Right bank Ifasy River, NE of Anaborano: Induan - Smithian, Madagascar
collected 1990

List of taxa
Where & when
Taphonomy & methods
Metadata & references
Taxonomic list
cf. Tertremoides madagascariensis (Lehman 1966)
1 specimen
MSNM V2993, incomplete skull
Aphaneramma gavialimimus n. sp. Fortuny et al. 2017
Fortuny et al. 2017 1 specimen
Edingerella madagascariensis (Lehman 1961)
Maganuco et al. 2009 1 specimen
MSNM V2992, mould of complete skull
see common names

Country:Madagascar State/province:Diana County:Ambilobe
Coordinates: 13.5° South, 48.8° East (view map)
Paleocoordinates:44.5° South, 41.3° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Period:Triassic Epoch:Early/Lower Triassic
10 m.y. bin:Triassic 1
Key time interval:Induan - Smithian
Age range of interval:252.17000 - 247.20000 m.y. ago
Geological group:Sakamena Formation:Middle Sakamena
Stratigraphic resolution:group of beds
Stratigraphy comments: "All previously described tetrapod specimens from the Diego Basin have been collected from the 'Couches à Poissons et Ammonites' (Besairie 1971) suggesting that the material described herein also pertains to this horizon, though this cannot be stated with absolute certainty." The "Couches à Poissons et Ammonites" are equivalent to the Middle Sakamena Formation of the Morondava Basin. "Biostratigraphical correlations based on ammonite faunas indicate referral of the 'Couches à Poissons et Ammonites' to the local Gyronitien Stage (Besairie 1971), which is equivalent to the Induan Stage of the Lower Triassic. This conclusion is supported by comparisons between the vertebrate faunas of Madagascar and the African mainland, which suggest that the Middle Sakamena Formation is intermediate in age between the Lystrosaurus and Cynognathus Assemblage Zones (Battail et al. 1987). Palynological data is also consistent with these interpretations (Wescott and Diggens 1998)." (Ketchum & Barrett, 2004. Canadian Journal of Earth Sciences 41:1-8).
"The presence of Magniestheria truempyi in the same formation (Shen et al., 2002) indicates an Early Smithian (Olenekian, Early Triassic) age by correlation with the well-constrained Triassic strata of the Germanic Basin (Kozur and Weems, 2010)." (Falconnet et al. 2010. Comptes Rendus Palevol 11:357-369)
Given the conflicting suggestions of an Induan or early Olenekian age, an Induan–Smithian range is given here.
Lithology and environment
Primary lithology:nodular "siliciclastic"
Includes fossils?Y
Lithology description: Fossil preserved as a natural mould in a siliceous nodule
Environment:deltaic indet.
Modes of preservation:mold/impression,concretion
Size of fossils:macrofossils
Preservation of anatomical detail:medium
Collection methods and comments
Collection methods:surface (float),survey of museum collection
Reason for describing collection:taxonomic analysis
Collection dates:1990s
Collection method comments: MSNM: Museo di Storia Naturale di Milano
Database number:144100
Authorizer:R. Butler, G. Lloyd Enterer:R. Butler, G. Lloyd
Modifier:M. Carrano Research group:vertebrate
Created:2013-04-30 21:44:11 Last modified:2018-02-01 13:03:25
Access level:the public Released:2013-04-30 21:44:11
Creative Commons license:CC BY
Reference information

Primary reference:

46594. S. Maganuco and G. Pasini. 2009. A new specimen of trematosaurian temnospondyl from the Lower Triassic of NW Madagascar, with remarks on palatal anatomy and taxonomic affinities. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 150:91-112 [R. Butler/R. Butler/R. Butler]

Secondary references:

65071 J. Fortuny, S. Gastou, F. Escuillié, L. Ranivoharimanana, and J.-S. Steyer. 2017. A new extreme longirostrine temnospondyl from the Triassic of Madagascar: phylogenetic and palaeobiogeographical implications for trematosaurids. Journal of Systematic Palaeontology 16:675-688 [G. Lloyd/G. Lloyd]
46595 S. Maganuco, J. S. Steyer, G. Pasini, M. Boulay, S. Lorrain, A. Beneteau, and M. Auditore. 2009. An exquisite specimen of Edingerella madagascariensis (Temnospondyli) from the Lower Triassic of NW Madagascar; cranial anatomy, phylogeny, and restorations. Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 36:1-72 [R. Butler/R. Butler]