Iharkút mine, Bakony mountains (Unit 1): Santonian, Hungary
collected by A. Osi et al. 2000–

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Plantae indet. Haeckel 1866
Osi 2005
"carbonized plant fragments"
Osteichthyes
Osteichthyes indet. Huxley 1880
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteidae indet. Cuvier 1825
Osi 2008
Actinopteri - Pycnodontiformes
Pycnodontiformes indet. Berg 1937
Osi 2008
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpetontidae indet. Fox and Naylor 1982
Osi 2008 16 specimens
MTM V2008.22.1., V.2008.23.1., V2008.24.1., V2009.1.1., and V2010.288.1. (premaxillae); MTM V2008.28.1. and V2009.29.1. (maxillae); MTM V2008.25.1., V2008.26.1., V2008.29.1., V2009.8.1., V2009.9.1., V2009.26.1., V2009.32.1., and V2010.287.1. (dentaries) and MTM V2009.10.1. (indeterminate jaw fragment) - see Szentesi et al. (2013)
Albanerpetontidae informal indet. B Fox and Naylor 1982
Szentesi 2020 2 specimens
VER 2015.134.1.–2.
Albanerpetontidae informal indet. A Fox and Naylor 1982
Szentesi 2020 1 specimen
VER 2015.134.13.
Amphibia - Temnospondyli - Discoglossidae
Bakonybatrachus fedori n. gen., n. sp. Szentesi and Venczel 2012
Szentesi and Venczel 2012 4 specimens
MTM V 2010.283.1. - holotype (incomplete right ilium); referred material: fragmentary maxilla (MTM V 2009.34.1.), fragmentary angulosplenial (MTM V 2008.31.1.) and fragmentary scapula (MTM V 2008.30.1.)
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Osi 2004 12 specimens
"frog"
    = Hungarobatrachus szukacsi n. gen., n. sp. Szentesi and Venczel 2010
Szentesi and Venczel 2010
MTM V2008.16.1 - holotype (incomplete right ilium); referred material: five right ilia (MTM V2008.12.1, 2008.13.1, 2008.15.1, 2008.17.1, 2008.18.1), one left ilium (MTM V2008.14.1); one tibio-fibula (MTM V2008.32.1); and four fragmentary tibio-fibulae (MTM V2008.19.1, 2008.21.1, 33.1, 34.1)
Reptilia
Squamata indet. (Oppel 1811)
Prondvai et al. 2017
eggshell morphotype MT IV
Chromatogenys tiliquoides n. gen., n. sp. Makadi and Nydam 2015
Makadi and Nydam 2015 1 specimen
MTM V 2010.129.1 (holotype), partial mandible
Reptilia - Polyglyphanodontidae
Distortodon rhomboideus n. gen., n. sp. Makadi 2013
Makadi 2013 3 specimens
Holotype. MTM PAL2012.31.1. partial right maxilla; MTM PAL2012.32.1. right dentary (Fig. 3) and MTM PAL2012.33.1. partial left dentary
Bicuspidon aff. hatzegiensis Folie and Codrea 2005
Makádi 2006 7 specimens
MTM V.01.24., left dentary fragment; MTM V.01.27., right lower jaw fragment; MTM 2006.108.1, left dentary; MTM 2006.109.1, left dentary fragment; MTM 2006.110.1, dentary fragment; MTM 2006.111.1, left dentary; MTM 2006.112.1, left dentary; MTM 2006.114.1, jaw fragment
Reptilia - Chamopsiidae
Pelsochamops infrequens n. gen., n. sp. Makadi 2013
Makadi 2013 3 specimens
