Kotelnich, Vjatka River ("Sarmin" series), PIN 2212: Early/Lower Severodvinian, Russian Federation
collected by K. Grekhov, among others 1935, 1999

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Nochnitsa geminidens n. gen., n. sp. Kammerer and Masyutin 2018
Kammerer and Masyutin 2018 1 individual
KPM 310, a nearly complete skull and lower jaws with articulated vertebrae, ribs, and partial right forelimb
Viatkogorgon ivakhnenkoi n. gen., n. sp. Tatarinov 1999
Ivakhnenko 2003 1 individual
PIN 2212/61, skeleton (type)
Anomodontia
Suminia getmanovi n. gen., n. sp. Ivakhnenko 1994
Ivakhnenko 1994 15 individuals
PIN 2212/10 (type) - incomplete skeleton, 2212/29-33 - skull; 2212/18, /22, /34-39, /52-54 - jaws; 2212/23-28, /42-51 - individual teeth
Therocephalia - Kotelcephalonidae
Kotelcephalon viatkensis n. gen., n. sp. Tatarinov 1999
Ivakhnenko 2011 1 individual
PIN 2212/91 (holotype), skull
Therocephalia
Gorynychus masyutinae n. gen., n. sp. Kammerer and Masyutin 2018
Kammerer and Masyutin 2018 1 individual
KPM 346–349
Therocephalia - Scaloposauridae
Muchia microdenta n. sp. Ivakhnenko 2011
Kammerer and Masyutin 2018
Therocephalia - Perplexisauridae
Perplexisaurus foveatus n. gen., n. sp. Tatarinov 1997
Ivakhnenko 2011 3 specimens
PIN 2212/15 (holotype), skull; 2212/19, dentary; 2212/55, skull fragment
Chlynovia serridentatus n. gen., n. sp. Tatarinov 2000
Ivakhnenko 2011 1 specimen
synonym of Perplexisaurus foveatus
PIN 2212/90 (holotype), partial skull
Therocephalia - Karenitidae
Karenites ornamentatus n. gen., n. sp. Tatarinov 1995
Ivakhnenko 2011 8 specimens
PIN 2212/60 (holotype), partial skeleton. PIN 2212/20, 21, 40, 59, 93, 96, 98
Scalopodon tenuisfrons n. gen., n. sp. Tatarinov 1999
Ivakhnenko 2011 1 individual
synonym of Karenites ornamentatus
PIN 2212/97 (holotype), partial skull
Therocephalia - Ictidosuchidae
Scalopodontes kotelnichi n. gen., n. sp. Tatarinov 2000
Ivakhnenko 2011 1 individual
PIN 2212/16 (holotype), skull
Therocephalia - Whaitsiidae
Viatkosuchus sumini n. gen., n. sp. Tatarinov 1995
Ivakhnenko 2011 2 individuals
PIN 2212/13 (holotype), skeleton; PIN 2212/94, maxilla
Reptilia - Nycteroleteridae
Emeroleter levis n. gen., n. sp. Ivakhnenko 1997
Ivakhnenko 1997 7 individuals
PIN 2212/92, an isolated, dorsoventrally flattened skull. PIN 2212/14, partial skull; PIN 2212/89, fragmentary skull; KPM uncat/E1, complete articulated skeleton; KPM uncat/E2, mostly complete articulated skeleton and skull, part and counter part; KPM uncat/E3, mostly complete skull and fragmentary postcranial remains; KPM uncat/E4, partial articulated postcranial skeleton with partial skull.
Reptilia - Pareiasauridae
Anthodon rossicus n. sp. Hartmann-Weinberg 1937
1 individual
recombined as Deltavjatia rossicus
PIN 2212/2 (type); catalogue number given in Lee (2000)
Pareiasuchus vjatkensis n. sp. Hartmann-Weinberg 1937
9 individuals
synonym of Deltavjatia rossicus
PIN 2212/1 (type); catalogue number given in Lee (1997); taxon referred to as "Pareiasaurus viatkensis" in Olson (1957); PIN 2212/6; UMZC T1321; KPM unnumbered, KPM N11/99 (a neonate, reported by Kordikova & Khlyupin, 2001); for repository of additional material see comments on taxonomic list
Anthodon chlynoviensis n. sp. Efremov 1940
Efremov 1940
synonym of Deltavjatia rossicus
Amphibia - Temnospondyli - Dvinosauridae
Dvinosaurus sp. Amalitzky 1921
Olson 1957
see common names

