poorly preserved, laterally crushed skull
Gordonia annae n. sp. Amalitzky 1922
Amalitzky 1922
synonym of Vivaxosaurus trautscholdi
PIN 2005/5 a poorly preserved but nearly complete skull missing portions of the squamosals
Gordonia rossica n. sp. Amalitzky 1922
Amalitzky 1922
synonym of Vivaxosaurus trautscholdi
PIN 2005/3 a badly distorted skull
Dicynodon amalitzkii n. sp. Seeley 1908
Sushkin 1926
recombined as Peramodon amalitzkii
PIN 2005/38, a well preserved complete skull and lower jaws, axial column (missing the caudal region), pectoral girdle and fragmentary pelvic girdle
Gorgonopidae
Inostrancevia parva n. sp. Pravoslavlev 1927
Ivakhnenko 2003 1 individual
recombined as Pravoslavlevia parva
PIN 2005/1859 (holotype), skull
Inostranceviidae
Inostrancevia latifrons n. sp. Pravoslavlev 1927
Ivakhnenko 2003 1 individual
PIN 2005/1857 (type)
Amalitzkia vladimiri n. gen., n. sp. Pravoslavlev 1927
Ivakhnenko 2003 1 individual
synonym of Inostrancevia latifrons
PIN 2005/1856 (type)
Inostrancevia alexandri n. gen., n. sp. Amalitski 1922
Ivakhnenko 2003 3 individuals
PIN 2005/1587 (lectotype), skeleton. PIN 2005/1588, skeleton, PIN 2005/2099, skull roof
Inostrancevia proclivis n. sp. Pravoslavlev 1927
Ivakhnenko 2003 1 individual
synonym of Inostrancevia alexandri
PIN 2005/1858 (type)
Therocephalia - Annatherapsididae
Anna petri n. gen., n. sp. Amalitzky 1922
Ivakhnenko 2011 10 specimens
recombined as Annatherapsidus petri
PIN 2005/1993 (holotype), partial skull. PIN 2005/1562, 1991, 1992, 1994, 1995, 1996, 1997, 1998, 2155
Dviniidae
Dvinia prima n. gen., n. sp. Amalitzky 1922
Battail and Surkov 2000 2 specimens
PIN 2005/2465 (holotype), anterior half of skull. PIN 2245/1237, a right maxilla
Permocynodon sushkini n. gen., n. sp. Woodward 1932
Battail and Surkov 2000 1 specimen
synonym of Dvinia prima
PIN 2005/2469 (holotype), almost complete skull
Kotlassidae
Kotlassia prima n. gen., n. sp. Amalitzky 1898
Bulanov 2003 3 specimens
PIN 2005/74 (holotype). PIN 2005/75, 2689
Karpinskiosauridae
Kotlassia secunda n. sp. Amaltzki 1898
Bulanov 2003 2 individuals
recombined as Karpinskiosaurus secundus
PIN 2005/81 (holotype), 82
see common names

Geography
Country:Russian Federation State/province:Arkhangelsk County:Kotlas
Coordinates: 61.2° North, 46.6° East (view map)
Paleocoordinates:33.5° North, 37.4° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Permian
10 m.y. bin:Permian 4
*Period:Late/Upper Permian *Epoch:Zechstein
*International age/stage:Tartarian *Local age/stage:Vyatka
Key time interval:Late/Upper Severodvinian - Vyatkian Other zone: Scutosaurus karpinskii
Age range of interval:264.28000 - 256.80000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Salarevo
Stratigraphic resolution:group of beds
Stratigraphy comments: Sokolki Subassemblage of the Sokolki Assemblage; the later part of the Late Tatarian (late Severodvinian–Vyatkian) (Golubev 2000; Benton 2012, Geology).

"Vyatka Regional Stage" (Kurkin, 1999).
Chronostratigraphy originally entered as >>Zechstein<<.
Lithostratigraphic data are from Ivakhnenko (2008); Salarevo Formation therein referred to as "Salarevskaya Svita" (TL).
Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet. Tectonic setting:cratonic basin
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Articulated whole bodies:many
Associated major elements:many
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:occasional
Collection methods and comments
Collection methods:survey of museum collection
Reason for describing collection:taxonomic analysis
Museum repositories:PIN
Collectors:V. P. Amalitskii
Collection method comments: It appears that Amalitzky collected at the Sokolki site whereas Edemski excavated at the Zavrazhe locality (end of 19th and beginning of 20th century).
Taxonomic list comments:"Scutosaurus tuberculatus": Ivakhnenko (2008) lists PIN 2005/1578 among specimens of Scutosaurus karpinskii as well as under S. tuberculatus. Since Lee (1997) says that 2005/1578 is a juvenile and Ivakhnenko (2008) says the same about the specimen listed as S. tuberculatus is seems clear that 2005/1578 is meant to be referred to the latter taxon by Ivakhnenko (2008).

