Dolnice: MN 4, Czech Republic

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Squamata
Squamata informal indet. I Oppel 1811
Roček 1984 1 element
maxilla (DP FNSP 213)
Squamata informal indet. III Oppel 1811
Roček 1984 1 element
premaxilla (DP FNSP 196)
Reptilia - Squamata - Lacertidae
Lacertidae indet. Bonaparte 1831
Roček 1984 1 fragment
dentary fragment (DP FNSP 3773)
Lacerta sp. Linnaeus 1758
Roček 1984 52 elements
37 dentaries (DP FNSP 1, 3, 5, 7, 11, 14-17, 25, 27, 28, 30, 330, 50, 51, 63, 73, 77, 82, 83, 94, 214, 236, 265, 514,3772, 3775, 3780, 3787, 3800, 3824, 3849, 3851, 3857, 3895, 4368), 13 maxillae (DP FNSP 24, 43, 52, 74, 81, 98, 124, 1473, 3795, 3843, 3852, 3860), 1 pterygoid (DP FNSP 4320), 1 pelvis (DP FNSP 449)
? Lacerta sp. Linnaeus 1758
Roček 1984 42 fragments
specimens listed as Lacerta sp. but assigned tentatively to this taxon due to bad preservation: 35 dentaries (DP FNSP 12, 21, 23, 29, 38, 44, 47, 58, 62, 65, 68, 76, 88-90, 222, 242, 244, 245, 296, 518, 3778, 3784, 3789, 3790, 3810, 3822, 3827, 3828, 3830, 3834, 3835, 3876, 4305, 4309), 7 maxillae (DP FNSP 32, 70, 72, 3818, 3855, 3884, 4307)
Lacerta cf. viridis (Laurenti 1768)
Čerňanský 2010 3 specimens
Pb 02054, fused frontals; Pb 02055, fragment of left frontal; Pb 02024, fragment of posterior right portion of parietal
Miolacerta tenuis n. gen., n. sp. Roček 1984
Roček 1984 17 elements
Holotype: left dentary (DP FNSP 3785); paratypes: 2 dentaries (DP FNSP 3791, 4350); 14 referred specimens, mainly tooth bearing bones (DP FNSP 54, 55, 69, 75, 78, 130, 240, 243, 3811, 3847, 3850, 3868, 3873, 3910):
Amblyolacerta dolnicensis n. gen., n. sp. Roček 1984
Roček 1984 7 elements
Holotype: right maxilla (DP FNSP 4820); paratypes: 2 dentaries (DP FNSP 3809, 3837), 1 maxilla (DP FNSP 46); 3 referred specimens (DP FNSP 1485*, 3864, 4328; *"rather doubtful")
Reptilia - Squamata - Anguidae
Anguidae indet. Gray 1825
Roček 1984 54 elements
trunk vertebrae: 8 of "morphotype II" (DP FNSP 1286, 1287, 1299, 1313, 1315, 1418, 1454, 3961), 9 of "morphotype III" (DP FNSP 1308, 1415, 1487, 3934, 3943, 3948, 4006, 4013, 4015), 2 of "morphotype IV" (DP FNSP 3951, 4017), 1 of "morphotype VI" (DP FNSP 1310); caudal vertebrae: 9 of "morphotype I" (DP FNSP 1302, 1333, 1424, 1446, 3919, 3925, 3972, 3977, 3989), 12 of "morphotype II" (DP FNSP 1241, 1303, 1305, 1335, 1363, 1449, 1452, 1453, 3918, 3952, 3995, 3996), 2 of "morphotype III" (DP FNSP 1251, 3939), 2 of "morphotype IV" (DP FNSP 1244, 1289), 3 of "morphotype V" (DP FNSP 354, and probably also 3933, 3950), 1 of "morphotype VI" (DP FNSP 4002), 4 of "morphotype VII" (DP FNSP 1212, 1215, 1439, 3923), 1 of "morphotype VIII" (DP FNSP 3945)
Anguidae informal indet. I Gray 1825
Roček 1984 14 elements
14 dentigerous bones (DP FNSP 4, 45, 67, 1395, 3801, 3820, 3823, 3858, 3866, 4308, 4312, "and probably also" 35, 3786, 3816)
