Ukhaa Tolgod (AMNH): Late/Upper Campanian, Mongolia
collected 1992-1996

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia
Theria indet. Parker and Haswell 1897
"new genus D"
Mammalia - Deltatheroida - Deltatheridiidae
Deltatheridium pretrituberculare Gregory and Simpson 1926
Rougier et al. 1998 2 specimens
Mammalia
cf. Asioryctes sp. Kielan-Jaworowska 1975
    = Ukhaatherium nessovi n. gen., n. sp. Novacek et al. 1997
Novacek et al. 1997
PSS-MAE 102 - holotype; referred material: PSS-MAE 105 and 106; 3 individuals
Mammalia - Asioryctidae
Asioryctes nemegtensis Kielan-Jaworowska 1975
Novacek et al. 2022
PSS-MAE 366
Mammalia
Kennalestes cf. gobiensis Kielan-Jaworowska 1968
Mammalia - Zalambdalestidae
Zalambdalestes lechei Gregory and Simpson 1926
Mammalia - Proteutheria - Palaeoryctidae
cf. Hyotheridium sp. Gregory and Simpson 1926
Mammalia - Allotheria
Allotheria indet. Marsh 1880
"new genus C"
Mammalia - Multituberculata - Eucosmodontidae
Eucosmodontidae indet. Jepsen 1940
"new genus B"
Mammalia - Multituberculata
Nemegtbaatar gobiensis Kielan-Jaworowska 1974
Mammalia - Multituberculata - Djadochtatheriidae
Tombaatar sabuli n. gen., n. sp. Rougier et al. 1997
Rougier et al. 1997 1 individual
PSS-MAE 122 (partial skull)
Catopsbaatar sp. Kielan-Jaworowska 1994
This is a typo? "Catopsbaatar Catopsaloides"
Mammalia - Multituberculata - Sloanbaataridae
Sloanbaatar cf. mirabilis Kielan-Jaworowska 1969
Kamptobaatar cf. kuczynskii Kielan-Jaworowska 1969
Mammalia - Multituberculata
Chulsanbaatar cf. vulgaris Kielan-Jaworowska 1974
Bulganbaatar cf. nemegtbaataroides Kielan-Jaworowska 1974
"new genus A"
Reptilia
Myrmecodaptria microphagosa n. gen., n. sp. Gao and Norell 2000
Gao and Norell 2000 1 individual
Parmeosaurus scutatus n. gen., n. sp. Gao and Norell 2000
Gao and Norell 2000 3 individuals
IGM 3/138, 141, 143
Hymenosaurus clarki n. gen., n. sp. Gao and Norell 2000
Gao and Norell 2000 1 individual
Globaura venusta Borsuk-Bialynicka 1988
Gao and Norell 2000 4 individuals
IGM 3/160-3/163
Carusia intermedia (Borsuk-Bialynicka 1985)
Gao and Norell 1998 35 individuals
IGM 3/16-3/50
Reptilia - Carusiidae
Shinisauroides latipalatum Borsuk-Bialynicka 1985
Reptilia - Polyglyphanodontidae
Gobinatus arenosus Alifanov 1993
Gao and Norell 2000 1 individual
IGM 3/128
Adamisaurus magnidentatus Sulimski 1972
5 individuals
IGM 3/111-3/115
Macrocephalosaurus sp. Gilmore 1943
Gao and Norell 2000 2 individuals
replaced by Gilmoreteius
IGM 3/132-3/133
Reptilia - Slavoiidae
Slavoia darevskii Sulimski 1984
Gao and Norell 2000 9 individuals
IGM 3/145-3/153
Reptilia - Priscagamidae
Priscagama gobiensis Borsuk-Bialynicka and Moody 1984
2 individuals
IGM 3/77-3/78
? Priscagama gobiensis Borsuk-Bialynicka and Moody 1984
Gao and Norell 2000 1 individual
IGM 3/80
Phrynosomimus asper Alifanov 1996
Gao and Norell 2000 1 individual
IGM 3/83
Mimeosaurus crassus Gilmore 1943
Gao and Norell 2000 2 individuals
IGM 3/74-3/75
Reptilia
Saichangurvel davidsoni n. gen., n. sp. Conrad and Norell 2007
Conrad and Norell 2007 1 individual
IGM 3/858 (holotype skeleton)
Temujinia ellisoni n. gen., n. sp. Gao and Norell 2000
Gao and Norell 2000 7 specimens
IGM 3/63-3/69
Isodontosaurus gracilis Gilmore 1943
Gao and Norell 2000 4 individuals
IGM 3/85, 86, 88, 89
Gobiderma pulchrum (Borsuk-Bialynicka 1984)
Gao and Norell 2000 3 individuals
misspelling Gobiderma pulchra
IGM 3/56-3/58
Estesia mongoliensis Norell et al. 1992
1 individual
IGM 3/169 (a skull with mandible, preserved three-dimensionally) and IGM 3/760 (partial axial skeletons including dorsal vertebrae with articulating ribs and an incomplete left forelimb)
Varanoidea indet. Gray 1827
Gao and Norell 2000 1 individual
IGM 3/172
Reptilia - Testudines - Nanhsiungchelyidae
? Basilemys sp. Hay 1902
Reptilia - Paralligatoridae
Shamosuchus djadochtaensis Mook 1924
Pol et al. 2009
IGM 100/1195 (nearly complete skull, lower jaws, and associated postcranial remains including; all eight cervical vertebrae with three right cervical ribs (c5–c7) and one left cervical rib (c8), dorsal vertebrae d1 through d3 with four right dorsal ribs (d1–d3) and three left dorsal ribs (d2?–d6?), five anterior caudal vertebrae and two haemal arches, a complete right humerus, the distal half of the left humerus, the proximal portion of the right radius, a complete left radius in association with the left humerus, a complete left ulna, a fragmentary right ischium, a partial right femur consisting of a damaged diaphysis and distal condyle, a complete right tibia, a nearly complete left tibia, a complete right fibula, a partial left fibula, and a single left metatarsal and phalanx)
Reptilia - Protoceratopsidae
Bagaceratops sp. Maryanska and Osmólska 1975
Reptilia - Ankylosauridae
Ankylosauridae indet. Brown 1908
Reptilia - Dromaeosauridae
cf. Velociraptor sp. Osborn 1924
"neonate skulls"
Velociraptor mongoliensis Osborn 1924
Reptilia
cf. Gobipteryx sp. Elzanowski 1974
    = Gobipteryx minuta Elzanowski 1974
Chiappe et al. 2001
based on IGM-100/1011, collected in 1994 from "Ukhaa Tolgod" (no details given)
Reptilia - Oviraptoridae
Oviraptoridae indet. Barsbold 1976
"taxon B"
Reptilia - Alvarezsauridae
Mononykus olecranus (Perle et al. 1993)
1 specimen
IGM 100/975 (1 measurement)
    = Shuvuuia deserti n. gen., n. sp. Chiappe et al. 1998
Chiappe et al. 1998
Reptilia - Avetheropoda
Carnosauria indet. Huene 1920
see common names