S. karpinskii: Lee (1997) lists 9 specimens of S. karpinskii predominantly represented by single postcranial elements (PIN 2005/123, 455, 698, 1543, 1877, 2070, 2473, 2485, 2489), which are not included in the taxonomic list of the present collection.
Metadata
Also known as:Northern Dvina; Little Northern Dvina; Malaya Severnaya Dvina; Kotlas
Database number:28065
Authorizer:J. Alroy, R. Butler, J. Mueller Enterer:R. Whatley, R. Butler, T. Liebrecht, J. Benito Moreno
Modifier:R. Butler Research group:vertebrate
Created:2003-01-22 13:07:41 Last modified:2017-12-04 08:35:22
Access level:the public Released:2003-01-22 13:07:41
Creative Commons license:CC BY
Reference information

Primary reference:
7660.5% 52920A. A. Kurkin. 1999. Novyy ditsinodont iz raskopok na reke Maloy Severnoy Dvine. [New dicynodont from Little Northern Dvina River excavations]. Paleontologicheskii Zhurnal 1999(3):87-92 [J. Alroy/R. Whatley/T. Liebrecht]

Secondary references:

31389 V. P. Amalitzky. 1922. Diagnoses of the new forms of Vertebrates and Plants from the Upper Permian on North Dvina. Bulletin de l’Académie des Sciences de Russie, VIème série [Izvestiya Rossijskoj Akademii Nauk, VI seriya] 16:329-340 [J. Mueller/T. Liebrecht/T. Liebrecht]
28966 B. Battail and M. V. Surkov. 2000. Mammal-like reptiles from Russia. In M. J. Benton, M. A. Shishkin, D. M. Unwin & E. N. Kurochkin (eds.), The Age of Dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge 86-119 [J. Mueller/T. Liebrecht]
63657 V. V. Bulanov. 2003. Evolution and systematics of seymouriamorph parareptiles. Paleontological Journal 37(Suppl. 1):S1-S105 [R. Butler/R. Butler]
63602 V. K. Golubev. 1998. Revision of the Late Permian Chroniosuchians (Amphibia, Anthracosauromorpha) from Eastern Europe. Paleontological Journal 32(4):390-401 [R. Butler/R. Butler]
63997 V. K. Golubev. 2000. The Faunal Assemblages of Permian Terrestrial Vertebrates from Eastern Europe. Paleontological Journal 34:S211-S224 [R. Butler/R. Butler]
63003 M. F. Ivakhnenko. 2003. Eotherapsids from the East European Placket (Late Permian). Paleontological Journal 37(Suppl. 4):S339-S465 [R. Butler/R. Butler]
28657 M. F. Ivakhnenko. 2008. Podklass Parareptilia. In M. F. Ivakhnenko and E. N. Kurotchkin (eds.), Iskopaemye pozvonotchnye Rossii i sopredel'nykh stran: Iskopaemye reptilii i ptitsy, Tchast' 1 [Fossil vertebrates of Russia and adjacent countries: Fossil reptiles and birds, Part 1], GEOS, Moscow 49-85 [J. Mueller/T. Liebrecht/T. Liebrecht]
46409 M. F. Ivakhnenko. 2011. Permian and Triassic therocephals (Eutherapsida) of Eastern Europe. Paleontological Journal 45(9):981-1144 [R. Butler/R. Butler]
29616 M. S. Y. Lee. 1997. A taxonomic revision of pareiasaurian reptiles: implications for Permian terrestrial paleoecology. Modern Geology 21:231-298 [J. Mueller/T. Liebrecht/M. Carrano]
57795 R. R. Schoch and A. R. Milner. 2014. Handbook of Paleoherpetology Part 3A2 Temnospondyli I. [R. Butler/E. Dunne/E. Dunne]
51609 P. P. Sushkin. 1926. Notes on the pre-Jurassic Tetrapoda from Russia. I. Dicynodon amalitzkii, n. sp. — II. Contributions to the morphology and ethology of the Anomodontia. — III. On Seymouriamorphae from the Upper Permian of North Dvina. Palaeontologia Hungarica 1:323-344 [M. Carrano/M. Carrano/J. Benito Moreno]