PBDB Collection

Warner's Bay, Belmont (AMF coll): Changhsingian, Australia
collected by M.S. Stanley, O. Le M. Knight

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Insecta - Permosialidae

	Permosialis belmontensis n. sp. Riek 1968	Riek 1968	1 specimen
	(1 measurement)
Insecta - Lophioneurida - Zoropsocidae

	Zoropsocus stanleyi n. sp. Davis 1942		2 specimens
	(2 measurements)
Insecta - Lophioneurida - Lophioneuridae

	Lophiocypha maxima n. sp. Davis 1942		1 specimen
	(1 measurement)

	Lophiocypha stanleyi Tillyard 1935		5 specimens
	(5 measurements)

	Lophiocypha permiana Tillyard 1935		1 specimen
	(1 measurement)

	Austrocypha abrupta Tillyard 1935		1 specimen
	(1 measurement)
Insecta - Hemiptera - Stenoviciidae

	Stanleyana pulchra n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
	(1 measurement)

	Permovicia obscura n. gen., n. sp. Evans 1943	Evans 1943	1 specimen

	Permagra distincta n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
	AM.F39866 (coll. O.M. Knight, 1942) (2 measurements)

	Stenovicia angustata n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
	(1 measurement)

	Palaeovicia incerta n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
	(1 measurement)
Insecta - Hemiptera - Scytinopteridae

	Eochiliocycla angusta n. gen., n. sp. Davis 1942		1 specimen
	(1 measurement)
Insecta - Hemiptera

	Permocapitus globulus n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
Insecta - Hemiptera - Archescytinidae

	Protopincombea obscura n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
	(2 measurements)

	Bekkerscytina primitiva n. gen., n. sp. Evans 1958	Evans 1958	1 specimen
	(1 measurement)

	Austroscytina imperfecta n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
	(1 measurement)

	Permocephalus knighti n. gen., n. sp. Evans 1943	Evans 1943	4 specimens
	AM.F39865; F39944, 39945, 39967 in Evans 1943b (1 measurement)

	Eoscytina migdisovae n. gen., n. sp. Evans 1958	Evans 1958	2 specimens
	(1 measurement)

	Eoscytina incompleta n. sp. Evans 1958	Evans 1958	1 specimen
Insecta - Hemiptera - Prosbolidae

	Homaloscytina plana Tillyard 1926	Evans 1943	8 specimens
	(7 measurements)

	Permobrachus magnus n. sp. Evans 1943	Evans 1943	1 specimen
	(1 measurement)

	Anomaloscytina metapteryx n. gen., n. sp. Davis 1942		1 specimen
	(1 measurement)

	Anomaloscytina incompleta n. sp. Evans 1943	Evans 1943	1 specimen

	Orthoscytina spp.	Evans 1943	10 specimens
	(9 measurements)
Insecta - Hemiptera - Dunstaniidae

	Austroprosbole maculata n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
	(1 measurement)
Insecta - Hemiptera - Dysmorphoptilidae

	Belmontocarta perfecta n. gen., n. sp. Evans 1958	Evans 1958	1 specimen
	(1 measurement)
Insecta - Hemiptera

	Tychticoloides belmontensis n. gen., n. sp. Evans 1963	Evans 1963	1 specimen
Insecta - Hemiptera - Pincombeidae

	Eupincombea postica n. gen., n. sp. Davis 1942		1 specimen
	(1 measurement)

	Protopsyllops minuta n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
	recombined as Pincombea minuta
	(1 measurement)
Insecta - Hemiptera - Protopsyllidiidae

	Tripsyllidium wadei n. gen., n. sp. Evans 1956	Evans 1956	1 specimen
	AM F30980 (1 measurement)

	Protopsyllidium sinuatum n. sp. Davis 1942		3 specimens
	Evans specimens are P. setosum, part of Davis specimens also re-ID (9 measurements)

	Protopsyllidium australe Tillyard 1926	Evans 1943	1 specimen
	(1 measurement)

	Protopsyllidium setosum n. sp. Becker-Migdisova 1985	Becker-Migdisova 1985	8 specimens
	BMNH In. 45252, 45847, 45237, 45208, AM F40067, 39997, 39981 (3 measurements)
Insecta - Hemiptera - Permopsyllidiidae