MTM 2006.106.1 (holotype), partial left mandible; MTM PAL 2013.24.1, right dentary fragment; MTM PAL 2013.25.1, left dentary fragment
Reptilia
Mosasauroidea indet. ()
Osi 2008 4 individuals
    = Pannoniasaurus inexpectatus n. gen., n. sp. Makádi et al. 2012
Makádi et al. 2012
MTM 2011.43.1. (holotype) isolated right quadrate; MTM V.01.115., left quadrate; MTM 2007.31.1., fragmentary left quadrate; plus 2 isolated premaxillae; 3 maxillae; 2 postorbitofrontals; 2 quadrates; 3 dentaries; 3 splenials; 3 angulars; coronoid; 2 surangulars; articular; 91 isolated teeth; 20 cervical, 40 dorsal, 4 sacral, and 18 caudal vertebrae; 34 vertebral fragments; 3 ribs; 2 humeral fragments; 4 ilia
Reptilia - Lacertilia
Lacertilia indet. Owen 1842
"lizards"
Reptilia - Teiidae
Teiidae indet. Gray 1827
Osi 2008
Reptilia
Pterodactyloidea indet. (Plieninger 1901)
Osi et al. 2011 2 specimens
MTM V 2010.98.1 (L articular region of lower jaw), MTM V 2010.99.1 (MC IV)
Reptilia - Azhdarchidae
Pterosauria indet. Kaup 1834
13 specimens
    = Azhdarchidae indet. Nessov 1984
Osi et al. 2005
Atlas-axis complex (MTM V 2010.82.1.), posterior part of mid-cervical vertebra (MTM V 2010.81.1.), glenoid region of scapulocoracoid (MTM V 2010.79.1.); distal frag of humerus (MTM V 2010.77.1.), distal frags of 2 ulnae (MTM V 2010.78.1. MTM V 2010.75.1.1⁄4Gyn/453); ?third or ?fourth wing phalanx (MTM V 2010.83.1.); prox. tibia frag (MTM V 2010.76.1.); 4 cervical vertebrae (MTM V 2010.100.1.1⁄4Gyn/448, MTM V.01.51.1⁄4Gyn/449, MTM V 2010.101.1.1⁄4Gyn/450, MTM V. 2003.21.1⁄4Gyn/451), R radius (MTM V 2010.102.1.1⁄4Gyn/452), prox. half of first wing phalanx (MTM V.2002.04.)
Bakonydraco galaczi n. gen., n. sp. Osi et al. 2005
Osi et al. 2005 45 specimens
MTM Gyn/3 (holotype), nearly complete mandible. MTM Gyn/4, 21 dentary symphyseal fragments; MTM V 2010.80.1 (frag. premaxilla), 2010.74.1–74.22 (dentary symphyseal frags.)
Reptilia
Theropoda informal indet. 3 (Marsh 1881)
Osi 2004 1 specimen
"Morphotype 3"
Theropoda indet. (Marsh 1881)
Osi et al. 2010 8 specimens
MTM V 2009.48.1, sacrum fragment; MTM V 2009.47.1-6, caudal vertebra; MTM V 2009.49.1, distal left femur
Theropoda informal indet. 1 (Marsh 1881)
Osi 2004 58 specimens
"Morphotype 1"
    = Tetanurae indet. Gauthier 1986
Osi et al. 2010
58 teeth/tooth fragments: V.01.54, V.01.30, V.01.20, V.2003.04-08, V 2008.36.1-51
Maniraptora indet. Gauthier 1986
Prondvai et al. 2017 1 specimen
eggshell morphotype MT V
cf. Pseudogeckoolithus sp. Vianey-Liaud and López-Martínez 1997
Prondvai et al. 2017
eggshell morphotype MT I
    = Pseudogeckoolithus sp. Vianey-Liaud and López-Martínez 1997
Choi et al. 2020
Theropoda informal indet. 2 (Marsh 1881)
Osi 2004 29 specimens
"Morphotype 2"
    = Paraves indet. Sereno 1997
Osi et al. 2010
