PBDB

Terril No 7, Avion: Westphalian C - Westphalian D, France
collected by Patrick Roques

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Insecta - Enigmapteridae

	Enigmaptera magnifica n. gen., n. sp. Nel et al. 2019	Nel et al. 2019	1 specimen
	Avion 35
Insecta - Meganisoptera - Meganeuridae

	Gallotupus oudardi n. gen., n. sp. Nel et al. 2008		1 specimen
	(1 measurement)
Insecta

	Jacquesoudardia magnifica n. gen., n. sp. Prokop et al. 2014	Prokop et al. 2014	1 specimen
	Avion no. 9 (1 measurement)
Insecta - Apatohymenoptera - Avioxyelidae

	Avioxyela gallica n. gen., n. sp. Nel et al. 2013	Nel et al. 2013	1 specimen
	Avion no. 3 (1 measurement)
Insecta - Westphalopsocidae

	Westphalopsocus pumilio n. gen., n. sp. Azar et al. 2013	Nel et al. 2013	1 specimen
	Avion no. 4 (1 measurement)
Insecta - Hemiptera - Aviorrhynchidae

	Aviorrhyncha magnifica n. gen., n. sp. Nel et al. 2013	Nel et al. 2013	1 specimen
	Avion no. 2 (1 measurement)
Insecta - Lophioneurida - Westphalothripidesidae

	Westphalothripides oudardi n. gen., n. sp. Nel et al. 2012	Nel et al. 2012	1 specimen
	MNHN-A31824 (1 measurement)
Insecta - Plecoptera - Gulouidae

	Gulou oudardi n. sp. Schubnel et al. 2019	Schubnel et al. 2019	1 specimen
	MNHN.F.A70112 (Avion 38)
Insecta - Plecoptera - Fatjanopteridae

	Avionptera communeaui n. gen., n. sp. Schubnel et al. 2019	Schubnel et al. 2019	1 specimen
	MNHN.F.A70113 (Avion 39)
Insecta - Cnemidolestodea - Tococladidae

	cf. Tococladus sp. Carpenter 1966	Coty et al. 2014	1 specimen
	Avion 00013
Insecta - Cnemidolestodea - Cnemidolestidae

	Piesbergopterum avionensis n. sp. Nel and Roques 2021	Nel and Roques 2021	1 specimen
	MNHN.F.A71354 (1 measurement)
Insecta - Cnemidolestodea

	Aviocladus pectinatus n. gen., n. sp. Prokop et al. 2014	Prokop et al. 2014	1 specimen
	Avion no. 7 (1 measurement)

	Aviohapaloptera bethouxi n. gen., n. sp. Prokop et al. 2014	Prokop et al. 2014	1 specimen
	Avion no. 8 (1 measurement)
Insecta - Cacurgidae

	Cacurgus avionensis n. sp. Schubnel et al. 2019	Schubnel et al. 2019	1 specimen
	MNHN.F.A70497 (1 measurement)
Insecta - Caloneurodea - Permobiellidae

	Aviobiella garroustei n. gen., n. sp. Nel and Roques 2021	Nel and Roques 2021	1 specimen
	MNHN.F.A71352
Insecta - Caloneurodea

	Aviogramma gracilis n. gen., n. sp. Prokop et al. 2014	Prokop et al. 2014	1 specimen
	Avion no. 10 (1 measurement)
Insecta - Eoblattidae

	Beloatta duquesnei n. gen., n. sp. Nel et al. 2020	Nel et al. 2020	1 specimen
	MNHN.F.A71343 (1 measurement)
Insecta

	Aviologus duquesnei n. gen., n. sp. Coty et al. 2014	Coty et al. 2014	1 specimen
	Avion 00034 (1 measurement)
Insecta - Nugonioneuridae

	Avionugonioneura jouaulti n. gen., n. sp. Nel and Roques 2021	Nel and Roques 2021	1 specimen
	MNHN.F.A71353 (1 measurement)
Insecta - Orthoptera - Theiatitanidae

	Theiatitan azari n. gen., n. sp. Schubnel et al. 2021	Schubnel et al. 2021	1 specimen
	MNHN.F.A70111 (Avion 37)
Insecta - Paoliida - Paoliidae

