Quarry L-41 (Jebel Qatrani Formation): Rupelian, Egypt
collected 1983, 1987, 1988

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Polyprotodonta
Polyprotodonta informal gen. nov. & sp. nov.
spelled with current rank as Polyprotodontia
Mammalia - Embrithopoda - Arsinoitheriidae
Arsinoitherium sp. Beadnell 1902
Mammalia - Hyracoidea - Pliohyracidae
Megalohyrax aff. eocaenus Andrews 1903
Geniohyus sp. n. sp. Andrews 1904
Thyrohyrax litholagos
    = Thyrohyrax litholagus Rasmussen and Simons 1991
Rasmussen and Simons 1991
Thyrohyrax meyeri Rasmussen and Simons 1991
Saghatherium bowni Rasmussen and Simons 1991
Bunohyrax sp. n. sp. Schlosser 1910
Mammalia - Hyracoidea - Titanohyracidae
Titanohyrax informal sp. nov. A Matsumoto 1922
    = Antilohyrax pectidens n. gen., n. sp. Rasmussen and Simons 2000
Rasmussen and Simons 2000
Titanohyrax informal sp. nov. B Matsumoto 1922
Mammalia
Widanelfarasia rasmusseni n. sp. Seiffert and Simons 2000
Seiffert and Simons 2000
Widanelfarasia bowni n. gen., n. sp. Seiffert and Simons 2000
Seiffert and Simons 2000
Mammalia - Macroscelidea - Macroscelididae
Herodotius pattersoni Simons et al. 1991
Mammalia - Chiroptera - Emballonuridae
Dhofarella sigei n. sp. Gunnell et al. 2008
Gunnell et al. 2008
Mammalia - Chiroptera - Vespertilionidae
Khonsunycteris aegypticus n. gen., n. sp. Gunnell et al. 2008
Gunnell et al. 2008
Mammalia - Chiroptera - Megadermatidae
Saharaderma pseudovampyrus n. gen., n. sp. Gunnell et al. 2008
Gunnell et al. 2008
Mammalia - Insectivora
Insectivora informal indet. Bowdich 1821
Mammalia - Artiodactyla
Artiodactyla informal gen. nov. & sp. nov. A
Artiodactyla informal gen. nov. & sp. nov. B
Mammalia - Creodonta
Creodonta indet. (Cope 1875)
Masrasector sp. n. sp. Simons and Gingerich 1974
    = Masrasector nananubis n. sp. Borths and Seiffert 2017
Borths and Seiffert 2017
CGM 83736 (formerly DPC 11383) - type
Brychotherium ephalmos n. gen., n. sp. Holroyd 1994
Borths et al. 2016
CGM 83750 - holotype; referred material: DPC 11569, 11990 and 17627
Pterodontinae informal new genus and species D
Holroyd 1999
synonym of Hyainailourinae
DPC 6545
Akhnatenavus aff. leptognathus (Osborn 1909)
Holroyd 1999
    = Akhnatenavus nefertiticyon n. sp. Borths et al. 2016
Borths et al. 2016
CGM 83735 - type; referred material: DPC 7765, 13518 and 18242
Apterodon sp. n. sp. Fischer 1880
"Metasinopa" sp. n. sp. Osborn 1909
Mammalia - Rodentia
Rodentia informal gen. nov. & sp. nov. A
Mammalia - Rodentia - Myophiomyidae
Phiomys lavocati Wood 1968
recombined as Talahphiomys lavocati
Mammalia - Rodentia
Mubhammys vadumensis n. gen., n. sp. Sallam and Seiffert 2016
Sallam and Seiffert 2016
CGM 66001 - holotype; referred material: DPC 14324, 13220 and 14141
Acritophiomys bowni n. gen., n. sp. Sallam et al. 2012
Sallam et al. 2012
recombined as Phiomys bowni
CGM 83705 - holotype; referred material: DPC specimens
cf. Birkamys sp. Sallam and Seiffert 2016
Sallam and Seiffert 2016
DPC 17995
Birkamys korai n. gen., n. sp. Sallam and Seiffert 2016
Sallam and Seiffert 2016
CGM 66000 - holotype; referred material: DPC 9276, 15625, 17457 and 22737
Mammalia - Rodentia - Gaudeamuridae
Gaudeamus hylaeus n. sp. Sallam et al. 2011
Sallam et al. 2011
CGM 66007 - holotype; referred material: DPC specimens
Gaudeamus aff. hylaeus Sallam et al. 2011
Sallam et al. 2011
DPC 17624 and 14487
Gaudeamus sp. n. sp. Wood 1968
    = Gaudeamus aslius n. sp. Sallam et al. 2011
Sallam et al. 2011
CGM 66006 - holotype; referred material: DPC specimens
Gaudeamus aff. aslius Sallam et al. 2011
Sallam et al. 2011
DPC 12990
Mammalia - Primates - Oligopithecidae
Catopithecus browni n. gen., n. sp. Simons 1989
Simons 1989
CGM 41885 - holotype; DPC 8701 - referred skull (1 measurement)
Mammalia - Primates - Proteopithecidae
Serapia eocaena n. gen., n. sp. Simons 1992
Simons 1992
Proteopithecus sylviae n. gen., n. sp. Simons 1989
Simons 1989
CGM 42209 and 42217, and DPC 16873 and17031 (2 measurements)
Mammalia - Primates
Arsinoea kallimos n. gen., n. sp. Simons 1992
Simons 1992
Mammalia - Primates - Parapithecidae
Abuqatrania basiodontos n. gen., n. sp. Simons et al. 2001
Simons et al. 2001
recombined as Qatrania basiodontos
Mammalia - Primates - Notharctidae
Anchomomys milleri n. sp. Simons 1997
Simons 1997
Mammalia - Primates
Aframonius diedies n. gen., n. sp. Simons et al. 1995
Simons et al. 1995
Plesiopithecus teras n. gen., n. sp. Simons 1992
Simons 1992
Mammalia - Primates - Galagidae
Wadilemur elegans n. gen., n. sp. Simons 1997
Simons 1997
Mammalia - Ptolemaiida - Ptolemaiidae
Ptolemaiidae indet. Osborn 1908
said to be a new genus and species
Reptilia - Ciconiiformes - Ciconiidae
Ciconiidae indet. Sundevall 1836
Miller et al. 1997 1 specimen
Reptilia - Eremopezidae
Eremopezus eocaenus Andrews 1904
Rasmussen et al. 2001
see common names

