PBDB Collection

Cambridge Greensand [non-specific locality data]: Late/Upper Albian, United Kingdom

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Reptilia

	Ornithocheirus scaphorhynchus n. sp. Seeley 1870	Seeley 1870	1 specimen
	nomen dubium belonging to Pterodactyloidea
	CAMSM B 54441, anterior portion of the rostrum

	Ornithocheirus crassidens n. sp. Seeley 1870	Seeley 1870	1 specimen
	nomen dubium belonging to Pterodactyloidea

	Pterodactylus woodwardi n. sp. Owen 1861	Owen 1861	1 specimen
	nomen dubium belonging to Pterodactyloidea
	CAMSM B 54433a, anterior portion of the rostrum

	Ornithocheirus oxyrhinus n. sp. Seeley 1870	Seeley 1870	1 specimen
	nomen dubium belonging to Pterodactyloidea
	CAMSM B 54612, anterior fragment of an upper jaw

	Ornithocheirus eurygnathus n. sp. Seeley 1870	Seeley 1870	1 specimen
	nomen dubium belonging to Pterodactyloidea
	CAMSM B54644, anterior fragment of an ?upper jaw

	Ornithocheirus dentatus n. sp. Seeley 1870	Seeley 1870	1 specimen
	nomen dubium belonging to Pterodactyloidea
	CAMSM B 54544, anterior portion of the rostrum

	Ornithocheirus enchorhynchus n. sp. Seeley 1870	Seeley 1870	1 specimen
	nomen dubium belonging to Pterodactyloidea
	CAMSM B 54444, anterior portion of the rostrum

	Ornithocheirus xyphorhynchus n. sp. Seeley 1870	Seeley 1870	1 specimen
	nomen dubium belonging to Pterodactyloidea
	anterior portion of the rostrum (collection data could not be recovered)

	Ornithocheirus tenuirostris n. sp. Seeley 1870	Seeley 1870	1 specimen
	nomen dubium belonging to Pterodactyloidea
	CAMSM B 54584, anterior portion of the rostrum

	Ornithostoma sedgwicki n. gen., n. sp. Seeley 1871	Seeley 1871	1 specimen

	Ornithocheirus brachyrhinus n. sp. Seeley 1870	Seeley 1870	1 specimen
	nomen dubium belonging to Pterodactyloidea
	CAMSM B54443, anterior portion of the rostrum
Reptilia - Anhangueridae

	Pterodactylus sedgwickii n. sp. Owen 1859		12 specimens
	recombined as Aerodraco sedgwickii
	CAMSM B54.422 (holotype), anterior rostrum

	Ornithocheirus capito n. sp. Seeley 1870	Seeley 1870	6 specimens
	recombined as Nicorhynchus capito
	CAMSM B54.625 (holotype), rostrum.

	Ornithocheirus reedi n. sp. Seeley 1870	Seeley 1870	1 specimen
	synonym of Nicorhynchus capito
Reptilia

	Pterodactylus fittoni n. sp. Owen 1859		7 specimens
	synonym of Ptenodactylus nasutus
	CAMSM B54.423 (holotype), anterior rostrum

	Ornithocheirus polyodon n. sp. Seeley 1870	Seeley 1870	1 specimen
	synonym of Ptenodactylus nasutus
	CAMSM B54440, anterior portion of the rostrum

	Ornithocheirus nasutus n. sp. (Seeley 1869)	Rodrigues and Kellner 2013	1 specimen
	original and current combination Ptenodactylus nasutus
	CAMSM B 54556 (holotype), anterior portion of the rostrum

	Pterodactylus cuvieri Bowerbank 1851	Seeley 1870	42 specimens
	recombined as Anhanguera cuvieri

	Ornithocheirus colorhinus n. sp. (Seeley 1869)	Seeley 1870	1 specimen
	original and current combination Ptenodactylus colorhinus

	Ornithocheirus denticulatus n. sp. Seeley 1870	Seeley 1870	1 specimen
	synonym of Anhanguera cuvieri
	Anterior portion of the rostrum (?CAMSM B 54794)

	Ornithocheirus sp. Seeley 1869	Unwin 2001	1 specimen
	CAMSM B54.890, possible new species

	Pterodactylus simus n. sp. Owen 1861	Owen 1861	18 specimens
	recombined as Ornithocheirus simus
	CAMSM B54.428 (holotype), rostrum. Additional rostra and mandibular symphyses; CAMSM B54552, CAMSM B54429, CAMSM B54677, MANCH L.10832, and NHMUK PV 35412

