Denmark Hill Insect Bed (Dunstan 1915-1916 collection): Norian, Australia
collected by Dunstan & Wilcox 1915-1916

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Eoblattida - Mesorthopteridae
Mesorthopteron locustoides Tillyard 1916
Tillyard 1922 10 specimens
GSQI258b, 72a-b, 75, 78a-b, 123, 224, 234, 241b; AM F39242, F39235 (from one of Dunstan's collections according to Riek, 1956) (1 measurement)
Insecta - Mecoptera
Mecoptera indet. Packard 1886
Tillyard 1919 1 individual
GSQI168
Insecta - Mecoptera - Mesopsychidae
Mesopsyche triareolata n. gen., n. sp. Tillyard 1917
1 individual
Dunstan 110 (AM Sydney) (1 measurement)
Aristopsyche superba n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
synonym of Mesopsyche triareolata
Dunstan 148 (AM Sydney) (1 measurement)
Neuropsyche elongata n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
synonym of Mesopsyche triareolata
Dunstan 228 (AM Sydney) (1 measurement)
Triassopsyche dunstani n. gen., n. sp. Tillyard 1917
1 individual
synonym of Mesopsyche triareolata
Dunstan 128a (AM Sydney) (1 measurement)
Insecta - Mecoptera - Orthophlebiidae
Orthophlebidae indet. Handlirsch 1906
Riek 1956 1 specimen
corrected as Orthophlebiidae
AM F39232 (from an unspecified Dunstan collection)
Insecta - Stereochoristidae
Stereochorista frustrata n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI218 (1 measurement)
Insecta - Neuroptera - Archeosmylidae
Archeosmylus sp. Riek 1953
Riek 1956 1 specimen
AM F39249 (from an unspecified Dunstan collection)
Insecta - Neuroptera - Osmylopsychopsidae
Archepsychops triassica n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
corrected as Archepsychops triassicus
GSQI137 (1 measurement)
Insecta - Neuroptera - Osmylopsychopidae
Osmylopsychops spillerae n. gen., n. sp. Tillyard 1923
Tillyard 1923 2 individuals
GSQI314, 283 (1 measurement)
Insecta - Neuroptera - Psychopsidae
Triassopsychops superba n. gen., n. sp. Tillyard 1922
Tillyard 1922 1 individual
corrected as Triassopsychops superbus
GSQI284 (1 measurement)
Insecta - Neuroptera
Protopsychopsis venosa n. gen., n. sp. Tillyard 1917
1 individual
Coll. QGS 160a (1 measurement)
Insecta - Coleoptera
Leioodes planum n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 348 (1 measurement)
Leioodes pygmaeum n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 287 (1 measurement)
Apheloodes rugosum n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 236 (1 measurement)
Apheloodes obliquum n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 143 (1 measurement)
Insecta - Coleoptera - Permosynidae
Platycrossos subtumidus n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 171 (1 measurement)
Platycrossos ligulatus n. sp. Dunstan 1923
Dunstan 1923 2 specimens
GSQ 268, 118 (1 measurement)
Insecta - Coleoptera
Polysitum punctatus n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 153 (1 measurement)
Polysitum minutus n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 336 (1 measurement)
Insecta - Coleoptera - Elateridae
Elaterites transversus n. sp. Dunstan 1923
Dunstan 1923 1 specimen
recombined as Elateridium transversum
GSQ 159 (1 measurement)
Elaterites subulatus n. sp. Dunstan 1923
Dunstan 1923 1 specimen
recombined as Elateridium subulatum
GSQ 263 (1 measurement)
Elaterium bipunctatum n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 292 (1 measurement)
Insecta - Coleoptera - Dermestidae
Tryoniopsis punctata n. gen., n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 250 (1 measurement)
Tryoniopsis granulata n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 161 (1 measurement)
Insecta - Coleoptera - Ademosynidae
Ademosyne lata n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 132 (1 measurement)
Ademosyne brevis n. sp. Dunstan 1923
Dunstan 1923 2 specimens
GSQ 339 (1 measurement)
Ademosyne vittamargina n. sp. Dunstan 1923
