PBDB Collection

São José de Itaborai: Early/Lower Eocene, Brazil
collected by Carlos de Paula Couto, Ney Vidal and others 1948-1950

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Mammalia - Jaskhadelphyidae

	Minusculodelphis minimus n. sp. Paula Couto 1962	Paula Couto 1962
	MNRJ 2342-V (holotype), 2503-V (6 measurements)

	Minusculodelphis modicum n. sp. Oliveira et al. 2016	Oliveira et al. 2016	6 specimens
	DGM 921a-M, an incomplete right dentary with complete m1–4 (holotype). MCT 2815-M, a complete right M3; MCN-PV 1792, a complete right M3; MN 2479-V, an incomplete right maxilla with complete M2 and incomplete M3; DGM 921b-M, an incomplete right dentary with roots of m1, m2, portion of m4, and with complete m3; DGM921c-M, a complete left femur.
Mammalia - Stagodontidae

	Eobrasilia coutoi Simpson 1947	Paula Couto 1962
	AMNH 39424 (holotype); Marshall (1984) adds the following specimens: DGM 919-M, DGM 896-M(a) (7 measurements)
Mammalia - Polydolopidae

	Polydolopidae indet. Ameghino 1897	Ladaveze and Muizon 2010
Mammalia - Bonapartheriidae

	Epidolops ameghinoi n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	DGM 321-M - holotype (almost complete but much crushed and flattened skull, lacking the upper part of the braincase and the part posterior to the basisphenoid, as well as the anterior end of the premaxillae; almost complete mandible, lacking the anterior end of the incisor border); referred material: DGM 170-197-M and 207-209-M (fragmentary right and left lower jaws with and without teeth), DGM 198-206-M (partial right and left maxillae with teeth), MNRJ 1372-1374-V, 1376-1378-V, 1381-1392-V, 1394-V and 1395-V, 1397-V to 1405-V (incomplete right and left lower jaws with and without teeth), 1407-V to 1416-V (portions of right and left maxillae with and without teeth), 1417-V to 1419-V (partial right P3), 1420-V (incomplete left P3), 1421-V (left M2), 1422-V (right Ml), 1423-V (anterior part of left M1), 1424-V (right I?2), AMNH 49806 to 49809, 49811 and 49813 (fragmentary lower jaws with teeth)

	Epidolops gracilis n. sp. Paula Couto 1952	Paula Couto 1952
	synonym of Epidolops ameghinoi
	DGM 188-M - type (fragment of right lower jaw); paratype: MNRJ 1400-V (partial left lower jaw); referred material: DGM 191-M (part of left lower jaw with P3, M1 present), 198-M (portion of left maxilla with P3, M1-2 present), MNRJ 1390-V and 1391-V (anterior parts, respectively, of right lower jaw with roots of P2 and with P3 present, and of left lower jaw with P2-3 present), 1412-V (part of right maxilla with P3, M1-2 present), 1404-V (fragment of left maxilla with P3 present), 1411-V (portion of right maxilla with P3 present), AMNH 49809, 49810, 49812 to 49814 and 49860 (lower jaw fragments with teeth)
Mammalia

	Zeusdelphys complicatus n. gen., n. sp. Marshall 1987
	MCT 2830-M - holotype (ex- DGM 896a-M, a left ?M1)

	aff. Bobbschaefferia sp. Paula Couto 1970
	MCN-PV 1797 (an incomplete right maxilla) and MCT 2816-M (right m3)

	Schaefferia fluminensis n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	replaced by Bobbschaefferia fluminensis
	MNRJ 1350-V - holotype (portion of left mandibular ramus); paratypes now comprise the type of Guggenheimia crocheti: DGM 314-M (incomplete left mandibular ramus) and 315 (fragment of left manibular ramus) (7 measurements)
Mammalia - Pediomyidae

	cf. Pediomyidae indet. (Simpson 1927)	Oliveira and Goin 2015
Mammalia - Sparassodonta - Borhyaenidae

	Borhyaenidae indet. Savage 1951	Ladaveze and Muizon 2010
Mammalia - Sparassodonta

	? Arminiheringia sp. Ameghino 1902	Paula Couto 1952
	MNRJ 1344-V (large left lower molar)
	= Nemolestes brasiliensis Rangel et al. 2023	Rangel et al. 2023
	MN 1344-V, a left m4 is assigned as hypodigm of Nemolestes brasiliensis sp. nov. The m4 of Nemolestes brasiliensis (MN 1344-V) was identified previously as Arminiheringia sp. by Paula Couto (1961), and cf. Nemolestes sp. by Marshall (1978) and Bergqvist et al. (2009). The m2 (MCT.M.657, holotype) was previously referred to as borhyaenid indet. by Paula Couto (1970). MCT.M.657 was excluded from Patene based on its much larger size (about 2 mm larger than the largest molar of Patene simpsoni; see Rangel et al. 2019 and Table 2). Based on the evident size difference (Fig. 4) and described morphology, MCT.M.657 and MN 1344-V are here assigned to Nemolestes brasiliensis sp. nov.
Mammalia - Sparassodonta - Hathlyacynidae

