Csarnota locality 2: Ruscinian - Villanyian, Hungary

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Megalobatrachidae indet.
2 specimens
Procamptoceros cf. brivatense
corrected as Procamptoceras
Myriapoda
Diplopoda indet. Gervais 1844
Gastropoda
Gastropoda indet. Cuvier 1797
Pisces
Pisces indet. Linnaeus 1758
synonym of Osteichthyes
Albanerpetontidae
Albanerpeton sp. Estes and Hoffstetter 1976
Venczel and Gardner 2003
"new species"
    = Albanerpeton pannonicus n. sp. Venczel and Gardner 2005
Venczel and Gardner 2005
MAFI V.22000 - holotype (fused pair of nearly complete premaxillae, articulated with anterior end of right maxilla); referred material: fragmentary skulls (n = 24), each consisting of either a maxillary arcade comprising a maxilla articulated with some combination of a jugal, lacrimal, prefrontal and palatal bones or an incomplete skull roof comprising fused frontals articulated with some combination of a prefrontal, lacrimal and nasal: MAFI V.22003 ⁄ 1, V.22011, V.22012 ⁄ 1–3, V.22015, V.22021 ⁄ 1–2, V.22033 ⁄ 2, V.22034, V.22039, V.22053, V.22055, V.22075, V.22076 ⁄ 1–2, V.22077, V.22093 ⁄ 1–3, V.22094 ⁄ 1, V.22100 ⁄ 1–2, V.22111; fused premaxillae (n = 6): MAFI V.22017 ⁄ 1–2, V.22023, V.22026 ⁄ 1, V.22041 ⁄ 1, V.22072 ⁄ 1; isolated premaxillae (n = 38): MAFI V.22006 ⁄ 1–5, V.22007, V.22018 ⁄ 1–3, V.22026 ⁄ 2, V.22030 ⁄ 1–2, V.22036 ⁄ 1–4, V.22041 ⁄ 2, V.22050, V.22057, V.22067, V.22072 ⁄ 2–3, V.22078 ⁄ 1–2, V.22083, V.22089 ⁄ 1–3, V.22095, V.22101 ⁄ 1–2, V.22105, V.22106 ⁄ 1–3, V.22113 ⁄ 1–3; isolated maxillae (n = 50): MAFI V.22002 ⁄ 1–6, V.22013 ⁄ 1–10, V.22014 ⁄ 1–2, V.22028 ⁄ 1, V.22038 ⁄ 1–3, V.22043 ⁄ 1–2, V.22048, V.22056 ⁄ 5, V.22070 ⁄ 1–4, V.22087 ⁄ 1–2, V.22092 ⁄ 1–4, V.22099 ⁄ 1–4, V.22110 ⁄ 1–10; fused frontals, not articulated with other bones (n = 39): MAFI V.22003 ⁄ 2–3; V.22004 ⁄ 1–3, V.22005 ⁄ 1–4, V.22016 ⁄ 1–7, V.22022, V.22025, V.22029, V.22035, V.22040, V.22045, V.22060, V.22063, V.22066 ⁄ 1–2, V.22071, V.22081, V.22082 ⁄ 1–2, V.22088, V.22094 ⁄ 2, V.22104 ⁄ 1–5, V.22112 ⁄ 1–2; mandibles (n = 20): MAFI V.22009, V.22010 ⁄ 1–4, V.22020 ⁄ 1–4, V.22032 ⁄ 1–2, V.22047, V.22052, V.22059, V.22062, V.22065, V.22080, V.22097, V.22108 ⁄ 1–2; dentaries (n = 302): MAFI V.22001 ⁄ 1– 18, V.22008 ⁄ 1–38, V.22019 ⁄ 1–37, V.22024 ⁄ 1–2, V.22027 ⁄ 1–8, V.22031 ⁄ 1–10, V.22037 ⁄ 1–24, V.22042 ⁄ 1–3, V.22044 ⁄ 1–6, V.22046 ⁄ 1–10, V.22049, V.22051 ⁄ 1–4, V.22056 ⁄ 1–4, V.22058 ⁄ 1– 3, V.22061 ⁄ 1–10, V.22064 ⁄ 1–3, V.22068 ⁄ 1–12, V.22073 ⁄ 1–4, V.22079, V.22084, V.22085 ⁄ 1–11, V.22090 ⁄ 1–17, V.22096 ⁄ 1–23, V.22102 ⁄ 1–11, V.22107 ⁄ 1–25, V.22114 ⁄ 1–16; presacral vertebrae (n = 10): MAFI V.22028 ⁄ 2, V.22054, V.22074 ⁄ 1, V.22086 ⁄ 1–2, V.22098 ⁄ 1, V.22103 ⁄ 1, V.22109 ⁄ 1–2, V.22114 ⁄ 17; humeri (n = 16): MAFI V.22033 ⁄ 1, V.22069, V.22074 ⁄ 2, V.22086 ⁄ 3, V.22091 ⁄ 1–5, V.22098 ⁄ 2–4, V.22103 ⁄ 2–3, V.22109 ⁄ 3–4.
Salientia
Anura indet. (Fischer von Waldheim 1813)
(minly Bufo (listed seperately) c. 20,000 specimens in total)
Salientia - Bufonidae
Bufo sp. Laurenti 1768
Urodela
Triturus sp. Rafinesque 1815
sporadic
Salamandra sp. Laurenti 1768
rare
Reptilia - Testudines - Testudinidae