Dunstan 1923 2 specimens
GSQ 341, 293 (1 measurement)
Ademosyne curvata n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 274 (1 measurement)
Ademosyne olliffi (Handlirsch 1906)
Dunstan 1923 1 specimen
GSQ 115
Ademosyne adunca n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 194 (1 measurement)
Ademosyne ramocostata n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 225 (1 measurement)
Ademosyne parva n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 312 (1 measurement)
Ademosyne intermedia n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 233 (1 measurement)
Ademosyne rugulosa n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 260 (1 measurement)
Grammositum bilineatus n. gen., n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 136 (1 measurement)
Insecta - Coleoptera - Tricoleidae
Willcoxia magnopunctata n. gen., n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 261 (1 measurement)
Insecta - Coleoptera
Tillyardiopsis tuberculata n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 133 (1 measurement)
Tillyardiopsis granulata n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 289 (1 measurement)
Insecta - Coleoptera - Schizocoleidae
Shepherdia quadrivittata n. gen., n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 130 (1 measurement)
Lobites granulatus n. sp. Dunstan 1923
Dunstan 1923 1 specimen
recombined as Mesolobites granulatus
GSQ 183 (1 measurement)
Lobites tuberculatus n. sp. Dunstan 1923
Dunstan 1923 1 specimen
recombined as Mesolobites tuberculatus
GSQ 342 (1 measurement)
Mesothoris tenuiclathrata n. sp. Dunstan 1923
Dunstan 1923 1 specimen
recombined as Mesolobites tenuiclathrata
GSQ 313 (1 measurement)
Lobites trivittatus n. sp. Dunstan 1923
Dunstan 1923 1 specimen
recombined as Mesolobites trivittatus
GSQ 164 (1 measurement)
Mesothoris quadripartita n. sp. Dunstan 1923
Dunstan 1923 1 specimen
recombined as Mesolobites quadripartita
GSQ 349 (1 measurement)
Reeveana minor n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 297 (1 measurement)
Reeveana intermedia n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 201 (1 measurement)
Reeveana major n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 251 (1 measurement)
Insecta - Coleoptera - Trachypachidae
Ademosynoides magnifica n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 199 (1 measurement)
Ademosynoides alternata n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 149 (1 measurement)
Ademosynoides abnormis n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 243 (1 measurement)
Insecta - Coleoptera - Taldycupedidae
Elaterium punctomarginum n. sp. Dunstan 1923
Dunstan 1923 1 specimen
recombined as Mesothoris punctomarginum
GSQ 200 (1 measurement)
Simmondsia subpiriformis n. sp. Dunstan 1923
Dunstan 1923 2 specimens
GSQ 135, 276 (1 measurement)
Simmondsia cylindrica n. sp. Dunstan 1923
Dunstan 1923 1 specimen
GSQ 87 (1 measurement)
Insecta - Archipanorpidae
Archipanorpa magnifica n. gen., n. sp. Tillyard 1917
2 specimens
GSQI106a, 120a (2 measurements)
Insecta - Plecoptera - Euxenoperlidae
Stenoperlidium triassicum Riek 1956
Riek 1956 1 specimen
recombined as Gondwanoperlidium triassicum
AM F39256 (from an unspecified Dunstan collection)
Insecta - Triassoblattidae
Triassoblatta typica n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI180 (1 measurement)
Triassoblatta insignita n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI124 (1 measurement)
? Triassoblatta intermedia n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI216 (1 measurement)
Insecta - Blattodea
Blattodea indet. (Latreille 1810)
Tillyard 1919 2 individuals
corrected as Blattaria
GSQI121, 152
Insecta - Blattodea - Mesoblattinidae
Austroblattula ipsviciensis n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI105 (1 measurement)