	Hathlyacynidae indet. (Ameghino 1894)	Ladaveze and Muizon 2010
	corrected as Hathlyacininae
Mammalia - Sparassodonta

	Patene simpsoni n. sp. Paula Couto 1952	Paula Couto 1952
	MNRJ 1331-V - type (incomplete right maxilla); paratypes: DGM 324-M (fragment of right lower jaw with alveolus of P2 and roots of P3, anterior root of M1, talonid and anterior root of M2, with M3, and with roots of M4 present), MNRJ 1332-V (right M4), 1335-V (right Ml or M2); referred material: MNRJ 1336-V to 1343-V and 1428-V (eight left lower molars, and one right lower molar, reduced to the trigonid) and DGM 331-M (part of left maxilla)

	Ischyrodidelphis castelllanosi n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	synonym of Patene simpsoni
	MNRJ 1351-V - holotype (median fragment of right mandibular ramus); paratypes: MNRJ 1352 and 1353 (right m2s), 1354 (left m2) and 1425 (right p3) (4 measurements)
Mammalia

	Australidelphia indet. Szalay 1982	Ladaveze and Muizon 2010
	"stem Australidelphia"
Mammalia - Microbiotheria - Microbiotheriidae

	Microbiotheriidae indet. (Ameghino 1887)	Ladaveze and Muizon 2010

	Mirandatherium alipioi Paula Couto 1952	Paula Couto 1962
	=Mirandaia ribeiroi, replacement name following Paula Couto (1952) (3 measurements)

	Mirandaia ribeiroi n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	replaced by Mirandatherium alipioi
	MNRJ 1359-V - holotype (median fragment of right mandibular ramus); paratypes: DGM 361 and AMNH 49859 (two partial right mandibular rami); referred material: MNRJ 1362 (median fragment of right lower jaw), 1363 (part of right lower jaw), AMNH 49802 (median portion of right lower jaw), DGM 360-M (posterior part of right mandibular ramus) and 362-M (posterior fragment of right lower jaw)
Mammalia - Dasyuromorphia - Malleodectidae

	Gaylordia mater n. sp. Oliveira and Goin 2015	Oliveira and Goin 2015
	Holotype. DGM 810-M, a partial right dentary with complete m2-4. Hypodigm. The type and MCT 2383-M (DGM 644-M), incomplete left dentary with m1-2 and m4; MCT 2384-M (DGM 645-M), incomplete left dentary with p3, m1-2, incomplete m3 and complete m4; MCT 2386-M, incomplete right dentary with m3 and alveoli of p3 (only the posterior alveolus is complete), m1, m2 and m4; MCT 2387-M, incomplete right dentary with complete p3 and m1, alveoli of p2, and m2; MCT 2774-M, complete left m4; MCT 2775- M, incomplete right m3; MCT 2776-M, complete right m3; MCT 2777-M, complete right M2; MCT 2778-M, incomplete left M2; MCT 2779-M, incomplete right M2; MCT 2780-M, complete left M3; MCT 2781-M, incomplete right M3.

	Gaylordia macrocynodonta n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	DGM 329-M - holotype (anterior part of skull and mesial fragment of left mandibular ramus); referred material: MNRJ 1366-V (median fragment of left lower jaw), 1357-1358 (left and right mandivular rami), AMNH 49801 (median portion of right lower jaw) and DGM 186-M (left mandibular ramus) (9 measurements)

	Xenodelphis doelloi n. sp. Paula Couto 1962	Paula Couto 1962
	synonym of Gaylordia macrocynodonta
	MNRJ 2504-V (holotype) (4 measurements)
Mammalia

	Monodelphopsis travassosi n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	MNRJ 1365-V - holotype (fragment of left lower jaw) (3 measurements)
Mammalia - Herpetotheriidae

	cf. Herpetotheriidae indet. Trouessart 1879	Oliveira and Goin 2015
Mammalia

	Gashternia carioca n. sp. Goin and Oliveira 2007	Goin and Oliveira 2007	1 specimen
	Holotype only. MCN-PV 1801, a fragment of left maxilla with complete P3 and M1, and roots M2.
Mammalia - Paucituberculata

	Paucituberculata indet. Ameghino 1894	Ladaveze and Muizon 2010

	Riolestes capricornicus n. gen., n. sp. Goin et al. 2009	Goin et al. 2009
	MCN-PV 1790 - holotype (an isolated m1)
Mammalia - Protodidelphidae

	Protodidelphidae indet. Marshall 1987	Ladaveze and Muizon 2010
	= Bergqvistherium primigenia n. gen., n. sp. Carneiro 2018	Carneiro 2018
	MCT 4387-M - holotype

	Periprotodidelphis bergqvistae n. gen., n. sp. Oliveira and Goin 2011
	DGM 806-M (left M?3)

	Carolocoutoia ferigoloi n. gen., n. sp. Goin et al. 1998
	MCN-PV 1802 - holotype (upper left molar, M3?)