Anguidae informal indet. II Gray 1825
Roček 1984 1 element
1 maxilla (DP FNSP 93)
cf. Xestops sp. Cope 1873
Roček 1984 30 elements
8 dentaries (DP FNSP 18, 39, 48, 60, 3781, 3783, 3794, 3859), 9 maxillae (DP FNSP 129, 212, 219, 3813, 3819, 3821, 3831, 3914, 4306), 6 premaxillae (DP FNSP 194, 195, 345, 1552, 3814, 3878), 7 fragments of dentigerous bone (DP FNSP 37*, 96*, 224*,277, 1396*, 3829*, 3841*; * = "with certain doubts")
Pseudopus sp. Merrem 1820
Roček 1984 2 fragments
2 bones with marginal dentition (DP FNSP 218, 896)
Pseudopus ahnikoviensis Klembara 2012
Klembara 2012 4 elements
parietal (DP FNSP 1016, paratype, identified as Ophisaurus aff. spinari by Klembara, 1981), frontal (DP FNSP 1017, paratype, identified as Ophisaurus sp. by Klembara, 1981), 2 dentaries (DP FNSP 1030 and 1031, paratypes, identified as Pseudopus sp. by Klembara, 1981)
Ophisaurus informal sp. I Daudin 1803
Roček 1984 1 element
dentary (DP FNSP 3844)
Ophisaurus informal sp. II Daudin 1803
Roček 1984 3 elements
3 dentaries (DP FNSP 84*, 1472, 3846; * = "probably")
cf. Ophisaurus sp. Daudin 1803
Roček 1984 53 elements
49 trunk vertebrae of "morphotype I" (DP FNSP 1187, 1209, 1210, 1234, 1240, 1242, 1246, 1292, 1293, 1306, 1309, 1314, 1409, 1410, 1412, 1416, 1417, 1420, 1423, 1425, 1426, 1429, 1442, 1445, 1455, 1456, 1460, 3928, 3941, 3942, 3944, 3946, 3949, 3955, 3958, 3960, 3963, 3965, 3968, 3973, 3992, 3997, 4001, 4007, 4010, 4015, 4019, 4024, 4027) and 4 trunk vertebrae of "morphotype V" (DP FNSP 1214, 1450, 3937, 4016)
Anguis robustus n. sp. Klembara 1979
Klembara and Rummel 2018 2 specimens
recombined as Ophisaurus robustus
Specimens listed as O. robustus in this paper. DP FNSP 1018 (holotype), parietal; 1020, prev. ID'd as O. spinari
Ophisaurus feifari n. sp. Klembara 1979
Klembara and Rummel 2018 3 specimens
DP FNSP 1013 (holotype), parietal; FNSP 1501, parietal; 1023, parietal (previously ID'd as O. spinari)
Ophisaurus cf. spinari Klembara 1979
Roček 1984 24 elements
14 dentaries (DP FNSP 79, 80, 99, 121-123, 128, 246, 290, 3774, 3788, 3874, 3896, 3911); 6 maxillae (DP FNSP 9, 64, 117, 120, 125, 127), 4 pterygoids (DP FNSP 1512, 1518, 1550, 1558)
? Ophisaurus cf. spinari Klembara 1979
Roček 1984 4 fragments
"perhaps" Ophisaurus cf. spinari: 3 maxillae (DP FNSP 19, 41, 3871), 1 pterygoid (DP FNSP 1551)
Ophisaurus spinari Klembara 1979
Klembara and Rummel 2018 6 specimens
Parietals DP FNSP 1014, DP FNSP 1019, DP FNSP 1021, 1022, DP FNSP 1025, DP FNSP 1028
Ophisaurus holeci Klembara 2015
Klembara and Rummel 2018 3 specimens
Parietals DP FNSP 1026, ?DP FNSP 1027, ?DP FNSP 1029
Reptilia - Squamata - Chamaeleonidae
Chamaeleo caroliquarti n. sp. Moody and Roček 1980
Moody and Roček 1980 19 elements
nomen dubium belonging to Chamaeleo
