Hainin well vertebrate fauna: Danian, Belgium
collected by Geological laboratory of the Faculté Polytechnique de Mons 1970

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Different species of anurans and urodeles (Folie 2007 [SVP abstract])
Amphibia - Temnospondyli - Urodela
Urodela indet. (Duméril 1805)
Different species of anurans and urodeles (Folie 2007 [SVP abstract])
Reptilia
Eusuchia indet. (Huxley 1875)
Puértolas-Pascual et al. 2016
Alethinophidia indet. Nopcsa 1923
Rage 1998 4 specimens
Three fragmentary vertebrae and one fragmentary rib; two referred to cf. Dunnophis by Van Dyck (1983)
Scincoideus haininensis n. gen., n. sp. Folie et al. 2005
Folie et al. 2005
IRSNB R 219 - holotype (right dentary); paratypes: four maxillae and maxillary fragments (IRSNB R 215-218), eight dentaries and dentary fragments (IRSNB R 220-227), one pair of frontals (IRSNB R 228), two isolated frontals (IRSNB R 229-230) and one incomplete parietal (IRSNB R 231)
Amphisbaenidae indet. (Gray 1865)
Augé 2005 4 specimens
H13, 70-14, 01; H14-14, 60, 03; H14, 60-15, 10, 03; H14, 60-15, 10, 01: four dorsal vertebrae
Reptilia - Polyodontobaenidae
Polyodontobaena belgica n. gen., n. sp. Folie et al. 2013
Folie et al. 2013
IRSNB R 283 (HA 605) - holotype (nearly complete left dentary); paratype: IRSNB R 284 (HA 606), a right dentary; referred material: 13.7-14 (01), 16.4-15.10 (01) and HA 607- 639 (37 trunk vertebrae including HA 607/IRSNB R 285); HA 640-642 (3 cervical or caudal vertebrae including HA 640/IRSNB R 286)
Reptilia - Lacertidae
Lacertidae indet. Bonaparte 1831
Augé 2005
Reptilia - Anguidae
Anguini indet. Augé 2005
Augé 2005
dentary fragments
Reptilia - Testudines - Trionychidae
Groessens - van Dyck 1984
carapace fragments
? Palaeotrionyx sp. (Schmidt 1945)
Groessens - van Dyck 1984
original and current combination Paleotrionyx
costal fragments
Reptilia
cf. Baenoidea indet.
Groessens - van Dyck 1984
shell fragments
    = Laurasichersis sp. Pérez-García 2020
Pérez-García 2020
Mammalia - Multituberculata - Kogaionidae
Hainina godfriauxi n. sp. Vianey-Liaud 1979
Vianey-Liaud 1979
IRSNB M1968 - type; referred material: IRSNB M1955, 1965-1969 and 2191-2192
Hainina belgica n. gen., n. sp. Vianey-Liaud 1979
Vianey-Liaud 1979
IRSNB M1964 - holotype; referred material: IRSNB M1956-1963, 2185-2190
Mammalia - Multituberculata - Boffiidae
Boffius splendidus n. gen., n. sp. Vianey-Liaud 1979
Vianey-Liaud 1979
IRSNB M1948 - holotype; referred material: IRSNB M1949-1954
Mammalia - Pseudorhyncocyonidae
Pseudorhyncocyonidae indet. Sigé 1974
De Bast and Smith 2017
IRSNB M2224
Mammalia - Cimolesta
Cimolesta indet. McKenna 1975
De Bast and Smith 2017
IRSNB M2225
Mammalia - Cimolestidae
cf. Cimolestes sp. Marsh 1889
De Bast and Smith 2017
IRSNB M2198
Mammalia - Pentacodontidae
Eurolestes dupuisi n. gen., n. sp. De Bast and Smith 2017
De Bast and Smith 2017
IRSNB M2203 - holotype; referred material: IRSNB M1945 and 2199-2205
Mammalia - Macroscelidea - Louisinidae
Cingulodon magioncaldai n. gen., n. sp. De Bast and Smith 2017
De Bast and Smith 2017
IRSNB M2223 - holotype
Monshyus praevius n. gen., n. sp. Sudre and Russell 1982
Sudre and Russell 1982
IRSNB M1970 - holotype (left M1 or M2); referred material: R1-7 (right DP4)
Mammalia - Condylarthra - Pleuraspidotheriidae
? Pleuraspidotheriidae indet. Zittel 1892
De Bast and Smith 2017
IRSNB M2222
Mammalia - Procreodi
Prolatidens waudruae n. gen., n. sp. Sudre and Russell 1982
Sudre and Russell 1982
IRSNB M1972 - holotype (left M1); referred material: IRSNB M2053 (= N2-15), left P4; IRSNB M2054 (= Q2-19), leftM1 orM2; IRSNBM2055 (= Q1-16), right M1 or M2; IRSNB M2056 (= O1-05), left dp4; IRSNB M2115 (= MN1.2-03), left m1; IRSNB M2116 (= N2-01), right m1; IRSNB M1973 (= O1-01), left m2; IRSNB M2117 (= N1-05), right m1 or m2 (De Bast and Smith 2013)
Mammalia
Quadratodon sigei n. gen., n. sp. De Bast and Smith 2017
De Bast and Smith 2017
IRSNB M2221 - holotype; paratypes: IRSNB M2218 and 2220; referred material: IRSNB M2216-2219
Mammalia - Primates - Picrodontidae
Russellodon haininense n. gen., n. sp. Sigé and Marandat 1997
De Bast and Smith 2017
IRSNB M1977 - holotype; referred material: IRSNB M2211 and 2212
Mammalia - Adapisoriculidae
Bustylus folieae n. sp. De Bast et al. 2012
IRSNB M1996 - type (left M1); and numerous referred material
Peradectes marandati n. sp. Crochet and Sigé 1983
Crochet and Sigé 1983
recombined as Bustylus marandati
IRSNB M1974 - type (right M2); and numerous referred material
? Adapisoriculus sp. Lemoine 1885
IRSNB M2009 (right M3)
    = cf. Adapisoriculus sp. Lemoine 1885
De Bast and Smith 2017
IRSNB M2001
Proremiculus lagnauxi n. gen., n. sp. De Bast et al. 2012
IRSNB M2006 - holotype (left M2); and numerous referred material
Afrodon gheerbranti n. sp. De Bast et al. 2012
IRSNB M1982 - type (right M2); paratypes: IRSNB M1982-1989
Mammalia - Proteutheria - Palaeoryctidae
Belgoryctes thaleri n. gen., n. sp. De Bast and Smith 2017
De Bast and Smith 2017
IRSNB M2193 - holotype; referred material: IRSNB M1947 and IRSNB M2194-2197
see common names

