Ikovo: Lutetian, Ukraine
collected 2010–2013

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
"Clerolamna" sp.
Udovichenko 2009
"Tobolamna" sp.
Udovichenko 2009
«Tobolamna» sp.
Sphyrenodus sp.
Udovichenko 2009
Osteichthyes indet. (Huxley 1880)
Zvonok and Skutschas 2011
Actinopteri - Scombriformes - Trichiuridae
Eutrichiurides sp. Casier 1944
Udovichenko 2009
Actinopteri - Scombriformes - Scombridae
Cybium sp. Cuvier 1829
Udovichenko 2009
Actinopteri - Labriformes - Labridae
Labrus sp. Linnaeus 1758
Udovichenko 2009
Actinopteri - Tetraodontiformes - Eotrigonodontidae
Eotrigonodon sp. Weiler 1929
Udovichenko 2009
Actinopteri - Tetraodontiformes - Triodontidae
Triodon sp. Cuvier 1829
Udovichenko 2009
Reptilia
Aves indet. Linnaeus 1758
Kievornis rogovitshi Averianov et al. 1990
Zvonok et al. 2015
Reptilia - Gaviidae
Colymbiculus udovichenkoi n. gen., n. sp. Mayr and Zvonok 2011
7 specimens
Reptilia - Odontopterygiformes - Pelagornithidae
? Dasornis sp. Owen 1870
6 specimens
cf. Odontopteryx toliapica Owen 1873
13 specimens
    = Lutetodontopteryx tethyensis n. gen., n. sp. Mayr and Zvonok 2012
Mayr and Zvonok 2012
Reptilia - Charadriiformes
Charadriiformes indet. Huxley 1867
Zvonok and Skutschas 2011
Reptilia
Crocodylia indet. (Owen 1842)
Kuzmin and Zvonok 2021
ZIN PH 2/145, 59/145, 61/145–74/145
Tomistominae indet. Eastman 1902
Zvonok and Skutschas 2011 1 specimen
ZIN PH 1/145 (anterior fragment of a right dentary)
cf. Diplocynodon sp. Pomel 1847
Kuzmin and Zvonok 2021
ZIN PH 52/145, 53/145, 55/145, 56/145, 57/145, 58/145, 60/145
Reptilia - Testudines
Testudinoidea indet. Gray 1825
Danilov et al. 2011
Argillochelys sp. Lydekker 1889
Zvonok et al. 2010
cf. Glossochelys sp. Seeley 1871
Звонок 2011
synonym of Euclastes
Reptilia - Testudines - Cheloniidae
Cheloniidae indet. Bonaparte 1832
Zvonok et al. 2013
The material of Cheloni idae gen. indet. (Fig. 2) includes the following ele ments: specimens ZIN PH, nos. 18/145 and 19/145, two postorbitals; specimen ZIN PH, no. 20/145, frag mentary right half of upper jaw; specimen ZIN PH, no. 21/145, costal plate IV; specimen ZIN PH, no.22/145, costal plate VII; specimens ZIN PH, nos. 23–27/145, fragments of five costal plates; speci men ZIN PH, no. 28/145, right peripheral plate II (?); specimen ZIN PH, no. 29/145, left peripheral plate II (?); specimen ZIN PH, no. 30/145, peripheral plate VI or VII; specimen ZIN PH, no. 31/145, posterior part of left epiplastron; specimen ZIN PH, no. 32/145, medial part of left hyoplastron; specimen ZIN PH, no. 33/145, medial part of right hypoplastron; speci men ZIN PH, no. 34/145, proximal part of left humerus; specimens ZIN PH, nos. 35/145 and 36/145, two right tibiae
Puppigerus nessovi Averianov 2005
Danilov et al. 2011
synonym of Puppigerus camperi
Eochelone sp. Dollo 1903
Звонок 2011
Reptilia - Testudines - Trionychidae
Trionychidae indet. Gray 1825
Zvonok et al. 2010
Trionyx ikoviensis n. sp. Danilov et al. 2011
Danilov et al. 2011 1 specimen
Acipenseriformes
Cylindracanthus sp. Leidy 1856
Udovichenko 2009
Chondrichthyes - Chimaeriformes - Edaphodontidae
Edaphodon sp. Buckland 1838
Zvonok and Удовиченко 2011
Chondrichthyes - Myliobatiformes - Aetobatidae
Aetobatus sp. Blainville 1816
Udovichenko 2009
Chondrichthyes - Myliobatiformes - Myliobatidae
Burnhamia sp. Cappetta 1976
Udovichenko 2009
Myliobatis sp. Cuvier 1816
Udovichenko 2009
Myliobatis sp. 1, 2
Chondrichthyes - Heterodontiformes - Heterodontidae
Heterodontus sp. de Blainville 1816
Udovichenko 2009
Heterodontus cf. vincenti (Leriche 1905)
Udovichenko 2009
original and current combination Cestracion vincenti
Chondrichthyes - Dalatiidae
Isistius trituratus (Winkler 1874)
Udovichenko 2009
Chondrichthyes - Squatinidae
Squatina cf. prima (Winkler 1874)
Udovichenko 2009
Chondrichthyes - Squalomorphii - Hexanchidae
Notorhynchus sp. Ayres 1855
Zvonok and Удовиченко 2011
Chondrichthyes - Carcharhiniformes - Scyliorhinidae
Abdounia lapierri
Udovichenko 2009
Abdounia cf. minutissima (Winkler 1874)
Udovichenko 2009
Chondrichthyes - Carcharhiniformes - Triakidae
Galeorhinus sp. de Blainville and Prévost 1816
Udovichenko 2009
Galeorhinus cf. ypresiensis (Casier 1946)
Udovichenko 2009
Chondrichthyes - Carcharhiniformes - Carcharhinidae
Rhizoprionodon gantourensis (Arambourg 1952)
Udovichenko 2009
misspelling Rhizoprionodon ganntourensis
Carcharhinus sp. de Blainville 1816
Udovichenko 2009
Physogaleus minor
Udovichenko 2009
Physogaleus secundus (Winkler 1876)
Udovichenko 2009
recombined as Physodon secundus
Chondrichthyes - Lamniformes
Isurolamna cf. bajarunasi
Udovichenko 2009
Chondrichthyes - Lamniformes - Mitsukurinidae
Striatolamia macrota (Agassiz 1838)
Udovichenko 2009
original and current combination Otodus macrotus
Chondrichthyes - Lamniformes - Lamnidae
Carcharocles auriculatus (Blainville 1818)
Zvonok and Удовиченко 2011
recombined as Carcharodon auriculatus
Isurus praecursor Thurmond and Jones 1981
Udovichenko 2009
Chondrichthyes - Lamniformes - Odontaspididae
Odontaspis winkleri Leriche 1905
Udovichenko 2009
Synodontaspis vincenti
Udovichenko 2009
Jaekelotodus aff. borystenicus
Udovichenko 2009
Hypotodus verticalis (Agassiz 1843)
Udovichenko 2009
see common names

