PBDB Collection

Mazon Creek (YPM collection): Westphalian D, Illinois
collected by S. S. Strong

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Coelacanthimorpha - Coelacanthiformes - Coelacanthidae

	Coelacanthus exiguus Eastman 1902	Eastman 1903
Dipnoi - Sagenodontidae

	Sagenodus cristatus n. sp. (Agassiz 1838)	Eastman 1903
	original and current combination Ctenodus cristatus
Haplolepidae

	Pyritocephalus gracilis Westoll 1944	Westoll 1944
	YPM 440
Elonichthyiformes - Elonichthyidae

	Elonichthys perpennatus Eastman 1902	Eastman 1903

	Elonichthys disjunctus n. sp. Eastman 1903	Eastman 1903
Palaeonisciformes - Platysomidae

	Platysomus circularis Newberry and Worthen 1870	Eastman 1903
Eurynotiformes - Amphicentridae

	Platysomus orbicularis Newberry and Worthen 1870	Eastman 1903
	recombined as Amphicentrum orbiculare
Insecta

	Sypharoptera pneuma n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 64 (2 measurements)
Insecta - Meganisoptera

	Paralogopsis longipes n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 65 (2 measurements)
Insecta - Syntonopteridae

	Syntonoptera schucherti n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 19 (1 measurement)
Insecta - Megasecoptera

	Lameereites curvipennis n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 66 "very shadowy fossil showing four wings of nymph…this highly interesting fossil…may be called in honor of my highly estimable opponent, Lameereites"
Insecta - Diaphanopterodea - Prochoropteridae

	Prochoroptera calopteryx n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 67 (3 measurements)
Insecta - Palaeodictyoptera

	Diexodus debilis n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 21
Insecta - Palaeodictyoptera - Homoiopteridae

	Scepasma gigas n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 22 (1 measurement)

	Ametretus laevis n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 23 (1 measurement)

	Amousus mazonus n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 20 (1 measurement)
Insecta - Palaeodictyoptera - Eubleptidae

	Athymodictya parva n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Eubleptus danielsi
	YPM 18 (2 measurements)
Insecta - Protorthoptera - Eucaenidae

	Eucaenus ovalis Scudder 1885	Handlirsch 1911	3 individuals
	YPM 47-49 (3 measurements)

	Eucaenus mazonus Melander 1903	Handlirsch 1911	1 individual
	synonym of Eucaenus ovalis
	YPM 51 (2 measurements)

	Eucaenus minor n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Eucaenus ovalis
	YPM 50 (1 measurement)
Insecta - Protorthoptera

	Eucaenus pusillus n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	nomen dubium belonging to Protorthoptera
	YPM 52 (2 measurements)
Insecta - Protorthoptera - Anthracothremmidae

	Pericalyphe longa n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 54 (3 measurements)

	Silphion latipenne n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 56 (1 measurement)

	Melinophlebia analis n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 55 (1 measurement)
Insecta - Protorthoptera

	Schuchertiella gracilis n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 32 (1 measurement)
Insecta - Protorthoptera - Apithanidae

	Apithanus jocularis n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 39 (1 measurement)
Insecta - Grylloblattodea - Epideigmatidae

	Epideigma elegans n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 45 (1 measurement)
Insecta - Cheliphlebiidae

	Cheliphlebia mazona n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 46 (2 measurements)
Insecta - Phyloblattidae

	Phyloblatta diversipennis n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 60 (1 measurement)
Insecta - Blattodea - Gyroblattidae

	Megablattina beecheri n. gen., n. sp. Sellards 1903	Sellards 1903	1 specimen
	nomen dubium belonging to Progonoblattina
	(1 measurement)
Insecta - Blattodea - Archimylacridae

	Etoblattina mazona Scudder 1882	Sellards 1904
	recombined as Asemoblatta mazona
Insecta - Blattodea - Mylacridae

	Platymylacris paucinervis n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 62 (1 measurement)

	Orthomylacris contorta n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 61 (1 measurement)

	Mylacris diplodiscus (Packard 1885)	Sellards 1904

	Mylacris elongata Scudder 1895	Sellards 1904
	(1 measurement)
	= ? Mylacris sellardsii n. sp. Handlirsch 1906	Handlirsch 1906
	corrected as Mylacris sellardsi
Insecta - Blattodea - Mesoblattinidae

	Blattoidea sellardsi n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	recombined as Pareinoblatta sellardsi
	M. diplodiscus in Sellards, 1904 fig 3
Insecta - Blattodea

	Blattoidea sp. Handlirsch 1906	Handlirsch 1906	3 specimens
	E. mazona in Sellards, 1904 fig 10-12

	Mylacris anceps n. sp. Sellards 1904	Sellards 1904
	recombined as Blattoidea anceps

