Andrès (MNHN-UL): Late/Upper Oxfordian - Late/Upper Kimmeridgian, Portugal
collected by J. Amorim 1988, 2005

List of taxa
Where & when
Taphonomy & methods
Metadata & references
Taxonomic list
Diplodocoidea indet. Upchurch 1995
Malafaia et al. 2006
    = Diplodocidae indet.
Malafaia et al. 2010
Bivalvia indet. Linnaeus 1758
Malafaia et al. 2006
Gastropoda indet. Cuvier 1797
Malafaia et al. 2006
Actinopteri - Lepisosteiformes
Lepidotes sp. (Agassiz 1843)
Malafaia et al. 2006 1 specimen
nearly complete individual
Actinopteri - Semionotiformes
Semionotiformes indet. Arambourg and Bertin 1958
Malafaia et al. 2006
Camarasaurus sp. Cope 1877
Malafaia et al. 2006
    = Neosauropoda indet. Bonaparte 1986
Malafaia et al. 2010
Titanosauriformes indet. Salgado et al. 1997
Malafaia et al. 2006
Reptilia - Avetheropoda - Allosauridae
Allosaurus fragilis Marsh 1877
2 individuals
    = Allosaurus europaeus Mateus et al. 2006
Mateus et al. 2006
Reptilia - Avetheropoda - Dromaeosauridae
Dromaeosauridae indet. Colbert and Russell 1969
Malafaia et al. 2006
Ornithopoda indet. Marsh 1881
Malafaia et al. 2006
Reptilia - Dryosauridae
Dryosauridae indet. Stefano 1903
Malafaia et al. 2006
    = aff. Dryosaurus sp. Marsh 1894
Malafaia et al. 2010
Reptilia - Camptosauridae
Camptosauridae indet. Marsh 1885
Malafaia et al. 2006
Reptilia - Rhamphorhynchidae
Pterosauria indet. Kaup 1834
Malafaia et al. 2006
    = cf. Rhamphorhynchus sp. Meyer 1847
Malafaia et al. 2010
Reptilia - Goniopholididae
Goniopholis sp. Owen 1842
Malafaia et al. 2006
Theriosuchus sp. Owen 1878
Malafaia et al. 2006
Lepidosauria indet. (Haeckel 1866)
Malafaia et al. 2006
Several individuals, incl. at least one partially complete skeleton
Reptilia - Rhynchocephalia
Sphenodontia indet. (Jaekel 1910)
Malafaia et al. 2006
Close relative of Opisthias
Amphibia indet. Linnaeus 1758
Malafaia et al. 2006
Plantae indet. Haeckel 1866
Malafaia et al. 2006
carbonized plant remains
see common names

Country:Portugal State/province:Estremdura County:Leiria
Coordinates: 39.9° North, 8.6° West (view map)
Paleocoordinates:33.8° North, 2.1° East
Basis of coordinate:based on nearby landmark
Altitude:212 meters
Geographic resolution:small collection
Period:Jurassic Epoch:Late/Upper Jurassic
10 m.y. bin:Jurassic 5
Key time interval:Late/Upper Oxfordian - Late/Upper Kimmeridgian
Age range of interval:161.20000 - 150.80000 m.y. ago
Stratigraphic resolution:bed
Stratigraphy comments: = Camadas de Alcobaça, could also be from lowermost Bombarral Formation (= Grès superiores); Malafaia et al 2010 consider the site to be ?upper Kimmeridgian to Tithonian in age
Lithology and environment
Primary lithology:paleosol/pedogenic,micaceous,gray carbonaceous sandstone
Secondary lithology:fine siltstone
Includes fossils?Y
Lithology description: "mainly represented by micaceous sandstones (as occur in Andrès), fine sands and silts", also described as "sandstones and marls that represent paleosols", also described as "fine-grained mud-sandstones, sometimes micaceous, with intercalations of some levels of marls, silts and clays, sometimes with abundant calcareous or limonitic nodules"
Environment:fluvial indet.
Geology comments: "deposited in a fluvial meandering environment with subsidiary deltaic and marginal marine environments.", also called a shallow, distal river setting
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Spatial orientation:life position
Preservation of anatomical detail:good
Articulated whole bodies:none
Associated major elements:many
Disassociated major elements:many
Disassociated minor elements:many
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:taxonomic analysis
Collectors:J. Amorim Collection dates:1988, 2005
Database number:38680
Authorizer:M. Carrano, P. Mannion Enterer:M. Carrano, J. Tennant
Modifier:M. Carrano Research group:vertebrate
Created:2004-04-28 10:46:44 Last modified:2019-09-25 09:26:01
Access level:the public Released:2004-04-28 10:46:44
Creative Commons license:CC BY
Reference information