Testudo cf. lambrechti
rare
Reptilia - Avetheropoda
Aves indet. Linnaeus 1758
Reptilia - Avetheropoda - Tetraonidae
Tetrao macropus
few
Reptilia - Avetheropoda - Phasianidae
Francolinus capeki Lambrecht 1933
sporadic. Francolinus capeki wezensis
Reptilia - Squamata - Lacertidae
Lacerta ruscinensis
several thousand
Lacerta aff. agilis Linneaus 1758
few
Reptilia - Squamata - Varanidae
Varanus deserticolus
few
Reptilia - Squamata - Anguidae
Anguis sp. Linnaeus 1758
several thousand
Ophisaurus intermedius
common
Reptilia - Squamata
Ophidia indet. Latreille 1804
abundant
Reptilia - Squamata - Colubridae
Elaphe quatuorlineata (Lacépède 1789)
Venczel and Vardai 2000 1 element
a single vertebra (GMH uncatalogued)
Mammalia - Primates - Cercopithecidae
cf. Dolichopithecus sp. Depéret 1889
cf. Dolichopithecus sp. indet.
Macaca sp. Lacépède 1799
Macaca sp. indet. sporadic
Mammalia - Rodentia - Muridae
Apodemus dominans
8000 specimens
c. 8000
Micromys praeminutus
few
Rhagapodemus frequens
common
Mammalia - Rodentia - Cricetidae
Propliomys hungaricus
abundant
Baranomys loczyi
few
Cseria gracilis
synonym of Mimomys
common
Prospalax priscus
600 specimens
c. 600
Promimomys cor
1 specimen
Dolomys nehringi
common
Cricetinus europaeus
few
Mammalia - Rodentia - Sciuridae
Sciurus sp. Linnaeus 1758
1 specimen
Sciurus sp. indet.
Pliopetes hungaricus
few
? Eutamias sp. Trouessart 1880
Eutamias sp. indet. few
Pliopetaurista pliocaenica
2 specimens
Mammalia - Rodentia - Gliridae
Glirurus gemmula
1 specimen
Dryomimus eliomyoides Kretzoi 1959
1000 specimens
c. 1000
Muscardinus sp. Kaup 1829
few
Glis minor Kowalski 1963
few
Mammalia - Lagomorpha - Leporidae
Hypolagus cf. beremendensis
few.
Mammalia - Lagomorpha - Ochotonidae
Ochotonoides csarnotanus
1-2 specimens
Mammalia - Lipotyphla - Erinaceidae
Hemiechinus cf. samsonowiczi
few
Mammalia - Lipotyphla - Talpidae
Desmana sp. Guldenstaedt 1777
Desmana sp. indet. I-II, rare
Talpa csarnotana
100 specimens
c. 100
Talpa cf. fossilis Petenyi 1864
rare
Mammalia - Soricidae
Sorex sp. Linnaeus 1758
Sorex s. 1. sp.
Petenyia hungarica Kormos 1934
100 specimens
c. 100
Episoriculus gibberodon Petenyi 1864
100 specimens
c. 100
Blarinoides mariae Sulimski 1959
few
Petenyiella gracilis
Beremendia fissidens (Petényi 1864)
400 specimens
c. 400
Mammalia - Chiroptera
Chiroptera indet. Blumenbach 1779
common
Mammalia - Carnivora - Ursidae
Ursus sp. Linnaeus 1758
few. Ursus s. 1. sp. indet.
Mammalia - Carnivora - Mustelidae
Baranogale beremendensis
few.
Mustela aff. palerminea
few.
Xenictis pilgrimi
few.
Mammalia - Carnivora - Canidae
Canidae indet. Fischer 1817
few. Canidae I-III indet.
Mammalia - Carnivora - Felidae
? Lynx sp. Kerr 1792
Lynx (?) sp. indet.
Felis sp. Linnaeus 1758
1 specimen
Felis s. 1. indet. (size of catus)
Mammalia - Ungulata - Bovidae
Hemitragus cf. bonali
Gazellospira cf. torticornis
Mammalia - Ungulata - Cervidae
"Alces" sp. Gray 1821
Alces sp. indet.
Cervus (? Rusa) sp. (Smith 1827)
synonym of Cervus
Cervus (?Rusa) sp. indet.
? Cervus pyrenaicus (Depéret 1890)
Small Cervidae (Cervus pyrenaicus Deperet?)
Megaceros cf. dupuisi
synonym of Megaloceros
(unlikely according to Kretzoi)
see common names