	Blattoidea schuchertiana n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	M. diplodiscus in Schuchert, 1897 fig 4, 5

	Dipeltis carri n. sp. Schuchert 1897	Handlirsch 1906	1 specimen
	recombined as Blattoidea carri

	"Blattoidea" melanderi Handlirsch 1906	Handlirsch 1911	1 individual
	YPM 63 unidentifiable larva (1 measurement)

	Blattoidea sellardsiana n. sp. Handlirsch 1906	Handlirsch 1906	1 specimen
	M. elongata in Sellards, 1904 fig 6, 7
Insecta - Notoptera - Adeloblattidae

	Etoblattina ? hilliana Scudder 1895	Sellards 1904	1 specimen
	(1 measurement)
	= Adeloblatta sellardsi Handlirsch 1906	Handlirsch 1911
	YPM 59
Insecta - Geraridae

	Gerarus collaris n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 34 (1 measurement)

	Gerarus longicollis n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Gerarus collaris
	YPM 36 (1 measurement)

	Gerarus latus n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Gerarus constrictus
	YPM 33 (1 measurement)

	? Gerarus reductus n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Gerarus constrictus
	YPM 35 (1 measurement)

	Anepitedius giraffa n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 38 (1 measurement)

	Gerarulus radialis n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 37 (1 measurement)
Insecta - Cacurgidae

	Spilomastax oligoneurus n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 42 (1 measurement)

	Cacurgus spilopterus n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 41 (1 measurement)
Insecta - Cnemidolestodea - Cnemidolestidae

	Narkema taeniatum n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 40 (1 measurement)
Insecta - Cnemidolestodea - Spanioderidae

	Spaniodera angusta n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Miamia bronsoni
	YPM 30 (1 measurement)

	Spaniodera parvula n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Miamia bronsoni
	YPM 29 (1 measurement)

	Spaniodera elatior n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Miamia bronsoni
	YPM 26 (1 measurement)

	Spaniodera acutipennis n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Miamia bronsoni
	YPM 28 (1 measurement)

	Spaniodera longicollis n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Miamia bronsoni
	YPM 24 (1 measurement)

	Spaniodera schucherti n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Miamia bronsoni
	YPM 27 (1 measurement)

	Spaniodera lata n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	synonym of Miamia bronsoni
	YPM 25 (1 measurement)

	Dieconeura mazona n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 31 (1 measurement)
Insecta - Eoblattidae

	Anegertus cubitalis n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 43 (1 measurement)
Insecta - Eoblattida

	"Protoblattoidea" sellardsi Handlirsch 1906	Handlirsch 1911	1 individual
	synonym of Eoblattida
	YPM 57 unidentifiable larva

	"Protoblattoidea" minor Handlirsch 1906	Handlirsch 1911	1 individual
	synonym of Eoblattida
	YPM 58 unidentifiable larva
Insecta - Eoblattida - Asyncritidae

	Asyncritus reticulatus n. gen., n. sp. Handlirsch 1911	Handlirsch 1911	1 individual
	YPM 44 (2 measurements)
Xiphosura - Xiphosurida - Belinuridae

	Pringlia bispinosa n. sp. Raymond 1944	Raymond 1944
	synonym of Liomesaspis laevis

	Euproops danae (Meek and Worthen 1865)	Haug and Rötzer 2018

	Euproops laevicula n. sp. Raymond 1944	Raymond 1944
	synonym of Euproops danae
Arachnida - Phalangiotarbida - Architarbidae

	Discotarbus deplanatus n. gen., n. sp. Petrunkevitch 1913	Petrunkevitch 1913	2 specimens
	YPM 174, 175

	Metatarbus triangularis n. gen., n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	YPM 182

	Architarbus rotundatus Scudder 1868	Petrunkevitch 1913	3 specimens
	YPM 185, 186, 187

	Architarbus minor n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	YPM 189

	Geratarbus minutus n. sp. Petrunkevitch 1913	Petrunkevitch 1913	6 specimens
	recombined as Orthotarbus minutus
	YPM 176-181
Arachnida - Trigonotarbida - Aphantomartidae

	Trigonomartus pustulatus (Scudder 1884)	Petrunkevitch 1913	1 specimen
	recombined as Aphantomartus pustulatus
	YPM 168
Arachnida - Trigonotarbida - Lissomartidae

	Trigonotarbus schucherti n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	recombined as Lissomartus schucherti
	YPM169
Arachnida - Thelyphonida - Thelyphonidae

	Geralinura gigantea Petrunkevitch 1913	Petrunkevitch 1913	2 specimens
	synonym of Geralinura carbonaria
	YPM 147; YPM 596 (formerly from Langford collection in Petrunkevitch 1949 and Tetlie & Dunlop 2008)
Arachnida - Araneae - Arthrolycosidae