	Guggenheimia brasiliensis n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	DGM 297-M - holotype (incomplete left mandibular ramus) (3 measurements)

	Guggenheimia crocheti n. sp. Oliveira and Goin 2011
	DGM 314-M - type (incomplete left dentary); hypodigm: DGM 315-M (incomplete left dentary), MNRJ 2899(a)-V (right m3), MCT 2783-M (left m3) and MCT 2782-M (left M1)

	Robertbutleria mastodontoidea n. gen., n. sp. Marshall 1987
	recombined as Protodidelphis mastodontoides
	DGM 896-M - holotype (left M1); referred material: MCT 2769-M (ex DGM 896b-M; complete left M2), MCN-PV 1799 (complete left dP3), MCN-PV 1806 (complete left M3), MNRJ 2897(b)-V (nearly complete right M2), MNRJ 2897(c)-V (incomplete right M2), MNRJ 2897(d)-V (incomplete left M3), AMNH 49858 (incomplete left dentary with p2-m2), MCT 2817-M (nearly complete left m2), MCT 2818-M (nearly complete left m2), MCT 2819-M (incomplete left m2), MNRJ 1371-V (complete left m1), MCN-PV 1819 (nearly complete left m3), MCN-PV 1816 (a nearly complete left m3), MCN-PV 1817 (incomplete left m3), MCN-PV 1820 (complete left m4)

	Protodidelphis vanzolinii n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	DGM 271-M - holotype (facial and palatal portion of an incomplete skull); paratype: DGM 250-M (right mandibular ramus); referred specimens: DGM 319-M (left mandibular ramus), 303-M (fragment of right maxilla), AMNH 49857 (right mandibular ramus), MNRJ 1371-V (left m1 or m2), 1426-V (left m1), 1427-V (right m3), AMNH 49803 (right m2) and 49858 (portion of left mandibular ramus) (6 measurements)
Mammalia - Didelphimorphia - Didelphidae

	Didelphidae indet. Gray 1821	Paula Couto 1952
	MNRJ 1367-V and1368-V (two very small posterior fragments of right lower jaws)
Mammalia - Sternbergiidae

	Carolopaulacoutoia itaboraiensis Paula Couto 1970	Oliveira and Goin 2015

	Didelphopsis cabrerai n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	MNRJ 1429-V - holotype (left maxilla); paratypes: DGM 243-M (right mandibular ramus) and MNRJ 1345-V (right m4); referred material: MNRJ 1346-V (m3?) and 1347-V (m4) (6 measurements)
Mammalia - Derorhynchidae

	Diogenesia brevirostris n. gen., n. sp. Oliveira et al. 2021	Oliveira et al. 2021
	Holotype: DGM 650-M, incomplete right dentary with alveoli of p1-2, roots of p3-m1, and complete m2-4.

	Derorhynchus singularis n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	MNRJ 1348-V - holotype (incomplete right mandibular ramus); paratype: AMNH 49826 (fragment of right mandibular ramus) (5 measurements)
Mammalia - Peradectidae

	Peradectidae indet. Crochet 1979	Ladaveze and Muizon 2010
Mammalia

	Procaroloameghinia pricei n. gen., n. sp. Marshall 1982
	DGM 805-M and 924-M - holotype (two pieces of the same right dentary) (7 measurements)

	Austropediomys marshalli n. gen., n. sp. Carneiro et al. 2018	Carneiro et al. 2018
	Holotype DGM 808-M, incomplete left maxilla with M2–4; MCT 2798-M, incomplete left M1; MCT 2799-M, complete left M3; MCT 2800- M, incomplete right M4; MCT 4386-M, left DP3

	Marmosopsis juradoi n. sp. Paula Couto 1962	Paula Couto 1962
	All the specimens housed at MNRJ: MNRJ 2343-V (holotype), 2344-V, 2345-V, 2346-V, 2348-V:2350-V, 2356-V:2357-V, 2359-V:2360-V, 2460-V: 2473-V, 2434-V (this record must be a mistyping for 2474 from the order of the hypodigm report), 2475-V:2481-V (MNRJ 2482-V is reported in the measurements section. However, it is missing in the hypodigmia section. Among the specimens reported as examined, only MNRJ 2481-V shows upper M1-4 present, so it is very likely that 2482 is but a mistyping for 2481. This requires confirmation by direct measurements of those specimens housed at MNRJ), 2483-V:2487-V, 2495-V:2502-V. (4 measurements)
Mammalia - Pucadelphyidae