	Ornithocheirus platyrhinus n. sp. Seeley 1870	Seeley 1870	1 specimen
	synonym of Ornithocheirus simus

	Ornithocheirus platystomus n. sp. Seeley 1870	Seeley 1870
	recombined as Lonchodectes platystomus

	Ornithocheirus microdon n. sp. (Seeley 1869)	Seeley 1870	1 specimen
	original and current combination Ptenodactylus microdon
	CAMSM B54486, anterior portion of the rostrum

	Ornithocheirus oweni n. sp. Seeley 1870	Seeley 1870	1 specimen
	synonym of Ptenodactylus microdon

	Ornithocheirus huxleyi n. sp. Seeley 1870	Seeley 1870	1 specimen
	synonym of Ptenodactylus microdon

	Ornithocheirus carteri n. sp. (Seeley 1870)	Seeley 1870	1 specimen
	nomen dubium belonging to Pterodactyloidea
	CAMSM B 54437, anterior portion of the rostrum
Reptilia - Plesiosauria

	Plesiosauria indet. (de Blainville 1835)	Seeley 1869

	Plesiosaurus pachyomus n. sp. Owen 1840	Seeley 1869
	nomen vanum belonging to Plesiosauria
	Humerus, reputedly the type (p. 42). There is also a femur marked "? femur of the Pl. pachyomus type" (p. 44)

	Plesiosaurus planus n. sp. Owen 1864	Owen 1864	3 individuals
	nomen vanum belonging to Plesiosauria
	p. 41, "series 5" [possibly more than one individual], "series 6", "series 7", mostly vertebrae

	Polyptychodon sp. Owen 1841	Seeley 1869	2 individuals
	nomen vanum belonging to Plesiosauria
	Partial axial columns of two individuals, plus 103 teeth (p. 45)
Reptilia

	Ichthyosauria indet. (de Blainville 1835)	Fischer 2013	104 specimens
	Teeth NOT counted as valid specimens

	Cetarthrosaurus walkeri n. gen., n. sp. (Seeley 1869)	Fischer et al. 2014	2 specimens
	nomen dubium belonging to Ichthyosaurus
	Two propodials: CAMSM B58069, CAMSM X50170
Reptilia - Ophthalmosauridae

	Ophthalmosauridae indet. Andrews 1910	Fischer 2013	44 specimens
	Teeth NOT counted as valid specimens

	Ophthalmosaurinae indet. Baur 1887	Fischer 2013	42 specimens
	NHMUK 32406 partim (tooth); NHMUK R16 partim (tooth); NHMUK 47268 (5 teeth); CAMSM B58042 (humerus); CAMSM B58045 (humerus); CAMSM B58050 (humerus); CAMSM B58053 (humerus); CAMSM B58055 (humerus); CAMSM TN1727 partim (humerus); CAMSM TN1755 partim (2 humeri); IRSNB GS3 (humerus); LEICT G65.1991 (humerus); NHMUK R2343 (3 humeri); NHMUK R4513 (2 humeri); NHMUK 35348 (humerus); NHMUK 43989 (humerus, holotype of Brachypterygius cantabrigiensis); IRSNB GS60 (anterior accessory epipodial element).

	Platypterygiinae indet. Arkhangelsky 2001	Fischer 2013	2 specimens
	holotype of Ichthyosaurus bonneyi (CAMSM B58062, a femur); holotype of I. platymerus (CAMSM B58057, a humerus)