Geography
Country:Mongolia State/province:Omnogov Aimak
Coordinates: 43.7° North, 100.9° East (view map)
Paleocoordinates:40.9° North, 90.2° East
Basis of coordinate:based on nearby landmark
Geographic resolution:local area
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
Key time interval:Late/Upper Campanian
Age range of interval:83.60000 - 72.10000 m.y. ago
Stratigraphy
Formation:Djadokhta
Stratigraphy comments: Age is not well constrained for the Djadokhta Formation (Clark et al. 2001). "Fossil vertebrate sites occur in the uppermost 65 m of a section..." see Fig. 1b, Dashzeveg et al. (1995).

Dashzeveg et al. 2005: The quick stratigraphic succession of normal and reversed magnetozones suggests, but does not clearly establish, that the sediments may have been deposited during the rapid sequence of polarity changes in the late part of the Campanian between about 75 to 71 Ma.
Lithology and environment
Primary lithology:"cross stratification",fine,coarse,red silty sandstone
Secondary lithology:red or brown mudstone
Includes fossils?Y
Includes fossils?Y
Lithology description: "Fossil vertebrate sites occur in the uppermost 65 m of a section dominated by red-orange, slightly silty, fine-to-coarse-grained sandstones with occasional beds of red-brown mudstones and grey-brown conglomerates (Fig. 1b)...The most fossiliferous sites are stratigraphically situated in a prominent cross-bedded unit of sandstone near the middle of the section." --Dashzeveg et al. 1995
Environment:eolian indet.
Taphonomy
Modes of preservation:body
Degree of concentration:-multiple events
Size of fossils:macrofossils,mesofossils
Spatial orientation:life position
Preservation of anatomical detail:excellent
Articulated whole bodies:some
Associated major elements:some
Disassociated major elements:some
Disassociated minor elements:some
Size sorting:poor
Fragmentation:occasional
Encrustation:none
Temporal resolution:snapshot
Spatial resolution:autochthonous
Collection methods and comments
Collection methods:surface (float),surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Collection dates:1992-1996
Collection method comments: Institute of Geology, Mongolia
Taxonomic list comments:Over 100 recovered and uncollected articulated skeletons of theropod, ankylosaurian, and protoceratopsian dinosaurs form the main 4 sq km collecting area. Also skulls (many with associated skeletons) of over 400 mammals and lizards. These are just the specimen numbers they include, there are certainly more in number.
Metadata
Database number:37184
Authorizer:J. Alroy, R. Benson, M. Carrano, P. Mannion Enterer:R. Whatley, R. Benson, P. Mannion, M. Carrano, J. Alroy
Modifier:G. Varnham Research group:vertebrate
Collections that are a subset of this one:37211, 37212, 37213, 45090, 45329, 68519, 68520, 68521, 68522, 68523, 68524, 68526, 222595
Created:2004-03-03 16:56:49 Last modified:2022-01-28 08:04:32
Access level:the public Released:2004-03-03 16:56:49
Creative Commons license:CC BY
Reference information