	Itaboraidelphys camposi Marshall and de Muizon 1984	Marshall and de Muizon 1984	7 specimens
	Holotype (DGM 817-M, left partial dentary with p3-m4); DGM 804-M, right dentary fragment with m2-3 and m1, m4 alveoli; DGM 814-M, right dentary fragment with m1-4; DGM 923-M, right dentary fragment with p2-m1 and m2 alveoli; DGM 926-M, maxillary fragment with M2-3 and M4 alveolus; MNRJ 2878-V a, isolated M2; MNRJ 2878-V b, isolated M3
Mammalia - Astrapotheria - Trigonostylopidae

	Trigonostylops apthomasi n. sp. Price and Paula Couto 1950	Paula Couto 1952
	DGM 154-M - type (right m1?); paratypes: DGM 155-M and 156-M (molars); numerous referred material
	= Tetragonostylops apthomasi Price and Paula Couto 1950	Paula Couto 1963
	All the material reported in Paula Couto (1952) is referred to Tetragonostylops apthomasi in Paula Couto (1963)

	Trigonostylops sp. Ameghino 1897	Paula Couto 1952
	MNRJ 1823-V-1839-V (small teeth)
	= Tetragonostylops apthomasi Price and Paula Couto 1950	Paula Couto 1963
	MNRJ 1823-V-1839-V (Explicitly stated in the 'Hypodigma' section)
Mammalia - Notoungulata

	Notoungulata indet. Roth 1903	Billet and de Muizon 2013
	MNHN-F-BRD 23 (left isolated petrosal)
Mammalia - Notoungulata - Henricosborniidae

	Nanolophodon tutuca n. gen., n. sp. Castro et al. 2021	Castro et al. 2021
	Holotype: MCT 4419M, a fragment of left maxilla with P3-M2 (moderate to advanced wear); MCT 4400M, isolated right P2; MCT 4397M, isolated left P3; MCT 4396M, isolated left P4; MCT 4390M, isolated right M1; MCT 4388M, isolated right M2; MCT 4389M, isolated right M2; MCT 4420M, isolated right M2; MCT 4421M, isolated left M3; MCT 4403M, isolated left lower molar; MCT 4422M, fragment of left jaw with dp2, dp3, dp4 and m1 (in alveolus).
Mammalia - Notoungulata - Oldfieldthomasiidae

	Homalostylops atavus n. sp. Paula Couto 1954	Paula Couto 1954
	synonym of Itaboraitherium atavum
	MNRJ 1992-V - type (posterior fragment of a right mandibular ramus with m1-3); paratypes: MNRJ 1990-V (2 right upper molars) and MNRJ 2245-V (one right and 3 left upper molars

	Henricosbornia magellanica n. sp. Price and Paula Couto 1950	Paula Couto 1952
	recombined as Colbertia magellanica
	DGM 152-M - type (left m3); paratype: DGM 153-M (fragment of left lower jaw); referred material: MNRJ 1865-V to 1909-V and 1944-V to 1977-V (partial right and left lower jaws with cheek teeth), 1910-V to 1925-V (fragmentary left and right maxillae with cheek teeth), 1926-V and 1927-V (series of associated left and right P2-M3 and P2-M2, respectively), 1928-V to 1943-V (isolated right and left upper cheek teeth), 1978-V to 1987-V (isolated right and left lower cheek teeth), DGM 236-M, 242-M, 252-M, 254-M, 256-M to 259-M, 264-M to 266-M, 269-M, 272-M, 274-M to 276-M, 279-M to 283-M, 285-M and 286-M, 289-M, 291-M, 293-M, 295-M, 298-M and 299-M, 316-M, 322-M, 377-M to 382-M (incomplete right and left lower jaws with cheek teeth), 310-M (almost complete mandible with right P2, P4-M3, and left P2-M3 [cast AMNH 49852]), 284-M, 288-M, 302-M, 306-M, 357-M, 372-M to 376-M (partial right and left maxillae with cheek teeth), 370-M and 371-M (fragmentary skulls with right P2-M3 and left P2-4, and right P1-Mi and left P4-M3, respectively), AMNH 49840, 49867 to 49877 (incomplete right and left maxillae with cheek teeth), 49841 to 49846, 49864 to 49866, 49878 to 49890 (partial right and left lower jaws with cheek teeth)

	Camargomendesia pristina n. gen., n. sp.	Bergqvist 2008	28 specimens
Mammalia - Xenungulata - Carodniidae

	Carodnia vieirai n. sp. Paula Couto 1952	Paula Couto 1952
	DGM 333-M - type (fragments of right and left maxillae; fragment of the occiput with the condyles; nearly complete mandible; all the cervical and the first two dorsal vertebrae; fragmentary first left rib; proximal fragment of left scapula; part of the bones of the anterior and posterior extremities, and other fragmentary skeletal remains); referred material: DGM 334-M (mandible of an immature individual), 335-M (fragment of right maxilla and anterior part of the skull of another individual), , 336-M- and 337-M (various skeletal remains), AMNH 49828 (left m3), MNRH 1843-V-1857-V (teeth)
Mammalia - Cingulata - Dasypodidae