Geography
Country:Russian Federation State/province:Kirov County:Kotelnich
Coordinates: 58.2° North, 48.4° East (view map)
Paleocoordinates:31.8° North, 40.5° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Permian Epoch:Guadalupian
Stage:Capitanian 10 m.y. bin:Permian 4
*Period:Early/Lower Permian - Middle Permian *Epoch:Early/Lower Rotliegendes - Middle Zechstein
Key time interval:Early/Lower Severodvinian Other zone: Deltavjatia vjatkensis
Age range of interval:264.28000 - 259.51000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Geological group:Kotelnich Formation:Urpalov Member:Vanyushonki
Stratigraphic resolution:formation
Stratigraphy comments: Kotelnich Subassemblage of the Sokolki Assemblage; the early part of the Late Tatarian (early Severodvinian) (Golubev 2000; Benton 2012, Geology).

"The geological age of the beds of the Kotelnitch locality is younger than that of the zone III (series I, acc. to Kassin --Urdzum series, acc. to A. N. Mazarovitich), and apparently they are synchronous with the lower horizons of Zone IV, or the Pareiasaurian zone, i. e., with the lower layers of the Sarmin series of Mazarovitch." (Efremov, 1940).
"The detailed geology and stratigraphy of the locality was studied by Coffa (1997 [PhD Thesis]). He recognized the Urpalov Formation at the locality as consisting of four members (Fig. 2), the oldest of which - Vanyushonkov Member (arrow in Fig. 2) composed of red calcareous clays and mudstones yields remains of pareiasaurs. At present, the Vanyushonkov Member correlates to the middle of the Urpalov Formation, Kotel'nich Group, Vishkil'skiy Horizon, Upper Tatarian Stage of the Upper Permian (Coffa 1997). The old Severo-Dvinsky Horizon is no longer in usage due to nomenclature problems." (Kordikova & Khlyupin, 2001).
Formation name, originally entered as >>Sarmin<<, which never was a lithostratigraphic term, was replaced by >>Urpalov<< (TL).
Stratigraphic age originally entered as >>Rotliegendes-Zechstein<< was refined: The Kotelnich locality is Lower Severodvinskian according to Ivakhnenko (2008). The Kotelnich Subassemblage of the Sokolki Assemblage and the Deltavjatia vjatkensis Assemblage Zone are regarded as early Upper Severodvinian by Golubev (2005, New Mexico Mus. Nat. Hist. Sci. Bull. 30).
Lithology and environment
Primary lithology:brown,red calcareous claystone
Secondary lithology: sandstone
Includes fossils?Y
Includes fossils?Y
Lithology description: Reddish-brown clay beds, red calcareous clays (Efremov, 1940). This primarily refers to the pareiasaur bearing Vanyushonkov Member of the Urpalov Formation (see Kordikova & Khlyupin, 2001). Olson (1957) and Rubidge & Sidor (2002) also mention sandstone as host rock, probably representing other fossiliferous levels than the Vanyushonkov Member.
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body,mold/impression,concretion
Size of fossils:macrofossils
Articulated whole bodies:many
Associated major elements:some
Collection methods and comments
Collection methods:survey of museum collection
Reason for describing collection:taxonomic analysis
Museum repositories:PIN
Collectors:K. Grekhov, among others Collection dates:1935, 1999
Collection method comments: The type material of Pareiasuchus vjatkensis and Anthodon rossicus was collected in 1935. The neonate Deltavjatia reported by Kordikova & Khlyupin (2001) was collected in 1999 by K. Grekhov.

KPM = Kotel'nich Paleontological Museum (= KMR in Lee, 1997, 2000)
UMZC = University Museum of Zoology, Cambridge, UK
Taxonomic list comments:"There are at least another four skulls of this animal [i.e., Deltavjatia vjatkensis]. One is in the Palaeontological Institute, Moscow. The labels have been mixed up, but the catalogue number is probably PIN 2212/3. Another is in the private collection of Mr. Terry Manning. Two more are currently undergoing preparation in the Department of Zoology, Erindale College, University of Toronto (Modesto. 1994)." (Lee, 1997, 2000).
Metadata
Also known as:Kotel'nich; Volki, Volky; Vanyushonki
Database number:28854
Authorizer:J. Alroy, J. Fröbisch, A. Dunhill, R. Butler, J. Mueller Enterer:R. Whatley, T. Liebrecht, R. Butler, B. Allen, M. Walther
Modifier:R. Butler Research group:vertebrate
Created:2003-02-25 13:52:31 Last modified:2018-06-19 17:25:03
Access level:the public Released:2003-02-25 13:52:31
Creative Commons license:CC BY
Reference information