Geography
Country:Belgium State/province:Hainaut
Coordinates: 50.4° North, 3.8° East (view map)
Paleocoordinates:43.8° North, 2.1° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Paleogene Epoch:Paleocene
Stage:Danian 10 m.y. bin:Cenozoic 1
Key time interval:Danian
Age range of interval:66.00000 - 61.60000 m.y. ago
Stratigraphy
Formation:Hainin
Stratigraphic resolution:group of beds
Stratigraphy comments: Reference-level MP1-5 of the mammalian biochronological scale for the European Palaeogene.

Although not precisely dated, has been correlated with the earliest Selandian based on stratigraphy (Steurbaut 1998), and more recently to the latest Danian (Steurbaut & Sztrakos 2008).

The borehole that yielded the fossils encountered continental deposits between 8.50 m and 38.52 m in depth. Of the 30 m of continental deposits, only the section between 11 and 25 m yielded vertebrates (De Bast and Smith 2016).
Lithology and environment
Primary lithology: claystone
Includes fossils?Y
Environment:lacustrine indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:core,chemical
Reason for describing collection:taxonomic analysis
Collectors:Geological laboratory of the Faculté Polytechnique de Mons Collection dates:Early 1970s
Collection method comments: The sediments collected from the Hainin well went through a first step of screen washing. The remaining hard parts were treated with formic acid (5%) with tricalcic phosphate (Ca3(PO4)2) used as a buffer. A second step of screenwashing was then performed.
Metadata
Also known as:Mons Basin
Database number:132949
Authorizer:P. Mannion, P. Barrett, E. Vlachos Enterer:P. Mannion, T. Cleary, E. Vlachos
Modifier:P. Mannion Research group:vertebrate
Created:2012-08-30 01:04:42 Last modified:2016-11-14 08:02:14
Access level:the public Released:2012-08-30 01:04:42
Creative Commons license:CC BY
Reference information