DP FNSP 101 (holotype), left dentary; 102, midsection of right dentary; 103, left maxillary; 104, right maxillary; 105, section of max with 2 teeth; 106, midsection left max; 107, anteriormost section of right dentary w 7 teeth; 108, ant section of right dentary w 14 teeth; 109, section of left max incl palatine process; 110, ant part of right dentary; 111, midsection left dentary; 112, posterior section left dentary. Add'l specimens from ref 43973: 5 fragments of unidentified dentigerous bone (DP FNSP 113, 279, 512, 3867, 3869); 1 partial right maxilla (DP FNSP 4316), 1 postorbital (DP FNSP 226)
Chamaeleo andrusovi n. sp. Čerňanský 2010
Čerňanský 2010 5 specimens
Pb 02056 (holotype), nearly complete parietal but lacking the posterior portion; Right jugal (Pb 02057); left jugal (Pb 02058); left prefrontal (Pb 02059); postfrontal portion of the left postorbitofrontal (Pb 02050)
Reptilia - Squamata - Viperidae
Viperidae indet. Oppel 1811
4 elements
4 vertebrae (DP FNSP 1158, 3929, 4032, 4529)
Reptilia - Squamata - Boidae
cf. Gongylophis sp. Wagler 1830
14 elements
14 precaudal vertebrae (DP FNSP 1147, 1148, 1157-1159, 1167, 1489, 1490, 3921, 3922, 4005, 4084, 4533, 5270)
    = Bavarioboa hermi Szyndlar and Schleich 1993
Szyndlar and Schleich 1993
Bransateryx septentrionalis n. sp. Szyndlar 1987
7 elements
replaced by Falseryx petersbuchi
a single posterior caudal vertebra (DP FNSP 4555 = holotype) and 6 further vertebrae (DP FNSP 3916, 3990, 4003, 4528, 4556, 4557)
Reptilia - Squamata
Colubroidea indet. Oppel 1811
5 elements
5 caudal vertebrae (DP FNSP 1211, 1427, 1444, 1447, 3971)
Reptilia - Squamata - Colubridae
Colubridae indet. Oppel 1811
11 elements
11 precaudal vertebrae (DP FNSP 1155, 1374, 1432, 3938, 3959, 3964, 3966, 4200, 4231, 4530, 5207)
Palaeonatrix lehmani (Rage and Roček 1983)
7 elements
7 precaudal vertebrae (DP FNSP 933, 1408, 1434, 1437, 4096, 4553, 4554); the author does not say whether this material comes (at least in part) from the same stratigraphic level as the type material of Dolniceophis lehmani (see PBDB coll. 68711) or not
Texasophis bohemiacus n. sp. Szyndlar 1987
3 elements
3 trunk vertebrae (holotype: DP FNSP 1238; referred specimens: DP FNSP 1239, 3927)
? Texasophis bohemiacus Szyndlar 1987
2 elements
2 trunk vertebrae (DP FNSP 3947, 4025)
Coluber dolnicensis n. sp. Szyndlar 1987
3 elements
2 trunk vertebrae (holotype: DP NFSP 4558; referred specimen 4559), 1 compound bone (DP FNSP 310)
Neonatrix nova n. sp. Szyndlar 1987
63 elements
63 precaudal vertebrae (holotype: DP FNSP 5197; referred specimens: DP FNSP 931, 932, 934-937, 1171, 1237, 1245, 1253, 1311, 1312, 1414, 1419, 1430, 1433, 1440, 1443, 1451, 1458, 3962, 3967, 3979, 3983, 3987, 3988, 4026, 4029, 4030, 4075, 4107, 4201, 4511, 4526, 4532, 5156-5159, 5162, 5164, 5169, 5171-5173, 5175, 5176, 5182, 5186-5188, 5192, 5194-5196, 5198, 5200, 5202, 5204, 5214, 5221, 5224)