	Riostegotherium yanei n. sp. Oliveira and Bergqvist 1998	Oliveira and Bergqvist 1998
	MCN-PV 1774 (holotype) and 1775, pelvic osteoderms; MCN-PV 1776, 1778, 1779, MCT 2081 M, MLP 75-XII-26-1, MLP 75-XII-26-2, isolated moveable osteoderms; MCN-PV 1777, caudal osteoderm
Mammalia

	Palaeocladosictis mosesi n. gen., n. sp. Paula Couto 1961	Marshall 1981	1 specimen
	MNRJ 2671-V, DP4 or P4
Mammalia - Panameriungulata - Protolipternidae

	Protolipterna ellipsodontoides n. gen., n. sp. Cifelli 1983	Cifelli 1983
	DNPM LE444A (cast, AMNH 109542), mandible with right C, alveoli of right and left lower P1-2, and with right and left P3-M3; associated right maxillary fragment with M1-3; hundreds of other specimens

	Asmithwoodwardia scotti n. sp. Paula Couto 1952	Paula Couto 1952
	DGM 358-M - type (badly crushed but nearly complete skull, with P3, M1-3 and left P3-4, M1-3, accompanied by its nearly complete mandible with all the teeth present, the tip of an upper C, and the greater part of the axis); paratype: AMNH 49817 (fragment of right lower jaw)

	Ernestokokenia parayirunhor n. sp. Paula Couto 1952	Paula Couto 1952
	recombined as Miguelsoria parayirunhor
	MNRJ 1468-V - type (almost complete right lower jaw); paratypes: DGM 305-M (part of left upper jaw) and 249-M (posterior part of left lower jaw); referred material: DGM 330-M (fragment of lower jaw; with the posterior part of the symphysis, and right C, P2-4, M1l2, and left C, P2 present), 246-M (part of lower jaw, with the posterior part of the symphysis, and the right P4, M1-3 present), 255-M (part of right lower jaw, with P4, M13 present), 290-M (part of left lower jaw, with M2-3 present), 296-M (part of left lower jaw, with M2-3 present), MNRJ 1469-V (anterior part of left lower jaw, with Pc-M1 present), 1470-V (part of left lower jaw, with Ml_2 present), 1466-V (part of left lower jaw, with M1-2 present), 1467-V (part of left lower jaw, with M2-3 present), 1861-V (part of left lower jaw, with M2 present), 1862-V (left M1,very worn) and AMNH 49815 (portion of left lower jaw, with M1.2 present)
Mammalia - Panameriungulata - Proterotheriidae

	Anisolambda prodromus n. sp. Paula Couto 1952	Paula Couto 1952
	recombined as Paranisolambda prodromus
	DGM 262-M - type (right lower jaw of a very young individual); paratypes: DGM 304-M (part of a right upper jaw), 239-M (partial left lower jaw) and 273-M (fragment of right and left upper jaw); referred material: MNRJ 1479-V (left M?2), 1493-V, 1494-V (right M?2), 1495-V, 1496-V (left M?2), 1858-V (fragment of left lower jaw with DM3-4 and Ml present), 1859-V (left M3), DGM 238-M (fragment of right lower jaw with M1-2 present), 307-M (partial left upper jaw with P2, DM3-4, M1 present), 310-M (fragment of right upper jaw with M2-3, and incomplete posterior part of the skull)
Mammalia - Panameriungulata - Macraucheniidae

	Victorlemoineia prototypica n. sp. Paula Couto 1952	Paula Couto 1952
	MNRJ 1470-V - type (right m2 or 3); paratype: MNRJ 1471-V (left m3); referred material: MNRJ 1472-V-1474-V (three left upper molars), 1475-V-1478-V (four right upper molars), 1480-V (fragment of left lower jaw with P4 present), 1481-V (right P?3), 1482-V-1486-V (right lower molars), 1487-V, 1488-V (left P3?s), 1489-V, 1490-V, 1492-V (left lower molars) and DGM 268-M (fragment of left lower jaw)
Mammalia - Panameriungulata - Didolodontidae

	Didolodontidae indet. Simpson 1934	Paula Couto 1952
	MNRJ 1841-V and 1842-V (left and right p4)