Primary reference:
31376. A. Hartmann-Weinberg. 1937. Pareiasauriden als Leitfossilien [Pareiasaurids as guide fossils]. Problemy paleontologii 213:649-712 [J. Mueller/T. Liebrecht/T. Liebrecht]

Secondary references:

78545% 47000J. A. Efremov. 1940. Predvaritel'noye opisaniye novykh form permskoy i triasovoy fauny nazemnykh pozvonochnykh SSSR [Preliminary description of new forms from the Permian and Triassic terrestrial vertebrate fauna of the USSR]. Trudy Paleontologicheskogo Instituta Akademiy Nauk SSSR 10(2):1-140 [J. Alroy/R. Whatley/M. Carrano]
63997 V. K. Golubev. 2000. The Faunal Assemblages of Permian Terrestrial Vertebrates from Eastern Europe. Paleontological Journal 34:S211-S224 [R. Butler/R. Butler]
38222 M. F. Ivakhnenko. 1994. A new Late Permian dromasaurian (Anomodontia) from Eastern Europe. Paleontological Journal 28(1):96-103 [J. Fröbisch/M. Walther]
26871 M. F. Ivakhnenko. 1997. New Late Permian nycteroleterids from Eastern Europe. Paleontological Journal 31(5):552-558 [J. Mueller/J. Dummasch]
63003 M. F. Ivakhnenko. 2003. Eotherapsids from the East European Placket (Late Permian). Paleontological Journal 37(Suppl. 4):S339-S465 [R. Butler/R. Butler]
28657 M. F. Ivakhnenko. 2008. Podklass Parareptilia. In M. F. Ivakhnenko and E. N. Kurotchkin (eds.), Iskopaemye pozvonotchnye Rossii i sopredel'nykh stran: Iskopaemye reptilii i ptitsy, Tchast' 1 [Fossil vertebrates of Russia and adjacent countries: Fossil reptiles and birds, Part 1], GEOS, Moscow 49-85 [J. Mueller/T. Liebrecht/T. Liebrecht]
46409 M. F. Ivakhnenko. 2011. Permian and Triassic therocephals (Eutherapsida) of Eastern Europe. Paleontological Journal 45(9):981-1144 [R. Butler/R. Butler]
38005 N. N. Kalandadze and A. A. Kurkin. 2000. A new Permian dicynodont and the question of the origin of the Kannemeyeroidae. Paleontological Journal 34(6):642-649 [J. Fröbisch/M. Walther]
66024 C. F. Kammerer and V. Masyutin. 2018. Gorgonopsian therapsids (Nochnitsa gen. nov. and Viatkogorgon) from the Permian Kotelnich locality of Russia. PeerJ 6:e4954 [R. Butler/R. Butler]
66025 C. F. Kammerer and V. Masyutin. 2018. A new therocephalian (Gorynychus masyutinae gen. et sp. nov.) from the Permian Kotelnich locality, Kirov Region, Russia. PeerJ 6:e4933 [R. Butler/R. Butler]
31378 E. G. Kordikova and A. J. Khlyupin. 2001. First evidence of a neonate dentition in pareiasaurs from the Upper Permian of Russia. Acta Palaeontologica Polonica 46(4):589-594 [J. Mueller/T. Liebrecht]
29616 M. S. Y. Lee. 1997. A taxonomic revision of pareiasaurian reptiles: implications for Permian terrestrial paleoecology. Modern Geology 21:231-298 [J. Mueller/T. Liebrecht/M. Carrano]
31395 M. S. Y. Lee. 2000. The Russian pareiasaurs. In M. J. Benton, M. A. Shishkin, D. M. Unwin & E. N. Kurochkin (eds.), The Age of Dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge 71-85 [J. Mueller/T. Liebrecht/R. Butler]
5921ETE 1702E. C. Olson. 1957. Catalogue of Localities of Permian and Triassic Terrestrial Vertebrates of the Territiories of the USSR. Journal of Geology 65(2):196-226 [A. Behrensmeyer/A. Behrensmeyer/T. Liebrecht]
29069 B. S. Rubidge and C. A. Sidor. 2002. On the Cranial Morphology of the Basal Therapsids Burnetia and Proburnetia (Therapsida: Burnetiidae). Journal of Vertebrate Paleontology 22(2):257-267 [J. Mueller/T. Liebrecht]