	Psocopsyllidium media n. gen., n. sp. Davis 1942		1 specimen
	(1 measurement)

	Permopsyllidium mitchelli Tillyard 1926	Evans 1943	1 specimen
	(2 measurements)
	= Permopsyllidium evansi Becker-Migdisova 1985	Becker-Migdisova 1985
	AM F40040

	Permopsyllidops stanleyi n. gen., n. sp. Davis 1942		1 specimen
	recombined as Permopsyllidium stanleyi
	(1 measurement)

	Permotheella scytinopteroides n. gen., n. sp. Davis 1942		3 specimens
	recombined as Permopsyllidium scytinopteroides
	1 in Evans, 1943 (3 measurements)

	Permopsyllidium affine Tillyard 1926	Becker-Migdisova 1985	2 specimens
	AM F39901, 40006 (P. mitchelli in Evans) (2 measurements)

	Clavopsyllidium minutum n. gen., n. sp. Davis 1942		1 specimen
	(1 measurement)

	Belpsylla reticulata n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
	(1 measurement)

	Permopsyllidium affine Tillyard 1926	Evans 1943	1 specimen
	(1 measurement)
	= Permopsyllidium permianum n. sp. Becker-Migdisova 1985	Becker-Migdisova 1985
	recombined as Permelania permiana

	Psocoscytina bifida n. gen., n. sp. Davis 1942		1 specimen
	(1 measurement)

	Eopsyllidium delicatulum n. gen., n. sp. Davis 1942		1 specimen
	(1 measurement)

	Psyllidella magna n. gen., n. sp. Evans 1943	Evans 1943	1 specimen
	(1 measurement)
Insecta - Miomoptera - Palaeomanteidae

	Belmomantis azari n. gen., n. sp. Prokop et al. 2017	Prokop et al. 2017	1 specimen
	AM F.142068/142069
Insecta - Permithonidae

	Permithone neoxenus n. sp. Riek 1953	Riek 1953	1 specimen
	recombined as Osmythone neoxenus
	(1 measurement)

	Permorapisma biserialis Tillyard 1926	Riek 1953	10 specimens

	Permithone belmontensis Tillyard 1922	Riek 1953	2 specimens

	Permithone oliarcoides Tillyard 1926	Riek 1953	2 specimens
Insecta - Neuroptera - Archeosmylidae

	Archeosmylus pectinatus n. gen., n. sp. Riek 1953	Riek 1953	3 specimens
	(1 measurement)
Insecta - Coleoptera - Ponomarenkiumidae

	Ponomarenkia belmonthensis n. gen., n. sp. Yan et al. 2017	Yan et al. 2017	2 specimens
	recombined as Ponomarenkium belmonthense
	AM F40278, 41618
Insecta - Trichoptera - Cladochoristidae

	Cladochorista belmontensis Tillyard 1926	Riek 1953	10 specimens
Insecta - Protomeroptera - Protomeropidae

	Permomerope nanus n. sp. Riek 1953	Riek 1953	1 specimen
	F40233 (1 measurement)

	Permomerope australis Tillyard 1926	Riek 1953	3 specimens
	F40108, F41239/41274, F43119/43120
Insecta - Mecoptera - Belmontiidae

	Belmontia mitchelli Tillyard 1919	Riek 1953	1 specimen
	F40970 (1 measurement)

	Parabelmontia permiana Tillyard 1922	Riek 1953	1 specimen
	synonym of Belmontia mitchelli
	F41188
Insecta - Mecoptera - Permochoristidae

	Mesochorista australica (Tillyard 1918)	Riek 1953	140 specimens

	Mesochorista phipa n. sp. Riek 1953	Riek 1953	1 specimen
	F40125/40180 (1 measurement)

	Neopetromantis australis n. gen., n. sp. Riek 1953	Riek 1953	4 specimens
	F40211, F40173, F41120, F43122

	Xenochorista sobrina n. sp. Riek 1953	Riek 1953	1 specimen
	F43123 (1 measurement)

	Xenochorista splendida n. gen., n. sp. Riek 1953	Riek 1953	1 specimen
	F40157 (1 measurement)

	Nannochoristella reducta n. gen., n. sp. Riek 1953	Riek 1953	1 specimen
	(1 measurement)

