PBDB Collection

Monte Bolca (Canossa collection): Ypresian, Italy

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

unclassified

	Mesileia emiliae	Jaekel 1894
Reptilia - Palaeophiidae

	Archaeophis proavus n. gen., n. sp. Massalongo 1859		1 specimen
Reptilia - Anomalophidae

	Archaeophis bolcensis n. gen., n. sp. Massalongo 1859		1 specimen
	replaced by Anomalophis bolcensis
Actinopteri

	Acronuroides eocaenicus n. gen., n. sp. Bannikov et al. 2019	Bannikov et al. 2017
Actinopteri - Caproiformes - Antigoniidae

	Antigonia veronensis Sorbini 1983	Sorbini and Bottura 1988
Actinopteri - Labriformes - Labridae

	Paralabrus rossiae n. gen., n. sp. Bannikov and Zorzin 2019	Bannikov and Zorzin 2019

	Zorzinilabrus furcatus n. gen., n. sp. Bannikov and Bellwood 2017	Bannikov and Bellwood 2017
Actinopteri - Perciformes - Gobioidei

	Carlomonnius quasigobius n. gen., n. sp. Bannikov and Carnevale 2016	Bannikov and Carnevale 2016
Actinopteri - Perciformes

	Pavarottia maiseyi n. sp. Bannikov 2016	Bannikov 2016

	Psettopsis latellai n. sp. Bannikov 2005	Bannikov 2005
	recombined as Latellopsis latellai
Actinopteri - Acanthuriformes - Acanthuridae

	Tauichthys padremenini n. gen., n. sp. Tyler 1999	Tyler 1999

	Gazolaichthys vestenanovae n. gen., n. sp. Blot and Tyler 1990	Blot and Tyler 1990

	Lehmanichthys lessiniensis n. gen., n. sp. Blot and Tyler 1990	Blot and Tyler 1990

	Metacanthurus veronensis n. gen., n. sp. Blot and Tyler 1990	Blot and Tyler 1990

	Eorandallius n. gen. rectifrons (Agassiz 1835)	Blot and Tyler 1990
	original and current combination Naseus rectifrons

	Eorandallius elegans n. sp. Blot and Tyler 1990	Blot and Tyler 1990

	Protozebrasoma bloti n. gen., n. sp. Sorbini and Tyler 1998	Sorbini and Tyler 1998

	Tylerichthys milani n. sp. Blot and Tyler 1990	Blot and Tyler 1990

	Tylerichthys nuchalis (Agassiz 1835)	Blot and Tyler 1990
	original and current combination Naseus nuchalis

	Sorbinithurus sorbinii n. gen., n. sp. Tyler 1999	Tyler 1999

	Pesciarichthys punctatus n. gen., n. sp. Blot and Tyler 1990	Blot and Tyler 1990

	Acanthuroides massalongoi n. gen., n. sp. Blot and Tyler 1990	Blot and Tyler 1990

	Acanthurus gazolai	Blot and Tyler 1990

	Acanthurus gaudryi de Zigno 1887	Blot and Tyler 1990

	Metaspisurus emmanueli n. gen., n. sp. Blot and Tyler 1990	Blot and Tyler 1990

	Proacanthurus ovalis (Agassiz 1839)	Blot and Tyler 1990
	original and current combination Acanthurus ovalis

	Proacanthurus bonatoi n. sp. Blot and Tyler 1990	Blot and Tyler 1990

	Proacanthurus n. gen. tenuis (Agassiz 1835)	Blot and Tyler 1990
	original and current combination Acanthurus tenuis

	Proacanthurus elongatus n. sp. Blot and Tyler 1990	Blot and Tyler 1990
Actinopteri - Tetraodontiformes - Balistidae

	Oligobalistes robustus Danil'chenko 1960	Tyler and Santini 2002

	Balistomorphus orbiculatus (Heer 1865)	Tyler and Santini 2002

	Balistomorphus ovalis (Agassiz 1842)	Tyler and Santini 2002

	Balistomorphus spinosus (Agassiz 1842)	Tyler and Santini 2002
Actinopteri - Tetraodontiformes - Monacanthidae