Geography
Country:Hungary State/province:Barany
Coordinates: 45.9° North, 18.2° East (view map)
Paleocoordinates:45.8° North, 18.1° East
Basis of coordinate:based on nearby landmark
Time
Period:Neogene Epoch:Pliocene
10 m.y. bin:Cenozoic 6
Key time interval:Ruscinian - Villanyian
Age range of interval:5.33300 - 2.58800 m.y. ago
Stratigraphy
Stratigraphy comments: Csarnotian stage, Mid. Pliocene, MN15 (according to Fejfar et al., 1997). Janossy puts it slightly later (in the Villanyian). Villafranchian.
Lithology and environment
Lithology description: 'red clay.'
Environment:karst indet.
Taphonomy
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:bulk,field collection
Reason for describing collection:biostratigraphic analysis
Collection method comments: This is a reconstructed faunal list, including several collections excavated from 1910 onwards. The original site lists (1910-1930s) do not give exact locality info., but the later excavations uncovered 25 strata in a trench 1m x 1.5m and 3m deep.
Taxonomic list comments:'Two Pliogallus species described by Galliard have been deleted from the list.'
Metadata
Also known as:Csarnóta 2
Database number:35354
Authorizer:A. Turner, J. Mueller, P. Mannion, M. Carrano Enterer:H. O'Regan, T. Liebrecht, P. Mannion, M. Carrano
Modifier:H. O'Regan Research group:PACED
Created:2003-12-05 04:26:08 Last modified:2003-12-05 06:26:08
Access level:the public Released:2003-12-05 04:26:08
Creative Commons license:CC BY
Reference information

Primary reference:

9534.PACED D. Janossy. 1986. Pleistocene vertebrate faunas of Hungary. Developments in Palaeontology and Stratigraphy, 8. Elsevier, Amsterdam 1-208 [A. Turner/H. O'Regan/H. O'Regan]

Secondary references:

23463ETE M. Venczel and J. Gardner. 2003. The geologically youngest albanerpetontid amphibian. Journal of Vertebrate Paleontology 23(3, suppl.):106A [M. Carrano/M. Carrano]
44192 M. Venczel and J. D. Gardner. 2005. The geologically youngest albanerpetontid amphibian, from the Lower Pliocene of Hungary. Palaeontology 48(6):1273-1300 [P. Mannion/P. Mannion]
54746 M. Venczel and G. Vardai. 2000. The Genus Elaphe in the Carpathian Basin: Fossil Record. Nymphaea – Folia Naturae Bihariae 28:65-82 [J. Mueller/T. Liebrecht]