Barrihonda–El Humero, RD-10: Late/Upper Kimmeridgian - Tithonian, Spain
collected 2005–2009

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Bivalvia - Trigoniida
? Trigonioidoidea indet. Cox 1952
Delvene et al. 2020 5 specimens
CPT-1516, 1517, 4403–4405
Bivalvia - Unionida
Unionoida indet. Stoliczka 1871
Delvene et al. 2020 3 specimens
corrected as Unionida
CPT-1514 (1-2), 1563
Osteichthyes
Osteichthyes indet. Huxley 1880
"fishes"
Reptilia - Pleurosternidae
Riodevemys inumbragigas n. gen., n. sp. Pérez-García et al. 2015
Pérez-García et al. 2015 1 individual
CPT-1423 (housed at MAP), a partial skeleton including the disjointed carapace, the articulated plastron, and several bones of the scapular and pelvic girdles
Reptilia - Testudines
Testudines indet. Batsch 1788
"turtles"
Reptilia - Crocodylomorpha
Crocodylomorpha indet. (Hay 1930)
"crocodylomorphs"
Reptilia
Dinosauria indet. (Owen 1842)
Luque et al. 2009 6 specimens
CPT-1179, 1142, 1121, 1132, 1228, 1137
Ornithopoda indet. Marsh 1881
1 individual
"isolated elements of ornithopods"
    = Ankylopollexia indet. Sereno 1986
Cobos et al. 2014
    = Oblitosaurus bunnueli n. gen., n. sp. Sánchez-Fenollosa et al. 2023
Sánchez-Fenollosa et al. 2023
A dentary tooth (CPT-1440), a right ungual pollex from manus (CPT-1444), and an almost complete left hindlimb: femur (MAP-8290), tibia (MAP-8291), fibula (MAP-8292), as- tragalus(MAP-8293),calcaneum(MAP-8294),metatarsalIV (MAP-8295), phalanx 1 of digit II (MAP-8296), phalanx 3 of digit IV (MAP-8297), ungual phalanx of digit II (MAP-8298), and ungual phalanx of digit IV (MAP-8299)
Reptilia - Stegosauridae
Stegosauria indet. Marsh 1877
3 individuals
"post-cranial remains of stegosaurs"
    = Dacentrurus sp. Lucas 1902
Cobos et al. 2008
CPT-1330, 1435; MAP-4680-4683 (vertebrae)
    = aff. Dacentrurus sp. Lucas 1902
Company et al. 2010
"Cervical and caudal vertebrae, iliosacral block, ischia, pubes, femur, tibia, fibula, calcaneum, astragalus, dermal plate (mainly undescribed)"
    = Dacentrurus armatus Marsh 1877
Cobos et al. 2014
Reptilia - Allosauridae
Theropoda indet. Marsh 1881
4 specimens
CPT-2331, 1326, 1437, 1518
    = Allosauridae indet. Marsh 1878
Gascó et al. 2012
"morphotype B"
Reptilia - Dromaeosauridae
Dromaeosauridae indet. Colbert and Russell 1969
Gascó et al. 2012 5 specimens
"morphotype C"; CPT-2126, 1283, 1327, 1433, 1443
Reptilia
Turiasaurus riodevensis n. gen., n. sp. Royo-Torres et al. 2006
1 individual
CPT-1195–1261
see common names