Geography
Country:Ukraine State/province:Luhansk
Coordinates: 49.5° North, 39.1° East (view map)
Paleocoordinates:47.5° North, 33.5° East
Basis of coordinate:stated in text
Geographic resolution:small collection
Time
Period:Paleogene Epoch:Eocene
Stage:Lutetian 10 m.y. bin:Cenozoic 2
Key time interval:Lutetian
Age range of interval:47.80000 - 41.20000 m.y. ago
Stratigraphy
Stratigraphic resolution:group of beds
Stratigraphy comments: early Lutetian
Lithology and environment
Primary lithology: sandstone
Includes fossils?Y
Environment:marine indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:general faunal/floral analysis
Collection dates:2010–2013
Taxonomic list comments:Redeposited from past epochs:
Megasqualus sp.
Paraorthacodus sp.
Isurolamna cf. inflata
Anacoracidae indet.
Metadata
Also known as:Novopskov
Database number:120885
Authorizer:M. Uhen, E. Vlachos, P. Mannion Enterer:M. Uhen, E. Vlachos, P. Mannion
Modifier:P. Mannion Research group:vertebrate
Created:2011-11-23 03:14:01 Last modified:2021-04-08 02:42:30
Access level:the public Released:2011-11-23 03:14:01
Creative Commons license:CC BY
Reference information