	Sisteronia seeleyi n. gen., n. sp. Fischer et al. 2014	Fischer et al. 2014	40 specimens
	Holotype. CAMSM B58257_67, an incomplete specimen, including partial basicranium, scapula, humerus, and 5 centra; Referred material: CAMSM B57943 (basioccipital); CAMSM B57945 (basioccipital); CAMSM B57948 (basioccipital); CAMSM B57950 (basioccipital); CAMSM B57947 (basioccipital); CAMSM B57941 (basioccipital); CAMSM B57951 (basioccipital); CAMSM B57946 (basioccipital); CAMSM B57956 (basioccipital); CAMSM B57954 (basioccipital); CAMSM B58314 (basioccipital); CAMSM TN1727 partim (basioccipital); CAMSM TN1735 partim (6 basioccipitals); CAMSM TN1739 partim (basioccipital); CAMSM TN1751 partim (6 basioccipitals); CAMSM TN1753 partim (basioccipital); IRSNB GS54 (basioccipital); IRSNB GS61 (basioccipital); LEICT G107.1991 (basioccipital); NHMUK 44159 (basioccipital); NHMUK 44159a (basioccipital); CAMSM B57908 (opisthotic); CAMSM B58077_78 (2 opisthotics); CAMSM TN1753 partim (opisthotic); NHMUK R2348 (opisthotic); IRSNB GS10 (opisthotic); CAMSM B58091 (tooth); CAMSM B58092 (tooth); CAMSM TN1716 partim (numerous teeth); CAMSM TN1778 partim (numerous teeth); CAMSM TN1779 partim (numerous teeth); CAMSM B58390 (tooth); NHMUK R1923 (tooth); IRSNB GS23 (tooth); IRSNB GS24 (tooth); IRSNB GS55 to GS58 (teeth); CAMSM TN1755 partim (humerus); CAMSM TN1757 partim (humerus)

	Platypterygius sp. von Huene 1922	Fischer et al. 2014	41 specimens
	CAMSM TN283* (rostrum and associated 112 teeth); CAMSM B42404_20* (basioccipital, centra); CAMSM B57939 (basioccipital); CAMSM B57940 (basioccipital); CAMSM B57944 (basioccipital); CAMSM B57959_60* (basioccipital, atlasaxis); CAMSM B58250_56* (quadrate, basioccipital, basisphenoid, supraoccipital, femur); CAMSM B75735 (basioccipital); CAMSM X50161 (basioccipital); CAMSM X50168 (basioccipital); CAMSM X50169 (basioccipital); CAMSM TN1729 partim (basioccipital); CAMSM TN1754 partim (basioccipital); CAMSM TN1755 partim (2 basioccipitals); NHMUK 35323 (basioccipital); several dozens of teeth, including CAMSM B57996_58009, CAMSM B58010 to B58027, CAMSM B58305_13; CAMSM B58379_87, CAMSM B76728_45, CAMSM TN1716 partim, CAMSM TN1778 partim; CAMSM TN1779 partim, NHMUK R625, NHMUK R133b partim, NHMUK R2336 partim (2 teeth), NHMUK 28110 partim, NHMUK 30253 partim, NHMUK 30254 (4 teeth), NHMUK 32406 partim, NHMUK 33242, NHMUK 35254 partim, NHMUK 35432_5, NHMUK 40358, NHMUK 41896, NHMUK 46381, NHMUK 47265_66* (teeth, centra), NHMUK 47269, IRSNB GS21, IRSNB GS25 to GS28, IRSNB GS32 to GS50, IRSNB GS53, IRSNB GS62; CAMSM B97401 partim (humerus morphotype 1); CAMSM B57987 (humerus morphotype 1); CAMSM B58043 (humerus morphotype 4); CAMSM B58048 (humerus morphotype 4); CAMSM B58056 (humerus morphotype 1); CAMSM B58057 (humerus morphotype 1; holotype of Ichthyosaurus platymerus); CAMSM B97401 partim (humerus morphotype 1); CAMSM TN1734 partim (humerus morphotype 4); CAMSM TN1751 partim (humerus morphotype 1); CAMSM TN1753 partim (one (humerus morphotype 1and one (humerus morphotype 4); CAMSM TN1757 partim (humerus morphotype 4); NHMUK R2342 partim (two humerus morphotype 4); CAMSM B58058 (femur); CAMSM B58060 (femur); CAMSM B58062 (femur; holotype of Ichthyosaurus bonneyi); CAMSM B58063_4 (femur); CAMSM B58361 (femur); CAMSM TN1749 partim (femur); CAMSM TN1748 partim (femur); CAMSM TN1757 partim (2 femora); NHMUK R23412 partim (femur); NHMUK R3510 (femur).
Reptilia - Testudines - Cheloniidae

	Chelone jessoni n. sp. Lydekker 1889	Lydekker 1889
	BMNH 35186 (holotype), mandible

	Lytoloma cantabrigiense n. sp. Lydekker 1889	Lydekker 1889
	BMNH 35178 (holotype), mandible
Reptilia

	Helochelys danubina von Meyer 1854	Joyce et al. 2011
	CAMSM B56424

	Plastremys rutteri (Andrews 1920)	Joyce 2022	50 specimens
	CAMSM B56029, a shell fragment (Figure 2g); CAMSM B56045, B56046, B56070, B56328, B56329, B56331, B56336, B56070, B56339, B56344, B56346, 11 shell fragments; CAMSM B56349, a neural (Figure 2e); CAMSM B56350–B56352, three partial neurals; CAMSM B56353–B56355, B56357–B56365, 12 shell fragments; CAMSM B56443, B56604, B56605, three plastral notch fragments; NHMUK R2251, 24 shell fragments, including one large neural (Figure 2f); NHMUK 35188, 35188a, 35188b, three shell fragments.