Reptilia - Squamata - Crotalidae
Vipera antiqua n. sp. Szyndlar 1987
6 elements
6 precaudal vertebrae (holotype: DP FNSP 4538; referred specimens: DP FNSP 938, 1114, 1213, 4014, 4057)
Vipera platyspondyla n. sp. Szyndlar 1987
8 elements
8 precaudal vertebrae (holotype: DP FNSP 940; referred specimens: 939, 1421, 1431, 1435, 3926, 4004, 4008)
Reptilia - Squamata
Scolecophidia indet. (Duméril and Bibron 1844)
1 element
a single broken vertebra (DP FNSP 1316)
Omoiotyphlops gracilis n. sp. Roček 1984
Roček 1984 41 elements
recombined as Blanus gracilis
Holotype: right dentary (DP FNSP 3771); paratypes: 2 left dentaries (DP FNSP 217, 3739) and 3 trunk vertebrae (DP FNSP 1317, 1336, 1364); referred material: 35 vertebrae (DP FNSP 1235, 1247, 1249, 1252, 1254, 1255, 1288, 1291, 1294-1296, 1298, 1300-1302, 1304, 1319, 1334, 1337, 1389, 1401, 1404, 1405, 1428, 1448, 1457, 3920, 3953, 3976, 3985, 3993, 3994, 3999)
Squamata informal indet. II Oppel 1811
Roček 1984 1 element
premaxilla (DP FNSP 317)
    = ? Blanus gracilis Roček 1984
Venczel and Stiuca 2008
belongs "quite probably" to this taxon
Reptilia - Squamata - Scincidae
cf. Scincidae indet. Gray 1825
Roček 1984 2 elements
2 maxillae (DP FNSP 59 "and perhaps" 61)
Reptilia - Squamata - Cordylidae
Palaeocordylus bohemicus n. gen., n. sp. Roček 1984
Roček 1984 28 elements
holotype: left dentary (DP FNSP 97); paratypes: 8 dentaries (DP FNSP 20, 57, 100, 119, 216, 294, 3798, 3804), 3 maxillae (DP FNSP 71, 86, 3802), 1 premaxilla (DP FNSP 114); referred specimens: 13 dentaries (DP FNSP 22, 49, 56, 85, 87, 3776, 3803, 3806, 3836, 3839, 3842, 38, 45, 3854), 2 maxillae (DP FNSP 3796, 3805)
? Palaeocordylus bohemicus Roček 1984
Roček 1984 13 fragments
specimens listed as P. bohemicus but "referred to this form with certain doubts" due to bad preservation: 9 dentaries (DP FNSP 40, 92, 268, 3779, 3815, 3840, 3848, 3856, 4350), 3 maxillae (DP FNSP 126, 232, 3792), 1 unidentified tooth bearing bone (DP FNSP 91)
see common names

Geography
Country:Czech Republic State/province:Karlovy Vary County:Cheb
Coordinates: 50.1° North, 12.4° East (view map)
Paleocoordinates:50.4° North, 10.9° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Burdigalian 10 m.y. bin:Cenozoic 5
Key time interval:MN 4
Age range of interval:16.90000 - 15.97000 m.y. ago
Stratigraphy
Stratigraphic resolution:member
Stratigraphy comments: The Dolnice locality belongs to the upper part of the so called "coal seam beds" and is located immediately above the main lignite seam with its grey and brown mudstones (so called "Kapuzin-Tone"). The "coal seam beds" are overlain by the muddy to clayey Cypris beds which are characterized by ostracode bearing freshwater limestone lenses (Fejfar, 1974).