	Arthrolycosa antiqua n. gen., n. sp. Harger 1874		2 specimens
	YPM 161; 162, 163 (in Petrunkevitch, 1913) (not 162, according to Selden 2021)
Arachnida - Araneae - Palaeothelidae

	Palaeothele onoi n. sp. Selden 2021	Selden 2021	1 specimen
	YPM IP.00162 (part of Arthrolycosa antiqua in Petrunkevitch 1913)
Arachnida - Solifugae

	Protosolpuga carbonaria n. gen., n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	YPM 155
Arachnida - Scorpiones - Eoctonidae

	Eoctonus miniatus n. gen., n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	YPM 131 (not YPM 132)
Arachnida - Scorpiones - Anthracoscorpionidae

	? Anthracoscorpio sp. Kušta 1889	Kjellesvig-Waering 1986	1 specimen
	YPM 126 (Eoscorpius typicus in Petrunkevitch 1913)
Arachnida - Scorpiones - Palaeobuthidae

	Eoscorpius typicus Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	YPM 127
	= Palaeobuthus distinctus Petrunkevitch 1913	Kjellesvig-Waering 1986

	Palaeobuthus distinctus n. gen., n. sp. Petrunkevitch 1913	Petrunkevitch 1913	2 specimens
	YPM 133; YPM 130 (in Kjellesvig-Waering 1986, Eoscorpius granulatus in Petrunkevitch, 1913)
Arachnida - Scorpiones - Palaeopisthacanthidae

	Palaeopisthacanthus schucherti n. gen., n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	YPM 140
Arachnida - Scorpiones - Paraisobuthidae

	Paraisobuthus virginiae n. sp. Kjellesvig-Waering 1986	Kjellesvig-Waering 1986	1 specimen
	YPM 132 (Eoctonus miniatus in Petrunkevitch, 1913)
Arachnida - Scorpiones - Telmatoscorpionidae

	Telmatoscorpio brevipectus n. gen., n. sp. Kjellesvig-Waering 1986	Kjellesvig-Waering 1986	1 specimen
	YPM 129 (Eoscorpius granulosus in Petrunkevitch, 1913)
Arachnida - Scorpiones - Eoscorpiidae

	Eoscorpius granulosus n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	synonym of Eoscorpius carbonarius
	YPM 128 (not YPM 129 or 130)
Arachnida - Opiliones - Nemastomoididae

	Protopilio longipes n. gen., n. sp. Petrunkevitch 1913	Petrunkevitch 1913	1 specimen
	recombined as Nemastomoides longipes
	YPM 171
Myriapoda - Pleurojulida - Pleurojulidae

	Pleurojulus cf. biornatus Fritsch 1899	Wilson and Hannibal 2005	1 specimen
	YPM 42861
Myriapoda - Zosterogrammida - Zosterogrammidae

	Zosterogrammus stichostethus n. gen., n. sp. Wilson 2005	Wilson 2005	4 specimens
	YPM 204068, 204067, 19819, 204066
see common names

Geography

Country:	United States	State/province:	Illinois	County:	Grundy
Coordinates:	41.3° North, 88.4° West (view map)
Paleocoordinates:	4.6° South, 23.4° West
Basis of coordinate:	estimated from map
Geographic resolution:	local area

Time

Period:	Carboniferous	Epoch:	Pennsylvanian
Stage:	Moscovian	10 m.y. bin:	Carboniferous 4
Key time interval:	Westphalian D
Age range of interval:	309.80000 - 307.40000 m.y. ago

Stratigraphy

Geological group:	Kewanee	Formation:	Carbondale	Member:	Francis Creek Shale
Local section:	lower Francis Creek Member	Local bed:	lower Francis Creek Member

Lithology and environment

Primary lithology:	concretionary,ferruginous lithified mudstone
Lithology description: ironstone concretions
Environment:	deltaic indet.	Tectonic setting:	cratonic basin
Geology comments: Illinois basin

Taphonomy

Modes of preservation:	mold/impression
Size of fossils:	macrofossils

Collection methods and comments

Collection size:	79 specimens
Reason for describing collection:	taxonomic analysis
Museum repositories:	YPM
Collectors:	S. S. Strong

Metadata

Database number:	112630
Authorizer:	M. Clapham, D. Smith, J. Flannery-Sutherland, C. Labandeira	Enterer:	J. Karr, A. Kinchloe, J. Flannery-Sutherland, P. Vazquez, M. Clapham, C. Labandeira
Modifier:	M. Clapham	Research group:	paleoentomology,vertebrate
Created:	2011-07-14 00:42:55	Last modified:	2020-12-21 18:56:05
Access level:	the public	Released:	2011-07-14 00:42:55
Creative Commons license:	CC BY

Reference information

Primary reference:

47345.