	Neochoristella optata n. gen., n. sp. Riek 1953	Riek 1953	1 specimen
	(1 measurement)

	Phipoides elegans n. gen., n. sp. Riek 1953	Riek 1953	2 specimens
	F40144, F40117 (1 measurement)

	Agetochoristella adscita n. gen., n. sp. Riek 1953	Riek 1953	9 specimens
	recombined as Agetopanorpa adscita
	F43656, F39890, F46233, F40159, F40183, F40185, F43124, F26975, F41018 (1 measurement)

	Neoageta elongata n. gen., n. sp. Riek 1953	Riek 1953	1 specimen
	recombined as Agetopanorpa elongata
	F43969 (1 measurement)

	Mesochorista jucunda (Tillyard 1926)	Riek 1953	10 specimens
	recombined as Kazakhopanorpa jucunda

	Mesochorista dubia n. sp. Riek 1953	Riek 1953	2 specimens
	recombined as Kazakhopanorpa dubia
	F41287, F42108/9 (1 measurement)
Insecta - Mecoptera - Parachoristidae

	Parachorista splendida Tillyard 1926	Riek 1953	2 specimens
	F41023, F41094

	Parachorista pincombeae Tillyard 1926	Riek 1953	1 specimen
	F43110
Insecta - Mecoptera - Mesopsychidae

	Prochoristella anagaura n. sp. Riek 1953	Riek 1953	11 specimens
	(1 measurement)

	Prochoristella pusilla n. sp. Riek 1953	Riek 1953	4 specimens
	(1 measurement)

	Prochoristella exilis n. sp. Riek 1953	Riek 1953	3 specimens
	(1 measurement)

	Prochoristella megaloprepia n. gen., n. sp. Riek 1953	Riek 1953	9 specimens
	F40123, F40766, F41060, F40127, F40214, F41594, F41113, F40129, F40081 (1 measurement)

	Prochoristella concinna n. sp. Riek 1953	Riek 1953	1 specimen
	(1 measurement)

	Prochoristella belli (Tillyard 1926)	Riek 1953	4 specimens

	Mesopanorpodes belmontensis n. sp. Riek 1953	Riek 1953	6 specimens
	recombined as Permopsyche belmontensis
	F40166, F41237, F40962/1, F43480, F42499, F42498 (1 measurement)

	Mesopanorpodes robustus n. sp. Riek 1953	Riek 1953	1 specimen
	recombined as Permopsyche robustus
	F40787/8 (1 measurement)
Insecta - Mecoptera - Robinjohniidae

	Protodiptera indet. Tillyard 1937	Tillyard 1937	1 specimen
	AM F52312 (collected by Tillyard) (1 measurement)
	= Robinjohnia tillyardi n. gen., n. sp. Martynova 1948	Martynova 1948
Insecta - Mecoptera - Permotanyderidae

	Permotanyderus ableptus n. gen., n. sp. Riek 1953	Riek 1953	4 specimens
	(1 measurement)

	Choristotanyderus nanus n. gen., n. sp. Riek 1953	Riek 1953	1 specimen
	recombined as Permotanyderus nanus
	(1 measurement)
Insecta - Glosselytrodea - Jurinidae

	Permoberothella perplexa n. gen., n. sp. Riek 1953	Riek 1953	3 specimens
	(1 measurement)
see common names

Geography

Country:	Australia	State/province:	New South Wales
Coordinates:	33.0° South, 151.6° East (view map)
Paleocoordinates:	66.5° South, 144.3° East
Basis of coordinate:	estimated from map
Geographic resolution:	outcrop

Time

Period:	Permian	Epoch:	Lopingian
Stage:	Changhsingian	10 m.y. bin:	Permian 4
Key time interval:	Changhsingian
Age range of interval:	254.14000 - 251.90200 m.y. ago

Stratigraphy

Geological group:	Newcastle Coal Measures	Formation:	Croudace Bay	Member:	Belmont Conglomerate
Stratigraphic resolution:	bed
Stratigraphy comments: From horizon 70 feet below base of Fassifern Coal, according to Knight (1950). Fassifern coal dated at 253.38 +/- 0.08 Ma and the underlying Upper Pilot A seam at 253.55 +/- 0.06 Ma by Metcalf et al. (2015).