	Trachydermochelys phlyctaenus n. gen., n. sp. Seeley 1869	Seeley 1869
	CAMSM 56340–6, seven peripheral plates
see common names

Geography

Country:	United Kingdom	State/province:	England	County:	Cambridgeshire
Coordinates:	52.2° North, 0.1° East (view map)
Paleocoordinates:	40.6° North, 2.8° East
Basis of coordinate:	based on nearby landmark
Geographic resolution:	local area

Time

Period:	Cretaceous	Epoch:	Early/Lower Cretaceous
Stage:	Albian	10 m.y. bin:	Cretaceous 4
Key time interval:	Late/Upper Albian
Age range of interval:	106.20000 - 100.50000 m.y. ago

Stratigraphy

Geological group:	Chalk	Formation:	West Melbury Marly Chalk	Member:	Cambridge Greensand
Stratigraphic resolution:	member
Stratigraphy comments: Remanie fauna - reworked from late Albian into early Cenomanian deposits. Faunal assemblage is late Albian in age (age used here). "Although the deposit itself is generally considered to be of late Cretaceous, Cenomanian age, its macrofossil assemblage is thought to be derived largely from the underlying early Cretaceous, Albian, Gault Formation, on the basis of derived ammonites associated with the vertebrate fossils. An early Cenomanian age for deposition of the Cambridge Greensand is based on an apparently autochthonous foraminiferan assemblage, but Morter and Wood (1983) could not rule out the possibility of a very late Albian age due to the presence of the bivalve Aucinella. The age of the derived vertebrates is now considered to be no older than the Callihoplites auritus ammonite subzone and most likely no younger than upper Stoliczkaia dispar Zone, and they are thus of late Albian age." (Martill & Unwin 2011, Cretaceous Research, 10.1016/j.cretres.2011.09.003).

Lithology and environment

Primary lithology:	lag,glauconitic,micaceous,nodular,phosphatic silty marl
Includes fossils?	Y
Environment:	open shallow subtidal

Taphonomy

Modes of preservation:	body,replaced with phosphate
Degree of concentration:	-lag
Size of fossils:	macrofossils
Preservation of anatomical detail:	variable
Abundance in sediment:	common
Fragmentation:	frequent
Bioerosion:	frequent
Encrustation:	frequent
Temporal resolution:	condensed
Spatial resolution:	allochthonous

Collection methods and comments

Collection methods:	salvage,mechanical,survey of museum collection
Reason for describing collection:	taxonomic analysis
Collection method comments: CAMSM: Sedgwick Museum, Cambridge

Metadata

Database number:	94210
Authorizer:	R. Butler, V. Fischer, P. Mannion, A. Dunhill, E. Vlachos, R. Benson, P. Holroyd	Enterer:	R. Butler, P. Holroyd, E. Vlachos, R. Benson, V. Fischer, B. Allen, J. Tennant
Modifier:	M. Uhen	Research group:	vertebrate
Created:	2010-02-11 07:35:24	Last modified:	2020-10-16 14:46:23
Access level:	the public	Released:	2010-02-11 07:35:24
Creative Commons license:	CC BY

Reference information

Primary reference:

32019.