Primary reference:
42813. E. De Bast, B. Sigé, and T. Smith. 2012. Diversity of the adapisoriculid mammals from the early Palaeocene of Hainin, Belgium. Acta Palaeontologica Polonica 57(1):35-52 [P. Mannion/P. Mannion]

Secondary references:

59290 M. L. Augé. 2005. Evolution des lézards du Paléogène en Europe. Mémoires du Muséum national d'histoire naturelle 192:1-369 [P. Barrett/T. Cleary/T. Cleary]
61389 J.-Y. Crochet and B. Sigé. 1983. Les mammifères Montiens de Hainin (Paléocène moyen de Belgique) Part III : Marsupiaux. Palaeovertebrata 13(3):51-64 [P. Mannion/P. Mannion]
47335 E. De Bast and T. Smith. 2013. Reassessment of the small ‘arctocyonid’ Prolatidens waudruae from the early Paleocene of Belgium, and its phylogenetic relationships with ungulate-like mammals. Journal of Vertebrate Paleontology 33(4):964-976 [P. Mannion/P. Mannion]
60813 E. De Bast and T. Smith. 2017. The oldest Cenozoic mammal fauna of Europe: implication of the Hainin reference fauna for mammalian evolution and dispersals during the Paleocene. Journal of Systematic Palaeontology 15(9):741-785 [P. Mannion/P. Mannion/P. Mannion]
42814 A. Folie, B. Sigé, and T. Smith. 2005. A new scincomorph lizard from the Palaeocene of Belgium and the origin of Scincoidea in Europe. Naturwissenschaften 92:542-546 [P. Mannion/P. Mannion]
54400 A. Folie, R. Smith, and T. Smith. 2013. New amphisbaenian lizards from the Early Paleogene of Europe and their implications for the early evolution of modern amphisbaenians. Geologica Belgica 16(4):227-235 [P. Mannion/P. Mannion]
42887 E. Gheerbrant. 1991. Bustylus (Eutheria, Adapisoriculidae) and the absence of ascertained Marsupials in the Palaeocene of Europe. Terra Nova 3:586-592 [P. Mannion/P. Mannion]
65726 M. C. Groessens - van Dyck . 1984. Les tortues du Paleocene continental de Hainin et Vinalmont (Belqique). Studia Geologica Salmanticensia, volumen especial 1 (Studia Palaeocheloniologica I) 1:133-139 [E. Vlachos/E. Vlachos]
71657 A. Pérez-García. 2020. Surviving the Cretaceous-Paleogene mass extinction event: A terrestrial stem turtle in the Cenozoic of Laurasia. Scientific Reports 10:1489 [P. Mannion/P. Mannion]
76284 E. Puértolas-Pascual, A. Blanco, C. A. Brochu and J. I. Canudo. 2016. Review of the Late Cretaceous-early Paleogene crocodylomorphs of Europe: Extinction patterns across the K-PG boundary. Cretaceous Research 57:565-590 [P. Mannion/P. Mannion]
12895 J.-C. Rage. 1998. Fossil snakes from the Palaeocene of São José de Itaboraí, Brazil. Part I. Madtsoiidae, Aniliidae. Palaeovertebrata 27(3-4):109-144 [J. Head/J. Head/J. Head]
46225 J. Sudre and D. E. Russell. 1982. Les mammifères montiens de Hainin (Paléocène moyen de Belgique). Part 2: les condylarthres. Palaeovertebrata 12(6):173-184 [P. Mannion/P. Mannion]
61387 M. Vianey-Liaud. 1979. Les mammifères Montiens de Hainin (Paléocène moyen de Belgique) Part 1 : Multituberculés. Palaeovertebrata 9(4):117-131 [P. Mannion/P. Mannion]