Geography
Country:Egypt State/province:Fayum
Coordinates: 29.5° North, 30.5° East (view map)
Paleocoordinates:26.1° North, 27.2° East
Basis of coordinate:stated in text
Time
Period:Paleogene Epoch:Oligocene
Stage:Rupelian 10 m.y. bin:Cenozoic 4
Key time interval:Rupelian
Age range of interval:33.90000 - 27.82000 m.y. ago
Stratigraphy
Formation:Jebel Qatrani
Stratigraphy comments: "more likely" represents Chron 13r and therefore is "very latest Eocene in age" (Gunnell et al. 2008); Rupelian age after Coster et al. (2012), Chron 13R is Oligocene not Eocene
Lithology and environment
Primary lithology:very fine,green unlithified silty mudstone
Includes fossils?Y
Lithology description: Lithologic description based on direct observation by data enterer. Said to be a "green shale" or "green clay" by Simons (1989)
Environment:lacustrine indet.
Geology comments: "a lacustrine deposit; vertebrates may have been flushed into the area during large storms" (Gunnell et al. 2008)
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Associated major elements:many
Disassociated major elements:many
Disassociated minor elements:some
Spatial resolution:allochthonous
Collection methods and comments
Collection excludes:some genera
Collection methods:selective quarrying,surface (in situ),field collection,survey of museum collection
Reason for describing collection:paleoecologic analysis
Collection dates:1983, 1987, 1988
Metadata
Also known as:lower sequence; Fayum Faunal Zone 1
Database number:60127
Authorizer:P. Holroyd, P. Mannion, K. Beard, J. Alroy, M. Uhen, P. Coster Enterer:P. Holroyd, K. Beard, P. Mannion, J. Alroy, M. Uhen, P. Coster
Modifier:M. Uhen Research group:vertebrate
Created:2006-05-02 21:28:24 Last modified:2022-11-17 14:55:36
Access level:the public Released:2006-05-02 21:28:24
Creative Commons license:CC BY
Reference information

Primary reference:
17279. M. Gagnon. 1997. Ecological diversity and community ecology in the Fayum sequence (Egypt). Journal of Human Evolution 32:133-160 [P. Holroyd/P. Holroyd/P. Coster]

Secondary references:

60781 M. R. Borths, P. A. Holroyd, and E. R. Seiffert. 2016. Hyainailourine and teratodontine cranial material from the late Eocene of Egypt and the application of parsimony and Bayesian methods to the phylogeny and biogeography of Hyaenodonta (Placentalia, Mammalia). PeerJ 4:e2639:1-87 [P. Mannion/P. Mannion]
62204 M. R. Borths and E. R. Seiffert. 2017. Craniodental and humeral morphology of a new species of Masrasector (Teratodontinae, Hyaenodonta, Placentalia) from the late Eocene of Egypt and locomotor diversity in hyaenodonts. PLoS ONE 12(4):e0173527:1-60 [P. Mannion/P. Mannion]
28499 G. F. Gunnell, E. L. Simons, and E. R. Seiffert. 2008. New bats (Mammalia: Chiroptera) from the late Eocene and early Oligocene, Fayum Depression, Egypt. Journal of Vertebrate Paleontology 28(1):1-11 [J. Alroy/J. Alroy]
17283 P. A. Holroyd. 1999. New Pterodontinae (Creodonta: Hyaenodontidae) from the late Eocene-early Oligocene Jebel Qatrani Formation, Fayum province, Egypt. PaleoBios 19(2):1-18 [P. Holroyd/P. Holroyd/P. Mannion]
43809 E. R. Miller, D. T. Rasmussen, and E. L. Simons. 1997. Fossil storks (Ciconiidae) from the Late Eocene and Early Miocene of Egypt. The Ostrich 68(1):23-26 [M. Uhen/M. Uhen]
57461 D. Rasmussen and E. Simons. 1991. The oldest Egyptian Hyracoids (Mammalia: Pliohyracidae): New species of Saghatherium and Thyrohyrax from the Fayum. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 182:187-209 [P. Coster/P. Coster]
29520 D. T. Rasmussen and E. L. Simons. 2000. Ecomorphological diversity among Paleogene hyracoids (Mammalia): a new cursorial browser from the Fayum, Egypt. Journal of Vertebrate Paleontology 20(1):167-176 [J. Alroy/J. Alroy]
43795 D. T. Rasmussen, E. L. Simons, F. Hertel and A. Judd. 2001. Hindlimb of a giant terrestrial bird form the upper Eocene, Fayum, Egypt. Palaeontology 44(2):325-337 [M. Uhen/M. Uhen]
59581 H. M. Sallam, E. R. Seiffert, and E. L. Simons. 2011. Craniodental Morphology and Systematics of a New Family of Hystricognathous Rodents (Gaudeamuridae) from the Late Eocene and Early Oligocene of Egypt. PLoS ONE 6(2):e16525:1-29 [P. Mannion/P. Mannion]
59514 H. M. Sallam, E. R. Seiffert, and E. L. Simons. 2012. A basal phiomorph (Rodentia, Hystricognathi) from the late Eocene of the Fayum Depression, Egypt. Swiss Journal of Palaeontology 131(2):283-301 [P. Mannion/P. Mannion]
59515 H. M. Sallam and E. R. Seiffert. 2016. New phiomorph rodents from the latest Eocene of Egypt, and the impact of Bayesian "clock"-based phylogenetic methods on estimates of basal hystricognath relationships and biochronology. PeerJ 4:e1717:1-53 [P. Mannion/P. Mannion]
23722 E. R. Seiffert and E. L. Simons. 2000. Widanelfarasia, a diminutive placental from the late Eocene of Egypt. Proceedings of the National Academy of Sciences 97(6):2646-2651 [J. Alroy/J. Alroy]
27009 E. R. Seiffert, E. L. Simons, T. M. Ryan and Y. Attia. 2005. Additional remains of Wadilemur elegans, a primitive stem galagid from the Late Eocene of Egypt. Proceedings of the National Academy of Sciences 102:11396-11401 [J. Alroy/J. Alroy]
29400 E. L. Simons. 1989. Description of two genera and species of Late Eocene Anthropoidea from Egypt. Proceedings of the National Academy of Sciences 86:9956-9960 [J. Alroy/J. Alroy]
59535 E. L. Simons. 1990. Discovery of the Oldest Known Anthropoidean Skull from the Paleogene of Egypt. Science 247(4950):1567-1569 [P. Mannion/P. Mannion]
28178 E. L. Simons. 1992. Diversity in the early Tertiary anthropoidean radiation in Africa. Proceedings of the National Academy of Sciences 89:10743-10747 [J. Alroy/J. Alroy]
29548 E. L. Simons. 1997. Discovery of the Smallest Fayum Egyptian Primates (Anchomomyini, Adapidae). Proceedings of the National Academy of Sciences 94(1):180-184 [J. Alroy/J. Alroy]
29402 E. L. Simons and E. R. Miller. 1997. An upper dentition of Aframonius dieides (Primates) from the Fayum, Egyptian Eocene. Proceedings of the National Academy of Sciences 94:7993-7996 [J. Alroy/J. Alroy]
29560 E. L. Simons, D. T. Rasmussen, and P. D. Gingerich. 1995. New cercamoniine adapid from Fayum, Egypt. Journal of Human Evolution 29(6):577-589 [J. Alroy/J. Alroy]
58488 E. L. Simons and E. R. Seiffert. 1999. A partial skeleton of Proteopithecus sylviae (Primates, Anthropoidea): first associated dental and postcranial remains of an Eocene anthropoidean. Comptes Rendus de l'Académie des Sciences, Paris, Sciences de la terre et des planètes 329:921-927 [P. Mannion/P. Mannion]
32563 E. L. Simons, E. R. Seiffert, P. S. Chatrath and Y. Attia. 2001. Earliest record of a parapithecid anthropoid from the Jebel Qatrani Formation, northern Egypt. Folia Primatologica 72:316-331 [K. Beard/K. Beard]