Isolated teeth V.01.215, V.01.231, V.2000.35, V 2008.37.1-10, V.2000.02-06; distal caudal vertebrae V 2009.46.1-3; metacarpal V 2009.45.1; 3 ungual phalanges V 2008.40.1-3; phalanges V 2008.42.1-2; half of left tibia V 2008.31.1; MTM PAL 2011.18 (R tibiotarsus), MTM PAL 2011.19 (L? MC III)
Paraves indet. Sereno 1997
Prondvai et al. 2017
eggshell morphotype MT III
Aves indet. (Linnaeus 1758)
Dyke and Osi 2010 3 specimens
MTM V 2009.41.1, distal L femur; MTM V.2002.06, distal subadult femur; MTM V 2003.19, metatarsal III
Aves indet. Linnaeus 1758
5 specimens
MTM V.2002.06, fig. 2G-I, distal end of L femur; MTM V.2003.19, metatarsal III
    = Enantiornithes indet. Walker 1981
Osi 2005
MTM Gyn 439, near-complete R tarsometatarsus; MTM V 2009.39.1, distal L humerus; MTM V.2002.05, L femur
Reptilia - Avisauridae
? Bauxitornis sp. Dyke and Osi 2010
Osi and Buffetaut 2011 2 specimens
MTM PAL 2011.20 & 21 (2 femora)
Bauxitornis mindszentyae n. gen., n. sp. Dyke and Osi 2010
Dyke and Osi 2010 1 specimen
MTM V 2009.38.1, R tarsometatarsus
Reptilia
Pneumatoraptor fodori n. gen., n. sp. Osi et al. 2010
Osi et al. 2010 1 specimen
MTM V.2008.38.1. - holotype (left scapulacoracoid)
Reptilia - Abelisauridae
Abelisauridae indet. Bonaparte and Novas 1985
Osi et al. 2010 2 specimens
MTM V 2008.43.1 (pedal ungual); MTM PAL 2011.17 (R femur)
Reptilia
Titanosauriformes indet. Salgado et al. 1997
Ősi et al. 2017 1 specimen
MTM PAL 2017.1.1
Ajkaceratops kozmai n. gen., n. sp. Osi et al. 2010
Osi et al. 2010 5 specimens
MTM V 2009.192.1 (holotype), fused premaxillae and rostral bones. MTM V2009.193.1, V2009.194.1, V2009.195.1, V2009.196.1, predentaries
Reptilia - Rhabdodontidae
Rhabdodontidae indet. Weishampel et al. 2003
4 specimens
Mochlodon vorosi n. sp. Osi et al. 2012
Osi et al. 2012 71 specimens
Left complete dentary with four broken teeth (MTM V 2010.105.1; holotype); left postorbital (MTM 2012.14.1); two right quadrates (MTM V 2010.110.1, V 2010.111.1), two left (MTM V 2010.105.1., 2012.15.1) and two right (MTM V 2010.107.1., V 2010.109.1.) dentaries, all four of which are almost complete, six fragmentary dentaries (MTM V 2010 106.1, V 2010 107.1, V 2010 108.1, V 2010 109.1, V 2010.112.1, 2012.16.1), 15 maxillary and 23 dentary teeth (MTM V 2000.01., V 2000.32., V 2000.33., V 2003.10., V 01.161., V 2003.14,– V.2003.16, V 01.64., 2012.17.1, 2012.18.1), isolated cervical (MTM 2012.19.1), dorsal (MTM 2010.118.1.), and caudal (MTM 2012.20.1, 2012.21.1) vertebrae, almost complete but compressed sacrum (MTM V 2010.121.1.), three coracoids (MTM V 01.53., V 2010.122.1., V 2010.123.1.), one fragmentary scapula (MTM 2012.22.1), one fragmentary (MTM 2012.23.1) and one complete humerus (MTM V 2010.128.1.), one complete ulna (MTM 2012.24.1), two almost complete femora (MTM V 01.225., V 2010.126.1.), one fragmentary femur (MTM 2012.25.1), one complete tibia (MTM V 2010.127.1.), two fragmentary tibiae (MTM V 01.101., 2012.26.1), and two phalanges (MTM 2012.27.1, 2012.28.1).