	Ernestokokenia protocenica n. sp. Paula Couto 1952	Paula Couto 1952
	recombined as Ricardocifellia protocenica
	MNRJ 1430-V - type (fragment of left maxilla); paratype: MNRJ 1431-V (part of left lower jaw); referred material: MNRJ 1432-V (small part of a right lower jaw with M3), 1433-V, 1434-V (respectively left and right M3), 1435-V, 1436-V, 1438-V (all right M2), 1439-V (left M2), 1440-V-1442-V (all right M1), 1443-V, 1444-V (left Ml), 1445-V (right P?4), 1446-V-1448-V (left P?4), 1449-V, 1450-V (left P?s), 1451-V, 1452-V (left M3), 1453-V-1455-V (right M2), 1456-V (left M2), 1457-V (right Ml), 1458-V, 1459-V (left Ml), 1460-V, 1461-V (left P3), 1462-V (right M2-3), AMNH 49819 (partial right lower jaw with M2-3), 49820 (left P3), 49821 (left P?s), 49822 (left P?4), 49823 (four M1, unassociated), 49824 (two left M3 and one right M3), 49825 (two right M2 and one left M2) 49827 (one left M2 and one left M3), 49829 (three left Ml)

	Lamegoia conodonta n. gen., n. sp. Paula Couto 1952	Paula Couto 1952
	MNRJ 1463-V - holotype (left m2?); paratypes: MNRJ 1464-V and 1465-V (left m1 and right m2 respectively)
Reptilia

	Eocaiman itaboraiensis n. sp. Pinheiro et al. 2013	Pinheiro et al. 2013
	MCT 1791-R - type (a well-preserved small left anterior dentary fragment, broken at the level of the eleventh alveolus and bearing four fully preserved teeth); referred specimens: MCT 1792-R (small left anterior dentary fragment with no teeth); MCT 1793-R (small left anterior dentary fragment with no preserved teeth); tentatively referred specimen: MCT 1794-R (an isolated small blunt tooth [root and crown]). The specimens referred to E. itaboraiensis sp. nov. have been kept associated in the MCT collection but lack information regarding their precise provenance. However, the different colouration of these fossils and the carbonate matrix (varying from yellowish to greyish-white) suggests these specimens were recovered from different S2 fissures infills.

	Aves informal sp. A (Linnaeus 1758)	Mayr et al. 2011
	MN 4115V (right carpometacarpus)

	Aves informal sp. B (Linnaeus 1758)	Mayr et al. 2011
	MN 4119−V (distal left tibiotarsus)

	Aves informal sp. C (Linnaeus 1758)	Mayr et al. 2011
	MN 4116−V (distal right tibiotarsus)

	Aves informal sp. D (Linnaeus 1758)	Mayr et al. 2011
	MN 4117−V (distal left tibiotarsus). An additional fragmentary distal end of left tarsometatarsus (MN 4120−V) may belong to this species or species E.

	Aves informal sp. E (Linnaeus 1758)	Mayr et al. 2011
	MN 4118−V (distal left tibiotarsus). An additional fragmentary distal end of left tarsometatarsus (MN 4120−V) may belong to this species or species D.
Reptilia - Gruiformes

	Itaboravis elaphrocnemoides n. gen., n. sp. Mayr et al. 2011	Mayr et al. 2011
	MN 4114V - holotype (left coracoid); referred specimens: MN 4113V (right humerus) and MN 4121V (distal end of left humerus)
Reptilia - Gruiformes - Phorusrhacidae

	Paleopsilopterus itaboraiensis n. gen., n. sp. Alvarenga 1985	Alvarenga 1985
	MNRJ-4040-V - holotype (proximal portion of the right tarsometatarsus); paratypes: DGM-1431-R (left and right tibiotarsus, proximally deformed and incomplete)
Reptilia - Cuculiformes - Cuculidae

	Eutreptodactylus itaboraiensis n. gen., n. sp. Baird and Vickers-Rich 1997	Baird and Vickers-Rich 1997
	MNB 4083v - holotype [now lost - only a cast and original illustrations remain] (Incomplete distal fragment of a right tarsometatarsus)
see common names

Geography

Country:	Brazil	State/province:	Rio de Janeiro	County:	Cabucu
Coordinates:	22.8° South, 42.9° West (view map)
Paleocoordinates:	25.6° South, 31.7° West
Basis of coordinate:	stated in text
Altitude:	19 meters
Geographic resolution:	local area

Time

Period:	Paleogene	Epoch:	Eocene
Stage:	Ypresian	10 m.y. bin:	Cenozoic 2
Key time interval:	Early/Lower Eocene	Mammal zone:	Itaboraian
Age range of interval:	56.00000 - 47.80000 m.y. ago

Stratigraphy

Formation:	Itaboraí
Stratigraphic resolution:	group of beds
Stratigraphy comments: Late Paleocene marl fillings in Early Eocene karst Itaboraian SALMA Middle sequence (S2) of the Itaborai Basin sediments Oliveira and Goin argue that it is more likely to be late Paleocene-early Eocene in age Early Eocene (Itaboraian SALMA); 53-50 Ma according to Woodburne et al. (2014) and Goin et al. (2018).