	Monacanthidae indet.	Tyler and Santini 2002
Actinopteri - Tetraodontiformes - Moclaybalistidae

	Moclaybalistes danekrus n. gen., n. sp. Tyler and Santini 2002	Tyler and Santini 2002
Actinopteri - Tetraodontiformes - Ostraciidae

	Ostraciidae indet.	Tyler and Santini 2002

	Eolactoria sorbinii Tyler 1975	Tyler and Santini 2002
Actinopteri - Tetraodontiformes - Molidae

	Molidae indet.	Tyler and Santini 2002
Actinopteri - Tetraodontiformes - Aracanidae

	Proaracana dubia (Blainville 1818)	Tyler and Santini 2002
	original and current combination Balistes dubius
Actinopteri - Tetraodontiformes - Protobalistidae

	Spinacanthus cuneiformis (Blainville 1818)	Tyler and Santini 2002
	original and current combination Blennius cuneiformis

	Protobalistum imperiale (Massalongo 1857)	Tyler and Santini 2002
	recombined as Spinacanthus imperiale
Actinopteri - Tetraodontiformes - Triodontidae

	Triodontidae indet.	Tyler and Santini 2002
Actinopteri - Tetraodontiformes - Zignoichthyidae

	Zignoichthys oblongus (Zigno 1874)	Tyler and Santini 2002
	original and current combination Ostracion oblongus
Actinopteri - Tetraodontiformes - Bolcabalistidae

	Eospinus daniltshenkoi Tyler and Bannikov 1992	Tyler and Santini 2002

	Bolcabalistes varii Tyler and Sorbini 1998	Tyler and Santini 2002
Actinopteri - Tetraodontiformes - Tetraodontidae

	Tetraodontidae indet.	Tyler and Santini 2002

	Eotetraodon pygmaeus (Zigno 1887)	Tyler and Santini 2002
	original and current combination Tetrodon pygmaeus
Actinopteri - Tetraodontiformes - Diodontidae

	Pshekhadiodon parini Bannikov and Tyler 1997	Tyler and Santini 2002

	Heptadiodon echinus (Heckel 1853)	Tyler and Santini 2002
	original and current combination Enneodon echinus

	Zignodon fornasieroae n. gen., n. sp. Tyler and Santini 2002	Tyler and Santini 2002

	Prodiodon erinaceus (Agassiz 1844)	Tyler and Santini 2002
	original and current combination Diodon erinaceus

	Prodiodon tenuispinus (Agassiz 1833)	Tyler and Santini 2002
	original and current combination Diodon tenuispinus
Actinopteri - Tetraodontiformes - Triacanthidae

	Protacanthodes ombonii de Zigno 1887	Tyler and Santini 2002

	Protacanthodes nimesensis Tyler and Santini 2001	Tyler and Santini 2002
Actinopteri - Tetraodontiformes - Eoplectidae

	Eoplectus bloti Tyler 1975	Tyler and Santini 2002
Actinopteri - Lutjanidae

	Lessinia horrenda n. gen., n. sp. Bannikov and Zorzin 2014	Bannikov and Zorzin 2014
Actinopteri - Scombriformes - Scombridae

	Orcynus lanceolatus n. gen., n. sp. Agassiz 1835	Agassiz 1843
	recombined as Thunnus lanceolatus

	Orcynus latior n. sp. Agassiz 1835	Agassiz 1843
	recombined as Thunnus latior

	Thynnus bolcensis Agassiz 1835	Woodward 1901
Actinopteri - Zeiformes - Bajaichthyidae

	Bajaichthys elegans Sorbini 1983	Sorbini and Bottura 1988
Actinopteri - Ellimmichthyiformes - Paraclupeidae

	Eoellimmichthys superstes n. gen., n. sp. Marramà et al. 2019	Marramà et al. 2019
Actinopteri - Clupeiformes - Clupeoidei

	Eoalosa janvieri n. gen., n. sp. Marrama` and Carnevale 2018	Marrama` and Carnevale 2018
Chondrichthyes - Myliobatiformes

	Lessiniabatis aenigmatica n. gen., n. sp. Marramà et al. 2019	Marramà et al. 2019