Reptilia - Nodosauridae
Hungarosaurus sp. Osi 2005
Osi et al. 2014 1 specimen
PAL 2013.23.1, partial skull
Nodosauridae indet. Marsh 1890
600 specimens
MNI = 5 (5 measurements)
    = Hungarosaurus tormai n. gen., n. sp. Osi 2005
Osi 2005
MTM PAL 2012.30.1 (humerus); 6 other specimens
Reptilia
Atoposauridae indet. (Gervais 1871)
Martin et al. 2014
Maxillae and teeth
    = Neosuchia indet. Clark 1988
Tennant et al. 2016
Alligatoroidea indet. (Gray 1844)
Osi 2008
A few isolated skull fragments and hundreds of teeth
    = Eusuchia indet. Huxley 1875
Puértolas-Pascual et al. 2016
basal eusuchian possibly related to Allodaposuchus, Musturzabalsuchus, or Massaliasuchus
Crocodylia indet. (Owen 1842)
"crocodile"
    = Eusuchia indet. Huxley 1875
Osi et al. 2005
    = Iharkutosuchus makadii Osi et al. 2007
Osi et al. 2007
Holotype skull (MTM 2006.52.1); paratypes: nearly complete skull (MTM 2006.53.1), 3 fragmentary skulls (MTM 2006.54.1-56.1), skull fragments (MTM 2006.57.1-79.1), 148 isolated teeth (MTM 2006.80.1)
Crocodylia indet. (Owen 1842)
Prondvai et al. 2017
eggshell morphotypes MT IIa, b, c
Doratodon sp. Seeley 1881
Osi 2008 233 specimens
Single fragmentary dentary and several isolated teeth
    = Doratodon carcharidens Bunzel 1871
Rabi and Sebők 2015
R premaxilla (MTM PAL 2014.122.1) w/teeth, fragmentary L maxilla (MTM PAL 2013.65.1), fragmentary L dentary (MTM V 2010.237.1), fragmentary R dentary (MTM PAL 2013.66.1) and 229 isolated teeth (MTM PAL 2014.123.1–PAL 2014.123.10, MTM V 2010.226.1.1–V 2010.226.1.7)
Reptilia - Testudines - Dortokidae
Rabi et al. 2013 9 specimens
Reptilia - Testudines - Bothremydidae
Bothremydidae indet. Baur 1891
Osi 2008
Reptilia - Testudines
Testudines indet. Batsch 1788
Osi 2004 8 specimens
"turtle"
    = Foxemys trabanti n. sp. Rabi et al. 2012
Rabi et al. 2012
MTM V2010.86.1 (holotype), incomplete skull; MTM V2010.215.1., skull; MTM V2010.216.1., partial skull; MTM V2010.87.1., partial skull; MTM uncatalogued partial skull; MTM V2010.88.1., left otic chamber; MTM V2010.89.1., fragmentary left lower jaw; MTM V2010.219.1., right and left lower jaw; MTM V2010.220.1., left lower jaw.
Reptilia
cf. Kallokibotioninae indet. Nopsca 1923
Rabi et al. 2013 2 specimens
    = Kallokibotion sp. Nopcsa 1923
Joyce 2017
see common names

Geography
Country:Hungary State/province:Veszprém
Coordinates: 47.2° North, 17.6° East (view map)
Paleocoordinates:31.0° North, 18.6° East
Basis of coordinate:based on nearby landmark
Altitude:354 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Santonian 10 m.y. bin:Cretaceous 6
Key time interval:Santonian
Age range of interval:86.30000 - 83.60000 m.y. ago
Stratigraphy
Formation:Csehbánya
Stratigraphic resolution:group of beds
Stratigraphy comments: two bone-bearing beds within section, approx. 0.5 m apart
Lithology and environment
Primary lithology:green sandy claystone
Secondary lithology:pebbly,gray,green argillaceous sandstone
Includes fossils?Y
Includes fossils?Y
Lithology description: "The Csehbánya Formation consists of variegated clay and sandstone beds...The fauna is concentrated near the top of the outcrop in 20 cm thick sandy clay, sandstone bed...Most of the fossils derived from 5 cm thick sandy clay layer below the sandstone. Other teeth and bones occur disordered on the surface of the sandstone bed and are covered with clay less than 1 cm thick." "The vertebrate-bearing beds form 2–3 m thick cyclical units beginning with green clay covered by breccia containing clay clasts, sandstone and siltstone layers (Tuba et al. 2006). Most of the material, including the turtles comes from the matrix supported breccia consisting of ripped-up variegated clay clasts, dolomite pebbles, carbonized plant remains and sand forming channel-like structures with the isolated bones accumulated in lenses" (Rabi et al. 2012) SZ-6: "a 10–50-cm-thick basal breccia layer composed of gray-green sand, siltstone, clay clasts, and pebbles. The poorly sorted sandy breccia is interrupted by laminated siltstone horizons; these coarse- and fine-grained layers are repeated several times resulting in a stacked fining-upward series of units."
Environment:crevasse splay
Geology comments: "deposited in an alluvial plain"; fossils "are buried in mud- and sand-filled channel deposits"
"The palaeoenvironment is interpreted as an alluvial plain and the deposition of the breccia could be the result of the episodic filling of a channel or pond by debris flows, silt and sand" (Osi & Mindszenty, 2009).