Primary reference:
38603. G. Mayr and E. A. Zvonok. 2011. Middle Eocene Pelagornithidae and Gaviiformes (Aves) from the Ukrainian Paratethys. Palaeontology 54(6):1347-1359 [M. Uhen/M. Uhen/O. Alfaro]

Secondary references:

39898 I. G. Danilov, E. A. Zvonok, E. V. Syromyatnikova and N. I. Udovichenko. 2011. A new species of soft-shelled turtle (Trionychidae) from the Middle Eocene of Ukraine. Proceedings of the Zoological Institute RAS 315(4):399-411 [M. Uhen/M. Uhen/P. Holroyd]
76282 I. T. Kuzmin and E. A. Zvonok. 2021. Crocodylian assemblage from the middle Eocene Ikovo locality (Lugansk Province, Ukraine), with a discussion of the fossil record and geographic origins of crocodyliform fauna in the Paleogene of Europe. Geobios 65:7-27 [P. Mannion/P. Mannion/P. Mannion]
42367 G. Mayr and E. A. Zvonok. 2012. A new genus and species of Pelagornithidae with well-preserved pseudodentition and further avian remains from the middle Eocene of the Ukraine. Journal of Vertebrate Paleontology 32(4):914-925 [M. Uhen/M. Uhen/O. Alfaro]
72877 N. I. Udovichenko. 2009. Ichthyofauna and age of the Paleogene sands of Osinovo area, Lugansk region. Ископаемая фауна и флора Украины: палеоэкологический и стратиграфический аспекты. Сборник научных трудов Института геологических наук НАН Украины 255-261 [M. Uhen/M. Uhen]
39900 E. A. Zvonok, I. G. Danilov, E. V. Syromyatnikova and N. I. Udovichenko. 2010. About new findings of turtles in the Eocene of Ukraine. Modern Paleontology: Classical and New Methods. The Seventh All-Russian Scientific School for Young Scientists in Paleontology (4–6 October, 2010, Moscow, Russia). Abstracts 15-16 [M. Uhen/M. Uhen/M. Uhen]
64143 E. A. Zvonok, I. G. Danilov, E. V. Syromyatnikova and N. I. Udovichenko. 2013. Remains of Sea Turtles from the Ikovo Locality (Lugansk Region, Ukraine; Middle Eocene). Paleontological Journal 47(6):607-617 [E. Vlachos/E. Vlachos]
39895 E. A. Zvonok and P. P. Skutschas. 2011. On a Tomistomine Crocodile (Crocodylidae, Tomistominae) from the Middle Eocene of Ukraine. Paleontological Journal 45(6):661-664 [M. Uhen/M. Uhen]
72876 E. O. Zvonok and Н И Удовиченко. 2011. новом местонахождении палеогеновых позвоночных в Украине . Геологічні пам’ятки – яскраві свідчення еволюції Землі. Матеріали II Міжнародної науково-практичної конференції. - Каменець-Подільський 128-130 [M. Uhen/M. Uhen]
72293 Y. Zvonok, G. Mayr, and L. Gorobets. 2015. New material of the Eocene marine bird Kievornis Averianov et al., 1990 and a reassessment of the affinities of this taxon. Vertebrata PalAsiatica 53(7):238-244 [M. Uhen/M. Uhen/M. Uhen]
72873 E. O. Звонок. 2011. Середньо- і верхньоеоценові рептилії України: нові відомості про місцезнаходження й таксономічне розмаїття. Тези доповідей всеукраїнської наукової конференції «Значення комплексних літо- і біостратиграфічних досліджень під час пошуків нафти і газу» 2011:32-33 [M. Uhen/M. Uhen]