	Paolia sp. Smith 1871	Prokop et al. 2014	1 specimen
	Avion no. 6 (1 measurement)
Insecta - Paoliida - Blattinopsidae

	Avionblattinopsis oudardi n. gen., n. sp. Quispe et al. 2021	Quispe et al. 2021	1 specimen
	MNHN.F.A71321 (1 measurement)
Insecta - Palaeodictyoptera - Breyeriidae

	Breyeriidae indet. Handlirsch 1906	Prokop et al. 2013	3 specimens
	Avion no. 2, 3, 4

	Aviobreyeria gracilis n. gen., n. sp. Prokop et al. 2013	Prokop et al. 2013	1 specimen
	Avion no. 1 (1 measurement)
Insecta - Palaeodictyoptera - Homoiopteridae

	Homoiopteridae indet. Handlirsch 1906	Prokop et al. 2014	1 specimen
	Avion no. 12 (1 measurement)
Insecta - Palaeodictyoptera - Spilapteridae

	Spilaptera splendens n. sp. Prokop et al. 2014	Prokop et al. 2014	1 specimen
	Avion no. 11 (1 measurement)
Insecta - Diaphanopterodea

	Avionbrodia incompleta n. gen., n. sp. Schubnel et al. 2020	Schubnel et al. 2020	1 specimen
	MNHN.F.A71314 (Jacques Oudard P 1568-76) (1 measurement)
see common names

Geography

Country:	France
Coordinates:	50.4° North, 2.8° East (view map)
Paleocoordinates:	0.9° South, 12.4° East
Basis of coordinate:	estimated from map
Geographic resolution:	outcrop

Time

Period:	Carboniferous	Epoch:	Middle Pennsylvanian
Stage:	Moscovian	10 m.y. bin:	Carboniferous 4
Key time interval:	Westphalian C - Westphalian D
Age range of interval:	314.60000 - 306.95000 m.y. ago

Stratigraphy

Stratigraphic resolution:	group of beds
Stratigraphy comments: Bolsovian (Westphalian C, 308-311 Mya, “faisceaux de Ernestine”) / Asturian (Westphalian D, 306-308 Mya, ”veines Arago, Dusouich, Marthe”)

Lithology and environment

Primary lithology:	lithified "shale"
Environment:	terrestrial indet.	Tectonic setting:	foreland basin
Geology comments: Variscan foreland basin

Taphonomy

Modes of preservation:	mold/impression
Size of fossils:	macrofossils

Collection methods and comments

Reason for describing collection:	taxonomic analysis
Museum repositories:	MNHN
Collectors:	Patrick Roques
Collection method comments: Specimen in Patrick Roques private collection, cast in Laboratoire de Paleontologie, MNHN, Paris. Material ultimately to be deposited in the Musée Géologique Pierre Vetter, Decazeville, France.

Metadata

Database number:	122504
Authorizer:	M. Clapham	Enterer:	M. Clapham
Modifier:	M. Clapham	Research group:	paleoentomology
Created:	2011-12-31 12:24:00	Last modified:	2013-11-25 13:02:06
Access level:	the public	Released:	2011-12-31 12:24:00
Creative Commons license:	CC BY

Reference information

Primary reference:

39050.

A. Nel, R. Garrouste, and P. Roques. 2008. A new griffenfly genus from the Late Carboniferous of northern France (Odonatoptera: Meganeuridae). Insect Systematics & Evolution 39:231-239 [M. Clapham/M. Clapham/M. Clapham]

Secondary references:

54489		D. Coty, J. Háva, J. Prokop, P. Roques, and A. Nel. 2014. New archaeorthopteran insects from the Late Carboniferous of the Nord and Pas-de-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera). Zootaxa 3878:462-470 [M. Clapham/M. Clapham]
83678		A. Nel, R. Garrouste, and P. Roques. 2020. The first representative of the archaeorthopteran family Eoblattidae in the Konservat Lagerstätte of Avion, France (Insecta: Polyneoptera). Palaeoentomology 3:552-555 [M. Clapham/M. Clapham]
48955		A. Nel, P. Roques, P. Nel, A. A. Prokin, T. Bourgoin, J. Prokop, J. Szwedo, D. Azar, L. Desutter-Grandcolas, T. Wappler, R. Garrouste, D. Coty, D. Y. Huang, M. S. Engel, and A. G. Kirejtshuk. 2013. The earliest known holometabolous insects. Nature 503:257-261 [M. Clapham/M. Clapham]
69178		A. Nel, P. Roques, J. Prokop and R. Garrouste. 2019. A new, extraordinary 'damselfly-like' Odonatoptera from the Pennsylvanian of the Avion locality in Pas-de-Calais, France (Insecta: 'Exopterygota'). Alcheringa 43:241-245 [M. Clapham/M. Clapham/M. Clapham]
84004		A. Nel and P. Roques. 2021. The second Caloneurodea from the Moscovian of Avion, France (Insecta, Archaeorthoptera). Palaeoentomology 4:320-322 [M. Clapham/M. Clapham]
84010		A. Nel and P. Roques. 2021. The second species of the cnemidolestid genus Piesbergopterum from the Upper Carboniferous of Avion, Northern France (Archaeorthoptera: Cnemidolestidae). Palaeoentomology 4:323-325 [M. Clapham/M. Clapham]
82365		A. Nel and P. Roques. 2021. A new strange Archaeorthoptera from the Moscovian of Avion (France) (Insecta, Polyneoptera). Historical Biology 34:1769-1771 [M. Clapham/M. Clapham]
41609		P. Nel, D. Azar, J. Prokop, P. Roques, G. Hodebert and A. Nel. 2012. From Carboniferous to Recent: wing venation enlightens evolution of thysanopteran lineage. Journal of Systematic Palaeontology 10:385-399 [M. Clapham/M. Clapham]
50568		J. Prokop, P. Roques, and A. Nel. 2014. New non-holometabolous insects from the Pennsylvanian of the Avion locality in Pas-de-Calais, France (Insecta: 'Exopterygota'). Alcheringa 38:155-169 [M. Clapham/M. Clapham]
51486		J. Prokop, Z. Tippeltová, P. Roques and A. Nel. 2013. A new genus and species of Breyeriidae and wings of immature stages from the Upper Carboniferous, Nord-Pas-de-Calais, France (Insecta: Palaeodictyoptera). Insect Systematics & Evolution 44:117-128 [M. Clapham/M. Clapham]
80082		L. Quispe, P. Roques, R. Garrouste and A. Nel. 2021. Carboniferous Blattinopsidae: revision of Klebsiella and new genus and species from Avion (Insecta, Paoliida). Historical Biology 34:383-389 [M. Clapham/M. Clapham]
77349		T. Schubnel, F. Legendre, P. Roques, R. Garrouste, R. Cornette, M. Perreau, N. Perreau, L. Desutter-Grandcolas, and A. Nel. 2021. Sound vs. light: wing-based communication in Carboniferous insects. Communications Biology 4(794) [M. Clapham/M. Clapham]
69421		T. Schubnel, L. Perdu, P. Roques, R. Garrouste, and A. Nel. 2019. Two new stem-stoneflies discovered in the Pennsylvanian Avion locality, Pas-de-Calais, France (Insecta: 'Exopterygota'). Alcheringa 43:430-435 [M. Clapham/M. Clapham/M. Clapham]
71711		T. Schubnel, D. Roberts, P. Roques, R. Garrouste, L. Desutter-Grandcolas and A. Nel. 2019. Moscovian fossils shed light on the enigmatic polyneopteran families Cacurgidae and Eoblattidae (Insecta: 'Eoblattida', Archaeorthoptera). Journal of Systematic Palaeontology 18:499-511 [M. Clapham/M. Clapham/M. Clapham]
83522		T. Schubnel, P. Roques, J. Oudard, R. Garrouste, and A. Nel. 2020. A new paleopteran genus and species from the late Carboniferous of Avion France, (Insecta: Palaeodictyopterida: ?Diaphanopterodea). Palaeoentomology 3:32-35 [M. Clapham/M. Clapham]