"a flash-flood sequence deposition, which started with a high flow regime characterized by sandstone and clay clasts, whereas the siltstone layers are indicative of waning flow velocity and deposition from suspension in standing water, following the minor flow pulse. Along with its taphonomic characteristics, this sedimentology suggests that Unit 1 was deposited by ephemeral high-density flash-flood events probably triggered by episodic heavy rainfalls." (Prondvai et al. 2017)
Taphonomy
Modes of preservation:body
Lagerst�tten type:concentrate
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:medium
Articulated whole bodies:none
Associated major elements:some
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:occasional
Temporal resolution:time-averaged
Collection methods and comments
Collection methods:bulk,selective quarrying,surface (float),surface (in situ),sieve,field collection
Collection size:410 specimens
Reason for describing collection:general faunal/floral analysis
Collectors:A. Osi et al. Collection dates:2000–present
Taxonomic list comments:Fragmentary turtle plates are among the most abundant remains at the site.
Metadata
Also known as:Németbánya, Iharkut, SZ-6
Database number:48722
Authorizer:M. Carrano, P. Holroyd, R. Butler, P. Mannion, P. Barrett, R. Benson Enterer:M. Carrano, J. Tennant, P. Holroyd, K. Maguire, T. Cleary, R. Benson, P. Mannion, M. Oreska, R. Butler
Modifier:M. Carrano Research group:vertebrate
Created:2005-03-30 05:50:53 Last modified:2023-11-20 13:19:47
Access level:the public Released:2005-03-30 05:50:53
Creative Commons license:CC BY
Reference information

Primary reference:
15353.ETE A. Osi, C.-M. Jianu, and D. B. Weishampel. 2003. Dinosaurs from the Upper Cretaceous of Hungary. In A. Petculescu & E. Stiuca (eds.), Advances in Vertebrate Paleontology: Hen to Panta. Romanian Academy, "Emil Racovita" Institute of Speleology, Bucarest 117-120 [M. Carrano/M. Carrano]

Secondary references:

86530 G. Botfalvai, E. Prondvai, and A. 2021. Living alone or moving in herds? A holistic approach highlights complexity in the social lifestyle of Cretaceous ankylosaurs. Cretaceous Research 118:104633 [M. Carrano/M. Carrano]
81824 S. Choi, M. Moreno-Azanza, Z. Csiki-Sava, E. Prondvai, and Y.-N. Lee. 2020. Comparative crystallography suggests maniraptoran theropod affinities for latest Cretaceous European ‘geckoid’ eggshell. Papers in Palaeontology 6(2):265-292 [M. Carrano/M. Carrano]
33409ETE G. J. Dyke and A. Osi. 2010. A review of Late Cretaceous fossil birds from Hungary. Geological Journal 45:434-444 [M. Carrano/M. Oreska/P. Mannion]
80599 E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
62124 W. G. Joyce. 2017. A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58(1):65-113 [P. Holroyd/P. Holroyd/P. Mannion]
58011 L. Makádi. 2006. Bicuspidon aff. hatzegiensis (Squamata: Scincomorpha: Teiidae) from the Upper Cretaceous Csehbanya Formation (Hungary, Bakony Mts). Acta Geologica Hungarica 49(4):373-385 [P. Barrett/T. Cleary/T. Cleary]
58016 L. Makadi. 2013. The first known chamopsiid lizard (Squamata) from the Upper Cretaceous of Europe (Csehbanya Formation; Hungary, Bakony Mts). Annales de Paleontologie 99:261-274 [P. Barrett/T. Cleary]
49728 L. Makadi. 2013. A new polyglyphanodontine lizard (Squamata: Borioteiioidea) from the Late Cretacoues Iharkut locality (Santonian, Hungary). Cretaceous Research 46:166-176 [P. Mannion/J. Tennant/M. Carrano]
44074 L. Makádi, M. W. Caldwell, and A. Osi. 2012. The first freshwater mosasauroid (Upper Cretaceous, Hungary) and a new clade of basal mosasauroids. PLoS ONE 7(12):e51781 [R. Benson/R. Benson]