	Promyliobatis gazolae n. gen., n. sp. (de Zigno 1885)	Jaekel 1894
	recombined as Promyliobatis gazolai
Chondrichthyes - Myliobatiformes - Urolophidae

	Urolophus crassicaudula	Jaekel 1894
Chondrichthyes - Myliobatiformes - Dasyatidae

	Trygon dezignii	Jaekel 1894
	synonym of Dasyatis

	Trygon muricata (Volta 1796)	Jaekel 1894
	recombined as Trygon muricatus
Chondrichthyes - Galeidae

	Pseudogaleus voltai n. gen., n. sp. Jaekel 1894	Jaekel 1894
Chondrichthyes - Zanobatidae

	Platyrhina egertoni (de Zigno 1878)	Jaekel 1894
	recombined as Plesiozanobatus egertoni
Chondrichthyes - Rajiformes - Platyrhinidae

	Platyrhina bolcensis Heckel 1851	Jaekel 1894
Chondrichthyes - Rajiformes - Rhinobatidae

	Rhinobatus primaerus	Jaekel 1894

	Rhinobatus dezignii (Heckel 1853)	Jaekel 1894
	recombined as Pseudorhinobatos dezignii
Chondrichthyes - Torpediniformes - Narcinidae

	Narcine molini n. sp. Jaekel 1894	Jaekel 1894
Chondrichthyes - Torpediniformes - Torpedinidae

	Platyrhina gigantea (de Blainville 1818)	Jaekel 1894
	recombined as Torpedo gigantea
Chondrichthyes - Carcharhiniformes - Triakidae

	Galeus cuvieri n. sp. Agassiz 1835	Jaekel 1894
	recombined as Galeorhinus cuvieri
Chondrichthyes - Carcharhiniformes - Carcharhinidae

	Alopiopsis plejodon Lioy 1865	Jaekel 1894
Chondrichthyes - Lamniformes - Odontaspididae

	Brachycarcharias lerichei (Casier 1946)	Marramà et al. 2019
Malacostraca - Isopoda - Sphaeromatidae

	Sphaeroma gasparellai De Angeli and Lovato 2009	Beschin et al. 2016
Angiospermae - Alismatales - Posidoniaceae

	Posidocea frickingeri n. gen., n. sp.	Gregor 1991
see common names

Geography

Country:	Italy
Coordinates:	45.6° North, 11.2° East (view map)
Paleocoordinates:	37.3° North, 10.3° East
Basis of coordinate:	estimated from map
Geographic resolution:	outcrop

Time

Period:	Paleogene	Epoch:	Eocene
Stage:	Ypresian	10 m.y. bin:	Cenozoic 2
Key time interval:	Ypresian
Age range of interval:	56.00000 - 47.80000 m.y. ago

Stratigraphy

Stratigraphic resolution:	group of beds
Stratigraphy comments: Initially assigned to the Lutetian. However, calcareous nannofossils attribute the classic Pesciara locality to the Discoaster sublodoensis Zone (NP14 or CP12) of uppermost Ypresian to lowermost Lutetian age. Larger foraminifera indicate the Alveolina dainellii Zone, and have limited abrasion so do not appear to be reworked. Therefore, biostratigraphy restricts the classic Pesciara locality to a narrow interval between the base of NP14 and the top of SBZ11, of late Ypresian age (Papazzoni et al., 2014). The lower-middle part of Monte Postale was also preliminarily assigned to SBZ11 of Ypresian age (Papazzoni & Trevisiani, 2009).

Lithology and environment

Primary lithology:	planar lamination,shelly/skeletal "limestone"
Includes fossils?	Y
Lithology description: "The fossil-bearing section consits of several calcareous units (Figure 20.2). Exceptionally well preserved fossils are derived from units of laminated sheet limestones unlain by reefal limestones. Fish and plant fossils can be found at five levels. The beds containing fish and plant fossils show a varved structure in which relatively thick biomicritic laminae alternate in a regular pattern with very thin micritic laminae, which are homogenous and slightly clayey. The thicker biomicritic laminae contain silt-sized particles and foraminifera, ostracods, molluscs, and small thin-shelled gastropods. These biomicritic laminae are gradational with the micritic laminae, producing a graded bedding texture for each pair of laminae. Finely disseminated pyrite and bitumen can be found in these laminated beds. (pg. 369)"
Environment:	lagoonal/restricted shallow subtidal