Lithology and environment

Primary lithology:	fine,tuffaceous,gray lithified chert
Lithology description: hard, fine-grained tuffaceous chert
Environment:	pond	Tectonic setting:	foreland basin
Geology comments: Switch to contractional deformation in Hunter-Bowen orogeny occurred c. 265 Ma; Sydney basin transitioned into foreland basin

Taphonomy

Modes of preservation:	mold/impression
Size of fossils:	macrofossils,mesofossils

Collection methods and comments

Reason for describing collection:	taxonomic analysis
Collectors:	M.S. Stanley, O. Le M. Knight
Collection method comments: Collected by Malcolm Stanley and types and other specimens donated to the Australian Museum. Much of material in Riek (1953) was collected by O. Le M. Knight from Warner's Bay-Belmont area.

Metadata

Database number:	122433
Authorizer:	M. Clapham	Enterer:	M. Clapham
Modifier:	M. Clapham	Research group:	paleoentomology
Created:	2011-12-27 12:00:09	Last modified:	2017-05-23 16:34:15
Access level:	the public	Released:	2011-12-27 12:00:09
Creative Commons license:	CC BY

Reference information

Primary reference:

39006.

C. Davis. 1942. Hemiptera and Copeognatha from the Upper Permian of New South Wales. Proceedings of the Linnean Society of New South Wales 67:111-122 [M. Clapham/M. Clapham]

Secondary references:

41079		E. E. Becker-Migdisova. 1985. Iskopaemye Nasekomye Psillomorfy. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 206:1-92 [M. Clapham/M. Clapham/M. Clapham]
50983		J. W. Evans. 1943. Two interesting Upper Permian Homoptera from New South Wales. Transactions of the Royal Society of South Australia 67:7-9 [M. Clapham/M. Clapham]
39008		J. W. Evans. 1943. Upper Permian Homoptera from New South Wales. Records of the Australian Museum 21(4):180-198 [M. Clapham/M. Clapham]
49517		J. W. Evans. 1956. Palaeozoic and Mesozoic Hemiptera (Insecta). Australian Journal of Zoology 4:165-258 [M. Clapham/M. Clapham]
39035		J. W. Evans. 1958. New Upper Permian Homoptera from the Belmont beds (Homoptera: Insecta). Records of the Australian Museum 24(9):109-114 [M. Clapham/M. Clapham]
39038		J. W. Evans. 1963. The systematic position of the Ipsviciidae (Upper Triassic Hemiptera) and some new Upper Permian and Middle Triassic Hemiptera from Australia (Insecta). Journal of the Entomological Society of Queensland 2:17-23 [M. Clapham/M. Clapham]
50889		O. M. Martynova. 1948. Materialy po evolyutsii Mecoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 14:1-76 [M. Clapham/M. Clapham/M. Clapham]
63134		J. Prokop, M. Pecharová, R. Garrouste, R. Beattie, I. C. Chintauan-Marquier and A. Nel. 2017. Redefining the extinct orders Miomoptera and Hypoperlida as stem acercarian insects. BMC Evolutionary Biology 17(205):1-20 [M. Clapham/M. Clapham]
36534		E. F. Riek. 1953. Fossil mecopteroid insects from the Upper Permian of New South Wales. Records of the Australian Museum 23(2):55-87 [M. Clapham/M. Clapham]
40249		E. F. Riek. 1968. Robinjohnia tillyardi Martynova, a mecopteron from the Upper Permian of Belmont, New South Wales. Records of the Australian Museum 27(14):299-302 [M. Clapham/J. Karr]
39034		E. F. Riek. 1968. Undescribed fossil insects from the Upper Permian of Belmont, New South Wales (with an appendix listing the described species). Records of the Australian Museum 27:303-310 [M. Clapham/M. Clapham]
40756		R. J. Tillyard. 1937. The ancestors of the Diptera. Nature 139:66-67 [M. Clapham/M. Clapham]
64814		E. V. Yan, J. F. Lawrence, R. Beattie and R. G. Beutel. 2017. At the dawn of the great rise: †Ponomarenkia belmonthensis (Insecta: Coleoptera), a remarkable new Late Permian beetle from the Southern Hemisphere. Journal of Systematic Palaeontology 16:611-619 [M. Clapham/M. Clapham]