58020 L. Makadi and R. L. Nydam. 2015. A new durophagous scincomorphan lizard genus from the Late Cretaceous Iharkut locality (Hungary, Bakony Mts). Palaeontologische Zeitschrift 89(4):925-941 [P. Barrett/T. Cleary]
51121 J. E. Martin, M. Rabi, Z. Csiki-Sava and S. Vasile. 2014. Cranial morphology of Theriosuchus sympiestodon (Mesoeucrocodylia, Atoposauridae) and the widespread occurrence in the Late Cretaceous of Europe. Journal of Paleontology 88(3):444-456 [P. Mannion/J. Tennant/J. Tennant]
33406 H. Morlon, E. P. White, R. S. Etienne, J. L. Green, A. Ostling, D. Alonso, B. J. Enquist, F. He, A. Hurlbert, A. E. Magurran, B. A. Maurer, B. J. McGill, H. Olff, D. Storch, and T. Zillio. 2009. Taking species abundance distributions beyond individuals. Ecology Letters 12:488-501 [M. Carrano/M. Oreska]
12998ETE A. Osi. 2004. The first dinosaur remains from the Upper Cretaceous of Hungary (Csehbánya Formation, Bakony Mts). Géobios 37:749-753 [M. Carrano/M. Carrano/J. Alroy]
14093ETE A. Osi. 2005. Hungarosaurus tormai, a new ankylosaur (Dinosauria) from the Upper Cretaceous of Hungary. Journal of Vertebrate Paleontology 25(2):370-383 [M. Carrano/M. Carrano/M. Carrano]
36772 A. Osi. 2008. Cranial osteology of Iharkutosuchus makadii, a Late Cretaceous basal eusuchian crocodyliform from Hungary. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 248(3):279-299 [P. Mannion/P. Mannion]
46340 A. Osi. 2008. Enantiornithine bird remains from the Late Cretaceous of Hungary. Oryctos 7:55-60 [P. Mannion/J. Tennant]
32758ETE A. Osi, S. Apesteguia, and M. Kowalewski. 2010. Non-avian theropod dinosaurs from the early Late Cretaceous of central Europe. Cretaceous Research 31(3):304-320 [M. Carrano/M. Carrano]
68032 A. Osi and E. Buffetaut. 2011. Additional non-avian theropod and bird remains from the early Late Cretaceous (Santonian) of Hungary and a review of the European abelisauroid record. Annales de Paléontologie 97:34-49 [M. Carrano/M. Carrano]
68033 A. Osi, E. Buffetaut, and E. Prondvai. 2011. New pterosaurian remains from the Late Cretaceous (Santonian) of Hungary (Iharkút, Csehbánya Formation). Cretaceous Research 32:456-463 [M. Carrano/M. Carrano]
32660 A. Osi, R. J. Butler, and D. B. Weishampel. 2010. A Late Cretaceous ceratopsian dinosaur from Europe with Asian affinities. Nature 465:466-468 [R. Butler/R. Butler]
14526ETE A. Osi, J. M. Clark, and D. B. Weishampel. 2005. Mastication in the most primitive eusuchian crocodylian. Kaupia 14:91 [M. Carrano/K. Maguire/M. Carrano]
36771 A. Osi, J. M. Clark, and D. B. Weishampel. 2007. First report on a new basal eusuchian crocodyliform with multicusped teeth from the Upper Cretaceous (Santonian) of Hungary. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 243(2):169-177 [P. Mannion/P. Mannion]
62444 A. Ősi, Z. Csiki-Sava, and E. Prondvai. 2017. A sauropod tooth from the Santonian of Hungary and the European Late Cretaceous ‘sauropod hiatus’. Scientific Reports 7:3261:1-8 [P. Mannion/P. Mannion/M. Carrano]
32498 A. Osi and L. A. Makádi. 2009. New remains of Hungarosaurus tormai (Ankylosauria, Dinosauria) from the Upper Cretaceous of Hungary: skeletal reconstruction and body mass estimation. Palaeontologische Zeitschrift 83:227-245 [M. Carrano/M. Carrano]
49732 A. Osi, X. Pereda-Suberbiola, and T. Foldes. 2014. Partial skull and endocranial cast of the ankylosaurian dinosaur Hungarosaurus from the Late Cretaceous of Hungary: implications for locomotion. Palaeontologia Electronica 17(1):1-18 [P. Mannion/J. Tennant]