Insecta - Orthoptera - Proparagryllacrididae
Proparagryllacris crassifemur n. gen., n. sp. Riek 1956
Riek 1956 1 specimen
AM F39251 (from an unspecified Dunstan collection)
Insecta - Orthoptera - Triassomanteidae
Triassomantis pygmaeus n. gen., n. sp. Tillyard 1922
Tillyard 1922 1 individual
GSQI86 (1 measurement)
Insecta - Orthoptera - Locustavidae
Triassolocusta leptoptera n. gen., n. sp. Tillyard 1922
Tillyard 1922 1 individual
GSQI99 (1 measurement)
Insecta - Aeroplanidae
Aeroplana mirabilis n. gen., n. sp. Tillyard 1918
Tillyard 1918 1 individual
GSQI126 (2 measurements)
Insecta - Hemiptera
Homoptera indet. Leach 1815
Tillyard 1919 5 specimens
Insecta - Hemiptera - Mesogereonidae
Mesogereon compressum n. sp. Tillyard 1921
Tillyard 1921 1 individual
GSQI144 (1 measurement)
Mesogereon superbum n. sp. Tillyard 1921
Tillyard 1921 1 individual
GSQI169 (1 measurement)
Mesogereon shepherdi n. sp. Tillyard 1921
Tillyard 1921 1 individual
GSQI97 (1 measurement)
Mesogereon affine n. sp. Tillyard 1921
Tillyard 1921 2 specimens
GSQI206, GSQI207 (2 measurements)
Insecta - Hemiptera - Dunstaniidae
Dunstaniidae indet. Tillyard 1916
Tillyard 1922 1 individual
GSQI119
Paradunstania affinis n. gen., n. sp. Tillyard 1918
Tillyard 1918 1 individual
GSQI147 (1 measurement)
Dunstaniopsis triassica n. gen., n. sp. Tillyard 1918
Tillyard 1918 1 individual
GSQI107 (1 measurement)
Insecta - Hemiptera - Dysmorphoptilidae
Mesocixiodes orthoclada n. sp. Tillyard 1922
Tillyard 1922 4 individuals
recombined as Mesonirvana orthoclada
GSQI318; GSQ 91, 193, 324 in Lambkin, 2016 (1 measurement)
Mesocixius triassicus n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI215 (1 measurement)
Triassocixius australicus n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI267a (1 measurement)
Insecta - Hemiptera - Archijassidae
Triassojassus proavitus n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
synonym of Mesojassus ipsviciensis
GSQI191a (1 measurement)
Insecta - Hemiptera - Hylicellidae
Mesocixiodes termioneura n. gen., n. sp. Tillyard 1922
Tillyard 1922 1 individual
GSQI88 (1 measurement)
Mesocixiodes brachyclada n. sp. Tillyard 1922
Tillyard 1922 1 individual
synonym of Mesocixiodes termioneura
GSQI325
Insecta - Hemiptera - Chiliocyclidae
Chiliocycla scolopoides n. gen., n. sp. Tillyard 1919
Tillyard 1919 2 individuals
GSQI158a, GSQ327
Insecta - Hemiptera - Tettigarctidae
Mesodiphthera grandis n. gen., n. sp. Tillyard 1919
Tillyard 1919 11 individuals
GSQI213, 192 (1 measurement)
Mesodiphthera dunstani n. sp. Tillyard 1922
Tillyard 1922 1 individual
recombined as Tardilly dunstani
(1 measurement)
Mesodiphthera prosboloides n. sp. Tillyard 1922
Tillyard 1922 1 individual
recombined as Tardilly prosboloides
GSQI89 (1 measurement)
Insecta - Hemiptera - Scytinopteridae
Mesoscytina australis n. gen., n. sp. Tillyard 1919
Tillyard 1919 2 individuals
GSQI112a; GSQ 320 in Lambkin (2016) presumably from this collection (1 measurement)
Triassoscarta subcostalis n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
synonym of Mesoscytina australis
GSQI116a (1 measurement)
Mesoscytina affinis n. sp. Tillyard 1919
Tillyard 1919 1 individual
synonym of Mesoscytina australis
GSQI235 (1 measurement)
Eurymelidium australe n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI248a (1 measurement)
Apheloscyta mesocampta n. gen., n. sp. Tillyard 1922
Tillyard 1922
(1 measurement)
Insecta - Hemiptera - Ipsviciidae
Ipsviciopsis elegans n. gen., n. sp. Tillyard 1922
Tillyard 1922 2 individuals
GSQI178, 278 (1 measurement)
Ipsviciopsis magna n. sp. Tillyard 1922
Tillyard 1922 1 individual
synonym of Ipsviciopsis elegans
GSQI93 (1 measurement)
Ipsvicia jonesi n. gen., n. sp. Tillyard 1919
Tillyard 1919 3 individuals
GSQI122a, 285, 226 (2 measurements)
Ipsvicia maculata n. sp. Tillyard 1919
Tillyard 1919 1 individual