Primary reference:
5757.ETE 1515D. Dashzeveg, M. J. Novacek, M.A. Norell, J.M. Clark, L.M. Chiappe, A. Davidson, M.C. McKenna, L. Dingus, C. Swisher and P. Altangerel. 1995. Extraordinary preservation in a new vertebrate assemblage from the Late Cretaceous of Mongolia. Nature 374:446-449 [A. Behrensmeyer/A. Behrensmeyer/R. Whatley]

Secondary references:

13618ETE L. M. Chiappe, M. A. Norell, and J. M. Clark. 1998. The skull of a relative of the stem-group bird Mononykus. Nature 392:275-278 [M. Carrano/M. Carrano/M. Carrano]
28690 L. M. Chiappe, M. Norell, and J. Clark. 2001. A new skull of Gobipteryx minuta (Aves, Enantiornithes) from the Cretaceous of the Gobi Desert. American Museum Novitates 3346:1-15 [J. Alroy/J. Alroy]
38116 J. L. Conrad and M. A. Norell. 2007. A complete Late Cretaceous iguanian (Squamata, Reptilia) from the Gobi and identification of a new iguanian clade. American Museum Novitates 3584:1-47 [R. Benson/R. Benson]
15459ETE D. Dashzeveg, L. Dingus, D.B. Loope, C.C. Swisher, III, T. Dulam and M.R. Sweeney. 2005. New stratigraphic subdivision, depositional environment, and age estimate for the Upper Cretaceous Djadokhta Formation, southern Ulan Nur Basin, Mongolia. American Museum Novitates (3498)1-31 [M. Carrano/K. Maguire/G. Varnham]
197345% 54920K. Gao and M. A. Norell. 2000. Taxonomic composition and systematics of Late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert. Bulletin of the American Museum of Natural History 249:1-118 [J. Alroy/E. Leckey/R. Benson]
38066 K. -Q. Gao and M. A. Norell. 1998. Taxonomic revision of Carusia (Reptilia: Squamata) from the Late Cretaceous of the Gobi Desert and phylogenetic relationships of anguimorphan lizards. American Museum Novitates 3230:1-51 [R. Benson/R. Benson]
43987 I. Horovitz. 2003. Postcranial skeleton of Ukhaatherium nessovi (Eutheria, Mammalia) from the Late Cretaceous of Mongolia. Journal of Vertebrate Paleontology 23(4):857-868 [P. Mannion/P. Mannion]
85503 M. J. Novacek, E. A. Hoffman, and M. A. O'Leary. 2022. First occurrence of the eutherian mammal Asioryctes nemegtensis from the Upper Cretaceous Djadokhta Formation, Gobi Desert, Mongolia, and a revised alpha taxonomy based on the skull and dentition. Journal of Vertebrate Paleontology 42(4):e2196320 [P. Mannion/P. Mannion]
28809 M. J. Novacek, G. W. Rougier, J. R. Wible, M. C. McKenna, D. Dashzeveg and I. Horovitz. 1997. Epipubic bones in eutherian mammals from the Late Cretaceous of Mongolia. Nature 389:483-486 [J. Alroy/J. Alroy]
38082 D. Pol, A. H. Turner, and M. A. Norell. 2009. Morphology of the Late Cretaceous crocodylomorph Shamosuchus djadochtaensis and a discussion of neosuchian phylogeny as related to the origin of Eusuchia. Bulletin of the American Museum of Natural History 324:1-103 [P. Mannion/P. Mannion]
2986 G. W. Rougier, M. J. Novacek, and D. Dashzeveg. 1997. A new multituberculate from the Late Cretaceous locality Ukhaa Tolgod, Mongolia. Considerations on multituberculate interrelationships. American Museum Novitates 3191:1-26 [J. Alroy/J. Alroy/R. Benson]
99595% 2220G. W. Rougier, J. R. Wible, and M. J. Novacek. 1998. Implications of Deltatheridium specimens for early marsupial history. Nature 396(6710):459-463 [J. Alroy/R. Whatley/R. Whatley]
44530 H.-Y. Yi and M. A. Norell. 2013. New materials of Estesia mongoliensis (Squamata: Anguimorpha) and the evolution of venom grooves in lizards. American Museum Novitates 3767:1-31 [P. Mannion/P. Mannion]