O. Harger. 1874. Notice of a new fossil spider from the Coal Measures of Illinois. The American Journal of Science and Arts, Third Series 7:219-223 [M. Clapham/A. Gomez/M. Clapham]

Secondary references:

9323		J. A. Dunlop. 1995. Redescription of the Pennsylvanian trigonotarbid arachnid Lissomartus Petrunkevitch 1949 from Mazon Creek, Illinois. The Journal of Arachnology 23(2):118-124 [D. Smith/A. Kinchloe/M. Clapham]
42819		C. R. Eastman. 1903. Carboniferous fishes from the central western States. Bulletin of the Museum of Comparative Zoology 39:163-226 [M. Clapham/P. Vazquez/M. Clapham]
35209		A. Handlirsch. 1906. Die Fossilen Insekten und die Phylogenie der Rezenten Formen, parts I-IV. Ein Handbuch fur Palaontologen und Zoologen 1-640 [M. Clapham/M. Clapham/M. Clapham]
37094		A. Handlirsch. 1906. Revision of American Paleozoic insects. Proceedings of the United States National Museum 29(1441):661-820 [M. Clapham/M. Clapham]
36675		A. Handlirsch. 1911. New Paleozoic insects from the vicinity of Mazon Creek, Illinois. The American Journal of Science, Fourth Series 31:353-377 [M. Clapham/J. Karr/M. Clapham]
5080	ETE 792	A. Handlirsch. 1911. New Paleozoic insects from the vicinity of Mazon Creek, Illinois. American Journal of Science, Series 4 31:297-326 [C. Labandeira/C. Labandeira/M. Clapham]
74853		C. Haug and M. A. I. Rötzer. 2018. The ontogeny of the 300 million year old xiphosuran Euproops danae (Euchelicerata) and implications for resolving the Euproops species complex. Development Genes and Evolution 228:63-74 [M. Clapham/M. Clapham]
49610		E. N. Kjellesvig-Waering. 1986. A restudy of the fossil Scorpionida of the world. Palaeontographica Americana 55:1-287 [M. Clapham/M. Clapham]
9398		A. I. Petrunkevitch. 1913. A monograph of the terrestrial Palaeozoic Arachnida of North America. Transactions of the Connecticut Acadamy of Arts and Sciences 18:1-137 [D. Smith/A. Kinchloe/M. Clapham]
74929		P. E. Raymond. 1944. Late Paleozoic xiphosurans. Bulletin of the Museum of Comparative Zoology 94:475-508 [M. Clapham/M. Clapham]
76590		P. A. Selden. 2021. New spiders (Araneae: Mesothelae), from the Carboniferous of New Mexico and England, and a review of Paleozoic Araneae. New Mexico Museum of Natural History and Science Bulletin 84:317-358 [M. Clapham/M. Clapham]
48173		E. H. Sellards. 1903. Some new structural characters of Paleozoic cockroaches. The American Journal of Science, Fourth Series 15:307-315 [M. Clapham/M. Clapham/M. Clapham]
36704		E. H. Sellards. 1904. A study of the structure of Paleozoic Cockroaches, with descriptions of new forms from the Coal Measures. The American Journal of Science, Fourth Series 18(105):213-227 [M. Clapham/J. Karr/M. Clapham]
36710		E. H. Sellards. 1904. A study of the structure of Paleozoic Cockroaches, with descriptions of new forms from the Coal Measures. American Journal of Science, Series 4 18(104):113-134 [M. Clapham/J. Karr/M. Clapham]
49581		O. E. Tetlie and J. A. Dunlop. 2008. Geralinura carbonaria (Arachnida: Uropygi) from Mazon Creek, Illinois, USA, and the origin of subchelate pedipalps in whip scorpions. Journal of Paleontology 82:299-312 [M. Clapham/M. Clapham]
84231		T. S. Westoll. 1944. The Haplolepidae, a new family of late Carboniferous bony fishes : a study in taxonomy and evolution. Bulletin of the AMNH ; v. 83, article 1 . Bulletin of the American Museum of Natural History 83(1):1-122 [E. Dunne/S. Fasey]
51032		H. M. Wilson. 2005. Zosterogrammida, a new order of millipedes from the middle Silurian of Scotland and the upper Carboniferous of Euramerica. Palaeontology 48:1101-1110 [M. Clapham/M. Clapham]
51029		H. M. Wilson and J. T. Hannibal. 2005. Taxonomy and trunk-ring architecture of pleurojulid millipedes (Diplopoda: Chilognatha: Pleurojulida) from the Pennsylvanian of Europe and North America. Journal of Paleontology 79:1105-1119 [M. Clapham/M. Clapham]