46700 A. Ősi and E. Prondvai. 2009. Forgotten pterosaurus in Hungarian collections: first description of Rhamphorhynchus and Pterodactylus specimens. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 252(2):167-180 [P. Mannion/J. Tennant/M. Clapham]
43083 A. Osi, E. Prondvai, R. Butler and D. B. Weishampel. 2012. Phylogeny, histology and inferred body size evolution in a new rhabdodontid dinosaur from the Late Cretaceous of Hungary. PLoS ONE 7(9):e44318. [R. Benson/R. Benson]
46669 A. Ösi and E. Prondvai. 2013. Sympatry of two ankylosaurs (Hungarosaurus and cf. Struthiosaurus) in the Santonian of Hungary. Cretaceous Research 44:58-63 [P. Mannion/J. Tennant/M. Carrano]
31045 A. Osi, D. B. Weishampel, and C. M. Jianu. 2005. First evidence of azhdarchid pterosaurs from the Late Cretaceous of Hungary. Acta Palaeontologica Polonica 50(4):777-787 [R. Butler/R. Butler]
86182 E. Prondvai. 2014. Comparative bone histology of rhabdodontid dinosaurs. Palaeovertebrata 38(2):e1:1-31 [M. Carrano/M. Carrano]
81826 E. Prondvai, G. Botfalvai, K. Stein, Z. Szentesi, and A. Osi. 2017. Collection of the thinnest: a unique eggshell assemblage from the Late Cretaceous vertebrate locality of Iharkút (Hungary). Central European Geology 60(1):73-133 [M. Carrano/M. Carrano]
76284 E. Puértolas-Pascual, A. Blanco, C. A. Brochu and J. I. Canudo. 2016. Review of the Late Cretaceous-early Paleogene crocodylomorphs of Europe: Extinction patterns across the K-PG boundary. Cretaceous Research 57:565-590 [P. Mannion/P. Mannion]
53482 M. Rabi and N. Sebők. 2015. A revised Eurogondwana model: Late Cretaceous notosuchian crocodyliforms and other vertebrate taxa suggest the retention of episodic faunal links between Europe and Gondwana during most of the Cretaceous. Gondwana Research 28(3):1197-1211 [P. Mannion/P. Mannion/P. Mannion]
42082 M. Rabi, H. Tong, and G. Botfalvai. 2012. A new species of the side-necked turtle Foxemys (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Hungary and the historical biogeography of the Bothremydini. Geological Magazine 149(4):662-674 [R. Benson/R. Benson/M. Carrano]
59665 M. Rabi, M. Vremir, and H. Tong. 2013. Preliminary Overview of Late Cretaceous Turtle Diversity in Eastern Central Europe (Austria, Hungary, and Romania). In D. B. Brinkman, P. A. Holroyd, and J. D. Gardner (eds.), Morphology and Evolution of Turtles 307-336 [P. Holroyd/P. Holroyd]
77157 Z. Szentesi. 2020. Új albanerpetontida (Allocaudata: Albanerpetontidae) leletek az iharkúti késő-kréta szárazföldi ősgerinces lelőhelyről [New albanerpetontid remains (Allocaudata: Albanerpetontidae) from the Late Cretaceous terrestrial paleovertebrate locality of Iharkút]. Annales Musei Historico-Naturalis Hungarici 112:37-55 [M. Carrano/M. Carrano]
46398 Z. Szentesi, J. D. Gardner, and M. Venczel. 2013. Albanerpetontid amphibians from the Late Cretaceous (Santonian) of Iharkút, Hungary, with remarks on regional differences in Late Cretaceous Laurasian amphibian assemblages. Canadian Journal of Earth Sciences 50:268-281 [P. Mannion/P. Mannion]
38712 Z. Szentesi and M. Venczel. 2010. An advanced anuran from the Late Cretaceous (Santonian) of Hungary. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 256(3):291-302 [P. Mannion/P. Mannion]
39482 Z. Szentesi and M. Venczel. 2012. A new discoglossid frog from the Upper Cretaceous (Santonian) of Hungary. Cretaceous Research 34:327-333 [P. Mannion/P. Mannion]
59838 J. P. Tennant, P. D. Mannion, and P. Upchurch. 2016. Evolutionary relationships and systematics of Atoposauridae (Crocodylomorpha: Neosuchia): implications for the rise of Eusuchia. Zoological Journal of the Linnean Society 177:854-936 [P. Mannion/P. Mannion]