R. Owen. 1859. Supplement (No. I) to the Monograph on the Fossil Reptilia of the Cretaceous Formations. Palaeontographical Society Monographs 1-19 [R. Butler/R. Butler/M. Carrano]

Secondary references:

45995		A. O. Averianov. 2012. Ornithostoma sedgwicki - valid taxon of azhdarchoid pterosaurs. Proceedings of the Zoological Institute RAS 316(1):40-49 [P. Mannion/J. Tennant/J. Tennant]
55270	fossil record	V. Fischer. 2013. Origin, biodiversity, and extinction of Cretaceous ichthyosaurs. 1-576 [V. Fischer/V. Fischer]
49911		V. Fischer, N. Bardet, M. Guiomar and P. Godefroit. 2014. High diversity in Cretaceous ichthyosaurs from Europe prior to their extinction. PLoS ONE 9(1):e84709 [P. Mannion/J. Tennant]
32042		R. W. Hooley. 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens of the Cambridge Greensand in the Sedgwick Museum. Annals and Magazine of Natural History 8:529-557 [R. Butler/R. Butler]
82801		W. G. Joyce. 2022. A review of helochelydrid shell material from late Albian to early Cenomanian greensands of Southern England, United Kingdom. The Anatomical Record 1-13 [E. Vlachos/E. Vlachos]
39848		W. G. Joyce, S. D. Chapman, R. T. J. Moody and C. A. Walker. 2011. The skull of the solemydid turtle Helochelydra nopcsai from the Early Cretaceous of the Isle of Wight (UK) and a review of Solemydidae. Special Papers in Palaeontology 86:75-97 [R. Benson/R. Benson/D. Nicholson]
46828		R. Lydekker. 1888. Note on the classificaiton of the Ichthyopterygia (with a notice of two new species). Geological Magazine 5(7):309-314 [P. Mannion/J. Tennant]
14159	ETE	R. Lydekker. 1889. On remains of Eocene and Mesozoic Chelonia and a tooth of (?) Ornithopsis. Quarterly Journal of the Geological Society of London 45:227-246 [M. Carrano/M. Carrano/M. Uhen]
21694		S. Melmore. 1946. Catalogue of types and figured specimens in the Geological Department of the Yorkshire Museum. The North Western Naturalist 20(3/4):207-221 [G. Webster/G. Webster/M. Carrano]
32018		R. Owen. 1859. On remains of new and gigantic species of pterodactyle (Pter. fittoni and Pter. sedgwickii) from the Upper Greensand near Cambridge. Reports of the British Association for the Advancement of Science 28:98-103 [R. Butler/R. Butler]
32023		R. Owen. 1861. Monograph on the fossil Reptilia of the Cretaceous formations. Supplement no. III. Pterosauria (Pterodactylus) and Sauropterygia (Polyptychodon). Palaeontographical Society Monographs 1-25 [R. Butler/R. Butler]
44004		R. Owen. 1864. Monograph on the fossil Reptilia of the Cretaceous formations. Supplement no. IV. Sauropterygia (Plesiosaurus). Palaeontographical Society Monographs 16(70):1-18 [R. Benson/R. Benson]
47175		T. Rodrigues and A. W. A. Kellner. 2013. Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England. ZooKeys 308:1-112 [P. Mannion/J. Tennant]
32026		H. G. Seeley. 1864. On pterodactyles and on a new species of Pterodactylus machaerorhynchus. Proceedings of the Cambridge Philosophical Society 1864:228 [R. Butler/R. Butler]
32027		H. G. Seeley. 1864. On the osteology and classification of Pterodactyles, Part II, with descriptions of the new species P. Hopkinsi and P. Oweni. Proceedings of the Cambridge Philosophical Society 1864:228 [R. Butler/R. Butler]
32030		H. G. Seeley. 1865. On the pterodactyle as evidence of a new subclass of Vertebrata (Saurornia). Report of the British Association for the Advancement of Science 34:69 [R. Butler/R. Butler]
14179	ETE	H. G. Seeley. 1869. Index to the Fossil Remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata, Arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co, Cambridge 1-143 [M. Carrano/M. Carrano/M. Uhen]
32034		H. G. Seeley. 1870. The Ornithosauria: an elementary study of the bones of pterodactyles, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge, Deighton, Bell & Co, Cambridge [R. Butler/R. Butler]
32038		H. G. Seeley. 1871. Additional evidence on the structure of the head in ornithosaurs from the Cambridge Upper Greensand; being a supplement to "The Ornithosauria". Annals and Magazine of Natural History 7:20-36 [R. Butler/R. Butler]
74115		H. G. Seeley. 1873. On Cetarthrosaurus walkeri (Seeley), an ichthyosaurian from the Cambridge Upper Greensand. Quarterly Journal of the Geological Society of Lodon 29:505-507 [M. Uhen/M. Uhen]
31160		D. M. Unwin. 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen as dem Museum für Naturkunde, Berlin, Geowissenschaftliche Reihe 4:189-222 [R. Butler/R. Butler]