Taphonomy

Modes of preservation:	body,mold/impression,soft parts
Size of fossils:	macrofossils
Preservation of anatomical detail:	excellent
Abundance in sediment:	abundant
Articulated whole bodies:	many
Associated major elements:	many
Temporal resolution:	snapshot
Spatial resolution:	autochthonous

Collection methods and comments

Collection excludes:	all macrofossils
Collection methods:	selective quarrying,mechanical,survey of museum collection,observed (not collected)
Reason for describing collection:	taxonomic analysis
Museum repositories:	MCZ,MfN
Collection method comments: Repository: collection of Canossa (material later separated, some going to Museum fur Naturkunde and some to Museum of Comparative Zoology)

Metadata

Database number:	55239
Authorizer:	J. Head, P. Holroyd, M. Uhen	Enterer:	J. Head, P. Holroyd, M. Uhen
Modifier:	M. Clapham	Research group:	vertebrate
Created:	2005-09-30 14:06:02	Last modified:	2021-08-24 23:22:52
Access level:	the public	Released:	2005-09-30 14:06:02
Creative Commons license:	CC BY

Reference information

Primary reference:

14695.

A. Massalongo. 1859. Specimen photographicum animalium quorundam plantaumque fossilium agri veronensis [J. Head/J. Head/M. Uhen]

Secondary references:

38894		L. Agassiz. 1843. Recherches Sur Les Poissons Fossiles. Tome I (18me livraison) xxxii-188 [M. Carrano/M. Carrano/M. Clapham]
72922		A. F. Bannikov. 2005. A new species of Psettopsis (Perciformes, Monodactylidae) from the Eocene of northern Italy (Bolca) . Miscellanea Paleontologica n. 8. Studi e Ricerche sui Giacimenti Terziari di Bolca 11:141-148 [M. Uhen/M. Uhen]
73202		A. F. Bannikov. 2016. A new species of the percoid fish genus Pavarottia (Perciformes) from the Eocene of Bolca, northern Italy. Studi e ricerche sui giacimenti terziari di Bolca 17(14):5-11 [M. Uhen/M. Uhen]
73206		A. F. Bannikov and D. R. Bellwood. 2017. Zorzinilabrus furcatus, a new genus and species of labrid fish (Perciformes) from the Eocene of Bolca in northern Italy. Studi e ricerche sui giacimenti terziari di Bolca 18(15):5-14 [M. Uhen/M. Uhen]
73203		A. F. Bannikov and G. Carnevale. 2016. †Carlomonnius quasigobius gen. et sp. nov.: The first gobioid fish from the Eocene of Monte Bolca, Italy. Bulletin of Geosciences 91(1):13-22 [M. Uhen/M. Uhen/M. Uhen]
75762		A. F. Bannikov, J. C. Tyler, D. Arcila and G. Carnevale. 2017. A new family of gymnodont fish (Tetraodontiformes) from the earliest Eocene of the Peri-Tethys (Kabardino-Balkaria, northern Caucasus, Russia). Journal of Systematic Palaeontology 15(2):129-146 [M. Uhen/M. Uhen]
73199		A. F. Bannikov and R. Zorzin. 2014. A new genus and species of lutjanid fish (Perciformes) from the Eocene of Bolca, Northern Italy. Studi e ricerche sui giacimenti terziari di Bolca 40(12):15-22 [M. Uhen/M. Uhen]
73208		A. F. Bannikov and R. Zorzin. 2019. Paralabrus rossiae, a new genus and species of putative labroid fish (Perciformes) from the Eocene of Bolca in northern Italy. Studi e ricerche sui giacimenti terziari di Bolca 19(16):39-47 [M. Uhen/M. Uhen]
71455		C. Beschin, A. Busulini, G. Tessier and R. Zorzin. 2016. I crostacei associati a coralli nell’Eocene inferiore dell’area di Bolca (Verona e Vicenza, Italia nordorientale). Memorie del Museo Civico di Storia Naturale di Verona - 2. serie, Sezione Scienze della Terra 9:1-189 [M. Uhen/M. Uhen]
62349		J. Blot and J. C. Tyler. 1990. New genera and species of fossil surgeon fishes and their relatives (Acanturoidei, Teleostei) from the Eocene of Monte Bolca, Italy, with application of the Blot formula to both fossil and recent forms. Studi e ricerche sui giacimenti terziari di Bolca 6:13-92 [M. Uhen/M. Uhen]
40170		H. -J. Gregor. 1991. Ein neues fossiles Seegras - Posidocea frickingeri nov. gen. et spe. im Palaogen Oberitaliens (Verona). Documenta Naturae 65:1-11 [M. Uhen/M. Uhen]
30721		O. Jaekel. 1894. Die eocanen Selachier vom Monte Bolca 1-176 [M. Uhen/M. Uhen]
76015		G. Marramà, A. F. Bannikov, J. Kriwet and G. Carnevale. 2019. An Eocene paraclupeid fish (Teleosti, Ellimmichthyiformes) from Bolca, Italy: The youngest marine record of double-armoured herrings. Papers in Palaeontology 5(1):83-98 [M. Uhen/M. Uhen]
76013		G. Marramà, G. Carnevale, L. Giusberti, G. J. P. Naylor, and J. Kriwet. 2019. A bizarre eocene dasyatoid batomorph (elasmobranchii, Myliobatiformes) from the Bolca Lagerstätte (italy) reveals a new, extinct body plan for stingrays. Nature Scientific Reports 9:14087 [M. Uhen/M. Uhen]
72428		G. Marramà, A. Engelbrecht, G. Carnevale and J. Kriwet. 2019. Eocene sand tiger sharks (Lamniformes, Odontaspididae) from the Bolca Konservat- Lagerstätte, Italy: palaeobiology, palaeobiogeography and evolutionary significance. Historical Biology 31(2):102-116 [M. Uhen/M. Uhen]
67560		G. Marrama` and G. Carnevale. 2018. Eoalosa janvieri gen. et sp. nov., a new clupeid fish (Teleostei, Clupeiformes) from the Eocene of Monte Bolca, Italy. PalZ 92:107-120 [P. Holroyd/P. Holroyd]
47117		L. Sorbini and C. Bottura. 1988. Antigonia veronensis, an Eocene caproid form Bolca, Italy. Bollettino del Museo Civico di Storia Naturale de Verona 14:255-269 [M. Uhen/M. Uhen]
47118		L. Sorbini and C. Bottura. 1988. Bajaichthys elegans, and Eocene lampridiform from Bolca, Italy. Bollettino del Museo Civico di Storia Naturale de Verona 14:369-380 [M. Uhen/M. Uhen]
62352		L. Sorbini and J. C. Tyler. 1998. A new genus and species of Eocene surgeon fish (Acanthuridae) from Monte Bolca, Italy, with similarities to the Recent Zebrasoma. Studi e ricerche sui giacimenti terziari di Bolca 7:7-19 [M. Uhen/M. Uhen]
14688		C. M. Tang. 2002. Monte Bolca: An Eocene fishbowl. Exceptional fossil preservation, a unqiue view on the evolution of marine life 365-377 [J. Head/J. Head]
62348		J. C. Tyler. 1999. A new genus and species of surgeon fish (Acanthuridae) with four dorsal-fin spines from the Eocene of Monte Bolca, Italy. Studi e ricerche sui giacimenti terziari de Bolca 257-268 [M. Uhen/M. Uhen]
72451		J. C. Tyler and F. Santini. 2002. Review and reconstructions of the tetraodontiform fishes from the Eocene of Monte Bolca, Italy, with comments on related Tertiary taxa. Studi e Ricerche sui Giacimenti Terzari di Bolca. Museo Civico di Storia Naturale di Verona 9:47-119 [M. Uhen/M. Uhen]
45833		A. S. Woodward. 1901. Catalogue of Fossil Fishes in the British Museum (Natural History), Part IV 1-636 [M. Clapham/P. Vazquez/M. Clapham]