Primary reference:

10381. B. P. Pérez-Moreno, D. J. Chure, C. Pires, C. Marques Silva, V. Dos Santos, P. Dantas, L. Póvoas, M. Cachão, J. L. Sanz and A. M. Galopim Carvalho. 1999. On the presence of Allosaurus fragilis (Theropoda: Carnosauria) in the Upper Jurassic of Portugal: first evidence of an intercontinental dinosaur species. Journal of the Geological Society, London 156:449-452 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

13166ETE J. F. Bonaparte and O. Mateus. 1999. A new diplodocid, Dinheirosaurus lourinhanensis gen. et sp. nov., from the Late Jurassic beds of Portugal. Revista del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigación de las Ciencias Naturales, Paleontología 5(2):13-29 [M. Carrano/M. Carrano/M. Carrano]
25000ETE E. Malafaia, P. Dantas, F. Ortega, F. Escaso, J. M. Gasulla, B. Ribeiro, F. Barriga, I. Gromicho, M. García-Oliva, G. Ramalheiro, J. Santamaría, N. L. Pimentel, C. Moniz and A. M. G. Carvalho. 2006. Análisis preliminar de la diversidad faunística en el yacimiento de Andrés (Jurásico Superior. Pombal, Portugal) [Preliminary analysis of faunal diversity in the Andrés locality (Upper Jurassic. Pombal, Portugal)]. IV Encuentro de Jóvenes Investigadores en Paleontología, Salamanca. Libro de Resúmenes 91-92 [M. Carrano/M. Carrano/M. Carrano]
24989ETE E. Malafaia, P. Dantas, F. Ortega and F. Escaso. 2007. Nuevos restos de Allosaurus fragilis (Theropoda: Carnosauria) del yacimiento de Andrés (Jurásico Superior; centro-oeste de Portugal) [New remains of Allosaurus fragilis (Theropoda: Carnosauria) from the Andres locality (Upper Jurassic; west-central Portugal)]. In Cantera Paleontológica, Diputación Provincial de Cuenca, Cuenca, Spain 255-271 [M. Carrano/M. Carrano]
44312 E. Malafaia, F. Ortega, F. Escaso, P. Dantas, N. Pimentel, J. M. Gasulla, B. Ribeiro, F. Barriga, and J. L. Sanz. 2010. Vertebrate fauna at the Allosaurus fossil-site of Andres (Upper Jurassic), Pombal, Portugal. Journal of Iberian Geology 36(2):193-204 [P. Mannion/J. Tennant/M. Carrano]
42539 P. D. Mannion, P. Upchurch, O. Mateus, R. N. Barnes, and M. E. H. Jones. 2012. New information on the anatomy and systematic position of Dinheirosaurus lourinhanensis (Sauropoda: Diplodocoidea) from the Late Jurassic of Portugal, with a review of European diplodocoids. Journal of Systematic Palaeontology 10(3):521-551 [P. Mannion/P. Mannion/M. Carrano]
19890ETE O. Mateus, A. Walen, and M. T. Antunes. 2006. The large theropod fauna of the Lourinhã Formation (Portugal) and its similarity to that of the Morrison Formation, with a description of a new species of Allosaurus. In J. R. Foster & S. G. Lucas (ed.), Paleontology and Geology of the Upper Jurassic Morrison Formation. New Mexico Museum of Natural History and Science Bulletin 36:123-129 [M. Carrano/M. Carrano/M. Carrano]
70303 J. L. Sanz. 2000. Yacimientos ibéricos de dinosaurios [Iberian dinosaur localities]. Dinosaurios: Los Señores del Pasado [Dinosaurs: The Lords of the Land] 267-299 [M. Carrano/M. Carrano]