synonym of Ipsvicia jonesi
GSQI208 (1 measurement)
Ipsvicia acutipennis n. sp. Tillyard 1919
Tillyard 1919 1 individual
synonym of Ipsvicia jonesi
GSQI204 (1 measurement)
Insecta - Hemiptera - Triassocoridae
? Triassocoris sp. Tillyard 1922
Tillyard 1922 1 individual
GSQI184
Triassocoris ovalis n. sp. Tillyard 1923
Tillyard 1923 1 individual
GSQI129 (1 measurement)
Triassocoris scutulum n. sp. Tillyard 1922
Tillyard 1922 2 individuals
GSQI134, 179 (1 measurement)
Triassocoris myersi n. gen., n. sp. Tillyard 1922
Tillyard 1922 3 specimens
GSQI140, 167, 150 (2 measurements)
Triassocoris grandis n. sp. Tillyard 1923
Tillyard 1923 1 individual
GSQI196 (1 measurement)
Insecta - Hemiptera - Progonocimicidae
Pseudipsvicia ala n. gen., n. sp. Handlirsch 1939
Handlirsch 1939 1 specimen
GSQ 285a (hindwing of Ipsvicia jonesi in Tillyard, 1923, fig 90) (1 measurement)
Insecta - Glosselytrodea - Polycytellidae
Polycytella triassica n. gen., n. sp. Tillyard 1922
Tillyard 1922 2 individuals
GSQI81, GSQI154 (1 measurement)
Insecta - Blattaria - Caloblattinidae
Samaroblatta intercalata n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI262
Samaroblatta triassica n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI156 (1 measurement)
Samaroblatta jonesi n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI157 (1 measurement)
Samaroblatta blabelloides n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI131b (1 measurement)
Samaroblatta reticulata n. gen., n. sp. Tillyard 1919
Tillyard 1919 1 individual
GSQI155a (1 measurement)
Insecta
Perissophlebia multiseriata n. gen., n. sp. Tillyard 1918
Tillyard 1918 1 individual
GSQI205
Insecta - Odonata
Triassophlebia stigmatica n. gen., n. sp. Tillyard 1922
Tillyard 1922 1 individual
GSQI82 (1 measurement)
Insecta - Odonata - Triassolestidae
Triassolestes epiophlebioides n. gen., n. sp. Tillyard 1918
Tillyard 1918 1 individual
GSQI203 (1 measurement)
Insecta - Mesophlebiidae
Mesophlebia antinodalis Tillyard 1916
Tillyard 1922 1 individual
GSQI127 (now AM F39270 / NHMUK In.33397) (1 measurement)
    = Mesophlebia tillyardi n. sp. Handlirsch 1939
Handlirsch 1939
synonym of Mesophlebia antinodalis
Insecta - Protomyrmeleontidae
Triassagrion australiense n. gen., n. sp. Tillyard 1922
Tillyard 1922 1 individual
GSQI290 (1 measurement)
see common names

Geography
Country:Australia State/province:Queensland
Coordinates: 27.6° South, 152.8° East (view map)
Paleocoordinates:58.6° South, 101.0° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Triassic Epoch:Late/Upper Triassic
Stage:Norian 10 m.y. bin:Triassic 4
Key time interval:Norian
Age range of interval:227.00000 - 208.50000 m.y. ago
Stratigraphy
Geological group:Ipswich Coal Measures Formation:Blackstone
Stratigraphic resolution:bed
Stratigraphy comments: Insect bed is about 50 feet above the Bluff coal seam and 50 feet below the Aberdare coal seam. Fossil bed is 15 cm thick. Purdy & Cranfield (2013) reported an unpublished SHRIMP date of 226±2 Ma for the Brisbane Tuff. Therefore, the Blackstone Formation may be considered younger than 226 Ma in absolute age. Thus, the Ipswich Coal Measures above the Brisbane Tuff, which is to say, the succession from the Mount Crosby Formation to the top of the Blackstone Formation must now be considered Norian.
Lithology and environment
Primary lithology: lithified sandy siltstone
Includes fossils?Y
Lithology description: "The fossil insect bed of finely arenaceous shale. Near the outcrop the bed is of a brownish grey colour with iron-stained joints and planes." Dunstan, 1916 Fresher rock is a greenish grey colour.
Environment:pond Tectonic setting:rift
Geology comments: Interbedded sandstones and mudstones accumulated on levees bordering channels and graded laterally into floodplains where carbonaceous mudstones accumulated with thin crevasse splay sandstone beds.