Geography
Country:Spain State/province:Aragón County:Teruel
Coordinates: 40.1° North, 1.1° West (view map)
Paleocoordinates:31.2° North, 8.1° East
Basis of coordinate:stated in text
Geographic resolution:small collection
Time
Period:Jurassic Epoch:Late/Upper Jurassic
Key time interval:Late/Upper Kimmeridgian - Tithonian
Age range of interval:152.20000 - 145.00000 m.y. ago
Stratigraphy
Formation:Villar del Arzobispo
Stratigraphic resolution:bed
Stratigraphy comments: Age of Villar del Arzobispo Formation constrained to Kimmeridgian-Tithonian by Campos-Soto et al. (2017), rather than Tithonian-Berriasian of older literature
Lithology and environment
Primary lithology:gray claystone
Secondary lithology:quartzose sandstone
Includes fossils?Y
Environment:fine channel fill
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Spatial orientation:random
Preservation of anatomical detail:good
Articulated whole bodies:none
Associated major elements:many
Disassociated major elements:some
Disassociated minor elements:many
Size sorting:poor
Fragmentation:occasional
Feeding/predation traces:tooth marks
Temporal resolution:snapshot
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:taxonomic analysis
Collection dates:2005–2009
Collection method comments: specimens housed in the Museo de la Fundación Conjunto Paleontológico de Teruel-Dinópolis, Teruel
Metadata
Also known as:Riodeva
Database number:68019
Authorizer:J. Alroy, P. Mannion, M. Carrano Enterer:J. Alroy, M. Carrano, J. Tennant
Modifier:M. Carrano Research group:vertebrate
Created:2006-12-23 20:05:49 Last modified:2022-12-08 14:33:36
Access level:the public Released:2006-12-23 20:05:49
Creative Commons license:CC BY
Reference information

Primary reference:
19463. R. Royo-Torres, A. Cobos, and L. Alcalá. 2006. A giant European dinosaur and a new sauropod clade. Science 314:1925-1927 [J. Alroy/J. Alroy/J. Alroy]

Secondary references:

82566 L. Alcalá, L. Luque, A. Aberasturi, D. Ayala, A. Cobos, E. Espílez, F. Gascó, A. González, L. Mampel and R. Royo-Torres. 2010. El dinosaurio gigante de Riodeva: nacido para enseñar [Riodeva’s giant dinosaur: born to teach]. Enseñanza de las Ciencias de la Tierra 18(1):97-106 [M. Carrano/M. Carrano]
79837 S. Campos-Soto, A. Cobos, E. Caus, M. I. Benito, L. Fernández-Labrador, P. Suarez-Gonzalez, I. E. Quijada, R. Mas, R. Royo-Torres and L. Alcalá. 2017. Jurassic Coastal Park: A great diversity of palaeoenvironments for the dinosaurs of the Villar del Arzobispo Formation (Teruel, eastern Spain). Palaeogeography, Palaeoclimatology, Palaeoecology 485:154-177 [P. Mannion/G. Varnham/G. Varnham]
47244 A. Cobos and F. Gascó. 2013. New vertebral remains of the stegosaurian dinosaur Dacentrurus from Riodeva (Teruel, Spain). Geogaceta 53:17-20 [M. Carrano/M. Carrano]
73949 A. Cobos, M. G. Lockley, F. Gascó, R. Royo-Torres, and L. Alcalá. 2014. Megatheropods as apex predators in the typically Jurassic ecosystems of the Villar del Arzobispo Formation (Iberian Range, Spain). Palaeogeography, Palaeoclimatology, Palaeoecology 399:31-41 [M. Carrano/M. Carrano]
30763 A. Cobos, R. Royo-Torres, and L. Alcalá. 2008. Presencia del estegosaurio Dacentrurus en Riodeva (Teruel) [Presence of the stegosaur Dacentrurus in Riodeva (Teruel)]. In J I Ruiz-Omeñaca, L Piñuela and J C García-Ramos (eds), XXIV Jornadas de la Sociedad Española de Paleontología, 15-18 October 2008, Museo del Jurásico de Asturias (MUJA), Colunga, Spain, Libro de Resúmenes 89-90 [M. Carrano/M. Carrano]
83320 A. Cobos, R. Royo-Torres, F. Gascó and L. Alcalá. 2011. A new giant turiasaurian specimen from Riodeva (Teruel, Spain). In A. van der Geer & A. Athanassiou (ed.), European Association of Vertebrate Palaeontologists, 9th Annual Meeting, Program and Abstracts 18 [M. Carrano/M. Carrano]
82563 A. Cobos, R. Royo-Torres, and L. Alcalá. 2014. Nuevos restos fósiles del estegosaurio Dacentrurus en el yacimiento Barrihonda-El Humero de Riodeva (Teruel, España) [New fossil remains of the stegosaur Dacentrurus in the Barrihondo-El Humero locality of Riodeva (Teruel, Spain)]. In R. Royo-Torres, F. J. Verdú, & L. Alcalá (eds.), XXX Jornadas de la Sociedad Española de Paleontología. ¡Fundamental! 24:47-49 [M. Carrano/M. Carrano]
82562 J. Company, X. Pereda Suberbiola, and J. I. Ruiz-Omeñaca. 2010. New stegosaurian (Ornithischia, Thyreophora) remains from Jurassic-Cretaceous transition beds of Valencia province (Southwestern Iberian Range, Spain). Journal of Iberian Geology 36(2):243-252 [M. Carrano/M. Carrano]
82565 G. Delvene, M. Munt, R. Royo-Torres, A. Cobos, and L. Alcalá. 2020. Late Jurassic-Early Cretaceous freshwater bivalves from Turiasaurus riodevensis bearing strata of Teruel (Spain). Spanish Journal of Palaeontology 28(2):161-172 [M. Carrano/M. Carrano]
44510 F. Gascó, A. Cobos, R. Royo-Torres, L. Mampel, and L. Alcalá. 2012. Theropod teeth diversity from the Villar del Arzobispo Formation (Tithonian–Berriasian) at Riodeva (Teruel, Spain). Palaeobiodiversity and Palaeoenvironments [M. Carrano/M. Carrano]
78423 L. Luque, L. Alcalá, L. Mampel, M. D. Pesquero, R. Royo-Torres, A. Cobos, E. Espílez, A. González, D. Ayala, A. Aberasturi, P. Marzo and R. Alloza. 2009. Mineralogical, elemental and chemical composition of dinosaur bones from Teruel (Spain). Journal of Taphonomy 7(2-3):151-178 [M. Carrano/M. Carrano]
76801 F. Ortega, R. Royo Torres, F. Gascó, F. Escaso, and J. L. Sanz. 2010. New evidences of the sauropod Turiasaurus from the Portuguese Upper Jurassic. 8th Meeting of the European Association of Vertebrate Palaeontologists. Abstract Volume 62 [M. Carrano/M. Carrano]
82564 X. Pereda Suberbiola, J. I. Ruiz-Omeñaca, and J. Company. 2009. Primera descripción de restos esqueléticos de dinosaurio ornitópodo en la Formación Villar del Arzobispo (tránsito Jurásico-Cretácico): yacimiento de Cerrito del Olmo (Alpuente, Valencia) [First description of ornithopod dinosaur skeletal remains from the Villar del Arzobispo Formation (Jurassic-Cretaceous transition): Cerrito del Olmo locality (Alpuente, Valencia)]. Geogaceta 47:13-16 [M. Carrano/M. Carrano]
51381 A. Pérez-García, R. Royo-Torres, and A. Cobos. 2015. A new European Late Jurassic pleurosternid (Testudines, Paracryptodira) and a new hypothesis of paracryptodiran phylogeny. Journal of Systematic Palaeontology 13(4):351-369 [P. Mannion/J. Tennant/P. Mannion]
28737ETE R. Royo-Torres, A. Cobos, A. Aberasturi, E. Espílez, I. Fierro, A. González, L. Luque, L. Mampel, and L. Alcalá. 2007. Riodeva sites (Teruel, Spain) shedding light to European sauropod phylogeny. Geogaceta 41:183-186 [M. Carrano/M. Carrano]
42863 R. Royo-Torres and P. Upchurch. 2012. The cranial anatomy of the sauropod Turiasaurus riodevensis and implications for its phylogenetic relationships. Journal of Systematic Palaeontology 10(3):553-583 [M. Carrano/M. Carrano]
81570 S. Sánchez-Fenollosa, F. J. Verdú, and M. Suñer. 2021. Current knowledge of Late Jurassic ornithopod dinosaurs from Europe. Ciências da Terra Procedia 1:54-57 [M. Carrano/M. Carrano/M. Carrano]
85812 S. Sánchez-Fenollosa, F. J. Verdú, and A. Cobos. 2023. The largest ornithopod (Dinosauria: Ornithischia) from the Upper Jurassic of Europe sheds light on the evolutionary history of basal ankylopollexians. Zoological Journal of the Linnean Society [M. Carrano/M. Carrano]