Lithology and environment
Primary lithology: calcareous siltstone
Secondary lithology: silty,sandy marl
Includes fossils?Y
Includes fossils?Y
Environment:fluvial-lacustrine indet. Tectonic setting:rift
Geology comments: The locality is situated in the Cheb Basin at the intersection of the SW-NE trending Krušné Hory Graben (Ohře Graben, German: Eger-Graben) and the NW-SE trending Cheb-Domazlice Graben (see Reicherter et al., 2008, pp. 1233-1286 in McCann (ed.) The Geology of Central Europe, vol. 2).
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Fragmentation:frequent
Collection methods and comments
Collection methods:bulk,sieve,field collection,survey of museum collection
Reason for describing collection:taxonomic analysis
Collection method comments: This collection comprises fossils which have been collected during systematic excavations in the second half of the 20th century. All of the material is housed in the Department of Paleontology of the Faculty of Natural Sciences, Charles University, Prague (DP-FNSP).
Metadata
Also known as:Dölitz, Delitz
Database number:138666
Authorizer:J. Mueller, T. Cleary Enterer:T. Liebrecht, T. Cleary
Modifier:T. Liebrecht Research group:vertebrate
Collections that are a subset of this one:68711
Created:2013-01-25 00:01:26 Last modified:2013-01-29 15:03:26
Access level:the public Released:2013-01-25 00:01:26
Creative Commons license:CC BY
Reference information

Primary reference:

44422. Z. Szyndlar. 1987. Snakes from the Lower Miocene Locality of Dolnice (Czechoslovakia). Journal of Vertebrate Paleontology 7(1):55-71 [J. Mueller/T. Liebrecht]

Secondary references:

67867 A. Čerňanský. 2010. Earliest world record of green lizards (Lacertilia, Lacertidae) from the Lower Miocene of Central Europe. Biologia 65(4):737-741 [T. Cleary/T. Cleary]
67866 A. Čerňanský. 2010. A revision of chamaeleonids from the Lower Miocene of the Czech Republic with description of a new species of Chamaeleo (Squamata, Chamaeleonidae). Geobios 43:605-613 [T. Cleary/T. Cleary]
54648 J. Klembara. 2012. A new species of Pseudopus (Squamata, Anguidae) from the early Miocene of Northwest Bohemia (Czech Republic). Journal of Vertebrate Paleontology 32(4):854-866 [J. Mueller/T. Liebrecht]
69175 J. Klembara and M. Rummel. 2018. New material of Ophisaurus, Anguis and Pseudopus (Squamata, Anguidae, Anguinae) from the Miocene of the Czech Republic and Germany and systematic revision and palaeobiogeography of the Cenozoic Anguinae. Geological Magazine 155(1):20-44 [T. Cleary/T. Cleary]
28846 S. Moody and Z. Roček. 1980. Chamaeleo caroliquarti (Chamaeleonidae, Sauria): a new species from the Lower Miocene of Central Europe. Bulletin of the Czech Geological Survey 55(2):85-96 [J. Mueller/T. Liebrecht/M. Uhen]
43973 Z. Roček. 1984. Lizards (Reptilia: Sauria) from the Lower Miocene locality Dolnice (Bohemia, Czechoslovakia). Rozpravy Československé Akademie Věd / Řada matematických a přirodních věd 94(1):3-64 [J. Mueller/T. Liebrecht/M. Uhen]
7900 Z. Szyndlar and H. H. Schleich. 1993. Description of Miocene snake from Petersbuch 2 with comments on the lower and middle Miocene ophidian faunas of southern Germany. Stuttgarter Beitrage zur Naturkunde, Series B. Geologie und Palaontologie (192)1-47 [J. Head/J. Head/J. Head]
43465 M. Venczel and E. Stiuca. 2008. Late middle Miocene amphibians and squamate reptiles from Taut, Romania. Geodiversitas 30(4):731-763 [J. Mueller/T. Liebrecht/M. Uhen]