Ipswich basin; transtensional rift in intermontane setting. Basin geometry affected by strike-slip movement along West Ipswich fault. Ipswich coal measures deposited near end of rifting phase.
Taphonomy
Modes of preservation:mold/impression,adpression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:survey of museum collection
Collection size:200
Reason for describing collection:taxonomic analysis
Collectors:Dunstan & Wilcox Collection dates:1915-1916
Metadata
Database number:111558
Authorizer:M. Clapham Enterer:J. Karr, M. Clapham
Modifier:M. Clapham Research group:paleoentomology
Created:2011-06-25 16:45:14 Last modified:2017-05-23 16:34:15
Access level:the public Released:2011-06-25 16:45:14
Creative Commons license:CC BY
Reference information

Primary reference:

36514. R. J. Tillyard. 1917. Mesozoic Insects of Queensland No. 1 Planipennia, Trichoptera, and the new order Protomecoptera. The Proceedings of the Linnean Society of New South Wales 42(165):175-200 [M. Clapham/J. Karr]

Secondary references:

49881 B. Dunstan. 1923. Mesozoic insects of Queensland Part I - introduction and Coleoptera. Queensland Geological Survey Publication 273:1-89 [M. Clapham/M. Clapham]
37201 A. Handlirsch. 1939. Neue Untersuchungen über die fossilen Insekten mit Ergänzungen und Nachträgen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II Teil. Annalen des Naturhistorischen Museums in Wien 49:1-240 [M. Clapham/J. Karr/M. Clapham]
59991 K. J. Lambkin. 2016. Revision of the Dysmorphoptilidae (Hemiptera: Cicadomorpha: Prosboloidea) of the Queensland Triassic - Part 2. Zootaxa 4092:207-218 [M. Clapham/M. Clapham/M. Uhen]
49543 E. F. Riek. 1956. A re-examination of the mecopteroid and orthopteroid fossils (Insecta) from the Triassic beds at Denmark Hill, Queensland, with descriptions of further specimens. Australian Journal of Zoology 4:98-110 [M. Clapham/M. Clapham/M. Clapham]
36561 R. J. Tillyard. 1918. Mesozoic Insects of Queensland. No. 3 Odonata and Protodonata. The Proceedings of the Linnean Society of New South Wales 43:417-436 [M. Clapham/J. Karr]
36567 R. J. Tillyard. 1918. Mesozoic Insects of Queensland. No. 4 Hemiptera Heteroptera: The family Dunstaniidae. With a note on the origin of the Heteroptera. Proceedings of the Linnean Society of New South Wales 43:568-592 [M. Clapham/J. Karr/M. Clapham]
36584 R. J. Tillyard. 1919. Mesozoic Insects of Queensland. No. 6 Blattoidea. The Proceedings of the Linnean Society of New South Wales 44:358-382 [M. Clapham/J. Karr]
36586 R. J. Tillyard. 1919. Mesozoic Insects of Queensland. No. 7 Hemiptera Homoptera; with a note on the phylogeny of the suborder. The Proceedings of the Linnean Society of New South Wales 44:857-896 [M. Clapham/J. Karr]
36572 R. J. Tillyard. 1919. Mesozoic Insects of Queensland. No. 5 Mecoptera, the new order Paratrichoptera, and additions to Planipennia. The Proceedings of the Linnean Society of New South Wales 44:194-212 [M. Clapham/J. Karr]
36590 R. J. Tillyard. 1921. Mesozoic Insects of Queensland. No. 8 Hemiptera Homoptera (contd) The genus Mesogereon; with a discussion of its relationship with the Jurassic Palaeontinidae. The Proceedings of the Linnean Society of New South Wales 46:270-284 [M. Clapham/J. Karr]
36601 R. J. Tillyard. 1922. Mesozoic Insects of Queensland. No. 9 Orthoptera, and additions to the Protorthoptera, Odonata, Hemiptera and Planipennia. The Proceedings of the Linnean Society of New South Wales 47:447-470 [M. Clapham/J. Karr]
36649 R. J. Tillyard. 1923. Mesozoic Insects of Queensland. No. 10 Summary of the Upper Triassic Insect fauna of Ipswich, Q. (With an Appendix describing new Hemiptera and Planipennia). The Proceedings of the Linnean Society of New South Wales 48:481-498 [M. Clapham/J. Karr/R. Day]