Lithology and environment

Primary lithology:	marl
Includes fossils?	Y
Lithology description: Fissure filling deposits with marls and collapsed breccias whose composition is similar to the gray limestone facies, although poorly consolidated, and transported into these cavities by heavy rains and gravitational flows
Environment:	fissure fill

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils,mesofossils

Collection methods and comments

Reason for describing collection:	taxonomic analysis
Collectors:	Carlos de Paula Couto, Ney Vidal and others	Collection dates:	1948-1950
Collection method comments: Quarry under the management of the Companhia Nacional de Cimento Portland (Maui)

Metadata

Also known as:	Companhia Nacional de Cimento Portland quarry, 1968 fissure
Database number:	118314
Authorizer:	P. Mannion, C. Jaramillo, M. Uhen, R. Butler, J. Carrillo, J. Zijlstra	Enterer:	P. Mannion, K. Pino, M. Uhen, G. Ballen, J. Zijlstra, R. Butler, G. Varnham
Modifier:	G. Varnham	Research group:	vertebrate
Created:	2011-10-12 01:43:19	Last modified:	2022-05-13 07:44:11
Access level:	the public	Released:	2011-10-12 01:43:19
Creative Commons license:	CC BY

Reference information

Primary reference:

42973.

E. V. Oliveira and F. J. Goin. 2011. A reassessment of bunodont metatherians from the Paleogene of Itaborai (Brazil): Systematics and the age of the Itaborian SALMA. Revista Brasileira de Paleontologia 14(2):105-136 [P. Mannion/P. Mannion]

Secondary references:

38164		H. Alvarenga. 1985. Um novo Psilopteridae (Aves: Gruiformes) dos sedimentos Terciários de Itaboraí, Rio de Janeiro, Brasil. Congresso Brasileiro de Paleontologia 8:17-20 [P. Mannion/P. Mannion]
38103		R. F. Baird and P. Vickers-Rich. 1997. Eutreptodactylus itaboraiensis gen. et sp. nov., an early cuckoo (Aves: Cuculidae) from the Late Paleocene of Brazil. Alcheringa 21(2):123-127 [P. Mannion/P. Mannion]
73183		R. M. D. Beck. 2017. The Skull of Epidolops ameghinoi from the Early Eocene Itaboraí Fauna, Southeastern Brazil, and the Affinities of the Extinct Marsupialiform Order Polydolopimorphia. Journal of Mammalian Evolution 24:373-414 [P. Mannion/M. Kouvari/P. Mannion]
57767		L. P. Bergqvist. 2008. Postcranial skeleton of the Upper Paleocene (Itaboraian) "Condylarthra" (Mammalia) of Itaboraí Basin, Brazil. Mammalian Evolutionary Morphology. A Tribute to Frederick S. Szalay 107-133 [J. Zijlstra/J. Zijlstra]
45342		G. Billet and C. de Muizon. 2013. External and internal anatomy of a petrosal from the late Paleocene of Itaboraí, Brazil, referred to Notoungulata (Placentalia). Journal of Vertebrate Paleontology 33(2):455-469 [P. Mannion/P. Mannion]
67168		L. M. Carneiro. 2018. A new protodidelphid (Mammalia, Marsupialia, Didelphimorphia) from the Itaboraí Basin and its implications for the evolution of the Protodidelphidae. Anais da Academia Brasileira de Ciências [P. Mannion/P. Mannion]
81704		L. M. Carneiro, E. V. Oliveira, and F. J. Goin. 2018. Austropediomys marshalli gen. et sp. nov., a new Pediomyoidea (Mammalia, Metatheria) from the Paleogene of Brazil: paleobiogeographic implications. Revista Brasileira de Paleontologia 21(2):120-131 [P. Mannion/G. Varnham]
81657		L. O. R. Castro, D. A. García-López, L. Paglarelli Bergqvist and H. I. De Araújo-Júnior. 2021. A new basal Notoungulate from the Itaboraí Basin (Paleogene) of Brazil. Ameghiniana 58(3):272-288 [P. Mannion/G. Varnham]
57766		R. Cifelli. 1983. The origin and affinities of the South American Condylarthra and early Tertiary Litopterna (Mammalia). American Museum Novitates 2772:1-49 [J. Zijlstra/J. Zijlstra]
42837		F. J. Goin, A. M. Candela, M. A. Abello and E. V. Oliveira. 2009. Earliest South American paucituberculatans and their significance in the understanding of ‘pseudodiprotodont’ marsupial radiations. Zoological Journal of the Linnean Society 155:867-884 [P. Mannion/P. Mannion]
57765		F. J. Goin and E. V. Oliveira. 2007. A new species of Gashternia (Marsupialia) from Itaboraí (Brazil). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 245(3):309-313 [J. Zijlstra/J. Zijlstra]
41100		S. Ladaveze and C. Muizon. 2010. Evidence of early evolution of Australidelphia (Metatheria, Mammalia) in South America: phylogenetic relationships of the metatherians from the Late Palaeocene of Itaboraí (Brazil) based on teeth and petrosal bones. Zoological Journal of the Linnean Society 159(3):746-784 [M. Uhen/M. Uhen]
58306		L. G. Marshall. 1981. Review of the Hathlyacyninae, an extinct subfamily of South American "dog-like" marsupials. Fieldiana Geology 7:1-120 [J. Zijlstra/J. Zijlstra]
57557		L. G. Marshall. 1982. Systematics of the extinct South American marsupial family Polydolopidae. Fieldiana Geology 12:1-109 [R. Butler/R. Butler]
57769		L. G. Marshall and C. de Muizon. 1984. Un nouveau Marsupial didelphidé (Itaboraidelphys camposi nov. gen., nov. sp.) du Paléocène moyen (Itaboraïen) de Sao José de Itaborai (Brésil) l. Comptes Rendus de l'Académie des Sciences de Paris, series II 299(18):1297-1300 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
38768		G. Mayr, H. Alvarenga, and J. Clarke. 2011. An Elaphrocnemus−like landbird and other avian remains from the late Paleocene of Brazil. Acta Palaeontologica Polonica 56(4):679-684 [P. Mannion/P. Mannion]
57759		E. V. Oliveira and L. P. Bergqvist. 1998. A new Paleocene armadillo (Mammalia, Dasypodoidea) from the Itaboraí Basin, Brazil. Asociación Paleontológica Argentina, Publicación Especial 5:35-40 [J. Zijlstra/J. Zijlstra]
81658		E. V. Oliveira, L. M. Carneiro, and F. J. Goin. 2021. A new derorhynchid (Mammalia, Metatheria) from the early Eocene Itaboraí fauna of Brazil with comments on its affinities. Anais da Academia Brasileira de Ciências 93(Suppl.2):e20201554 [P. Mannion/G. Varnham/G. Varnham]
57764		E. V. Oliveira and F. J. Goin. 2015. A new species of Gaylordia Paula Couto (Mammalia, Metatheria) from Itaboraí, Brazil. Revista Brasileira de Paleontologia 18(1):97-108 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
57603		E. V. Oliveira, N. Zimicz, and F. J. Goin. 2016. Taxonomy, affinities, and paleobiology of the tiny metatherian mammal Minusculodelphis, from the early Eocene of South America. The Science of Nature 103 [R. Butler/R. Butler]
43586		C. Paula Couto. 1962. Didelfidos fosiles del Paleoceno de Brasil. Revista del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" 8:135-166 [C. Jaramillo/G. Ballen]
43313		C. Paula Couto. 1963. Um Trigonostylopidae do Paleoceno do Brasil. Anais da Academia Brasileira de Ciencias 35(3):339-351 [C. Jaramillo/G. Ballen]
42935		C. d. Paula Couto. 1952. Fossil mammals from the beginning of the Cenozoic in Brazil. Marsupialia: Didelphidae. American Museum Novitates 1567:1-26 [P. Mannion/P. Mannion/P. Mannion]
42936		C. d. Paula Couto. 1952. Fossil mammals from the beginning of the Cenozoic in Brazil. Marsupialia: Polydolopidae and Borhyaenidae. American Museum Novitates 1559:1-27 [P. Mannion/P. Mannion/P. Mannion]
42938		C. d. Paula Couto. 1952. Fossil mammals from the beginning of the Cenozoic in Brazil. Notoungulata. American Museum Novitates 1568:1-16 [P. Mannion/P. Mannion/M. Uhen]
42939		C. d. Paula Couto. 1952. Fossil mammals from the beginning of the Cenozoic in Brazil. Condylarthra, Litopterna, Xenungulata, and Astrapotheria. Bulletin of the American Museum of Natural History 99(6):355-394 [P. Mannion/P. Mannion]
42934		C. d. Paula Couto. 1954. On a Notostylopid from the Paleocene of Itaborai, Brazil. American Museum Novitates 1693:1-5 [P. Mannion/P. Mannion/P. Mannion]
44088		A. E. P. Pinheiro, D. C. Fortier, D. Pol, D. A. Campos, and L. P. Bergqvist. 2013. A new Eocaiman (Alligatoridae, Crocodylia) from the Itaborai Basin, Paleogene of Rio de Janeiro, Brazil. Historical Biology 25(3):327-337 [P. Mannion/P. Mannion/P. Mannion]
85086		C. C. Rangel, L. M. Carneiro, M. F. Tejedor, L. P. Bergqvist, and E. V. Oliveira. 2023. A reassessment of Nemolestes (Mammalia, Metatheria): Systematics and evolutionary implications for Sparassodonta. Journal of Mammalian Evolution [J. Carrillo/K. Pino/K. Pino]