Sainte-Barbe coal mine, Bernissart: Barremian - Early/Lower Aptian, Belgium
collected by Fagès, Sohier, Sonnet, De Pauw 1878-1881

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Lycopodites sp.
Quinet 1969
Matonidium goepperti
Quinet 1969
Laccopteris dunkeri
Quinet 1969
Protorhipis roemeri
Quinet 1969
Ruffordia goepperti
Quinet 1969
Leckenbya valdensis
Quinet 1969
Pinites solmsi
Quinet 1969
Pinites cf. solmsi
Quinet 1969
Sphenopteris goepperti Dunker 1846
Dupont 1892
Enoplea - Rhabditida - Ascarididae
Ascarites gerus n. sp. Poinar and Boucot 2006
Poinar and Boucot 2006 1 specimen
Oregon State University Y2 852 (2), from IRSNB
Ascarites priscus n. gen., n. sp. Poinar and Boucot 2006
Poinar and Boucot 2006 2 specimens
Oregon State University Y2 852 (2), from IRSNB
Insecta - Hemiptera - Tettigarctidae
Hylaeoneura lignei n. gen., n. sp. Lameere and Severin 1897
Lameere and Severin 1897 1 specimen
Trematoda
Digenites proterus n. gen., n. sp. Poinar and Boucot 2006
Poinar and Boucot 2006 1 specimen
Oregon State University Y2 852 (2), from IRSNB
Actinopteri - Palaeonisciformes - Coccolepidae
Coccolepis macropterus n. sp. Traquair 1911
Quinet 1969
Actinopteri - Aspidorhynchidae
Belonostomus sp. Agassiz 1843
du Pauw 1902
Actinopteri - Elopiformes - Elopidae
Anaethalion robustus n. sp.
Quinet 1969
Actinopteri - Pholidophoriformes - Pholidophoridae
Pholidophorus sp. Agassiz 1843
du Pauw 1902
Pholidophorus obesus n. sp.
Quinet 1969
Actinopteri - Leptolepidiformes - Leptolepididae
Leptolepis formosus n. sp.
Quinet 1969
Leptolepis attenuatus n. sp.
Quinet 1969
Leptolepis brevis n. sp.
Quinet 1969
Actinopteri - Ichthyodectiformes
Pachythrissops vectensis (Woodward 1890)
Quinet 1969
Actinopteri - Pycnodontiformes - Pycnodontidae
Mesodon bernissartensis n. sp.
Quinet 1969
replaced by Typodus
Microdon sp. Agassiz 1843
Woodward 1885
    = Microdon radiatus
du Pauw 1902
replaced by Polypsephis
Actinopteri - Amiiformes - Caturidae
Caturus sp. Agassiz 1843
du Pauw 1902
Actinopteri - Amiiformes - Amiidae
Amiopsis dolloi n. sp. Traquair 1911
Quinet 1969
Amiopsis lata n. sp. Traquair 1911
Quinet 1969
synonym of Amiopsis dolloi
Actinopteri - Lepisosteiformes
Lepidotes sp. (Agassiz 1843)
Woodward 1885
    = Lepidotus mantelli Agassiz 1833
du Pauw 1902
recombined as Scheenstia mantelli
Lepidotes bernissartensis n. sp.
Quinet 1969
Lepidotes brevifulcratus n. sp.
Quinet 1969
Lepidotes arcuatus n. sp.
Quinet 1969
Actinopteri - Semionotiformes - Macrosemiidae
Notagogus parvus n. sp.
Quinet 1969
Actinopteri - Ophiopsiidae
Ophiopsis sp. Agassiz 1843
Woodward 1885
    = Ophiopsis penicillatus
du Pauw 1902
Ophiopsis dorsalis Agassiz 1844
Dupont 1892
Actinopteri - Ionoscopiformes - Furidae
Callopterus insignis n. sp.
Quinet 1969
unclassified
Hylaeobatrachus croyii n. gen., n. sp. Dollo 1884
Dollo 1884
Urodela - Urodela
Urodela indet. (Duméril 1805)
Dollo 1883
Reptilia - Testudines - Peltochelyidae
Peltochelys duchastelii n. sp. Dollo 1885
Dollo 1885 1 specimen
Lectotype shell, IRSNB Ct. R. 16; Dollo (1984) also originally referred two juvenile shells from this locality but they were transferred to Chelonii indet. by Perez-Garcia (2011)
Chelonii indet. Latreille 1800
Pérez-García 2011 2 specimens
IRSNB Ct. R. 17 and IRSNB Ct. R. 18 (juveniles originally referred to Peltochelys duchastelii by Dollo (1884))
    = Peltochelys duchastelli Dollo 1885
Karl et al. 2012
original and current combination Peltochelys duchastelii
Reptilia - Testudines
Chitracephalus dumonii n. sp. Dollo 1885
Dollo 1885 1 specimen
IRSNB R11 and IRSNB R12, an almost complete skeleton
Reptilia
Megalosaurus dunkeri Dames 1884
Quinet 1969 1 individual
phalanx
    = Theropoda indet. Marsh 1881
Yans et al. 2006
Reptilia - Iguanodontidae
Iguanodon atherfieldensis Hooley 1925
Yans et al. 2006 1 individual
IRSNB 1551 (12 measurements)
    = Dollodon bampingi n. gen., n. sp. Paul 2008
Paul 2008
synonym of Mantellisaurus atherfieldensis
Reptilia
Iguanodon sp. Mantell 1825
28 individuals
(15 measurements)
    = Iguanodon bernissartensis n. sp. Boulenger 1881
Boulenger 1881
Reptilia - Goniopholididae
Anteophthalmosuchus epikrator Ristevski et al. 2018
Ristevski et al. 2018
IRSNB R47 (previously referred to Anteophthalmosuchus hooleyi by Martin et al. 2016)
Goniopholis simus Owen 1879
Dollo 1883 1 individual
IRSNB R1537 (partial skeleton including skull and mandible) and IRSNB R1539 (postcranial skeleton)
    = Goniopholis sp. Owen 1842
Salisbury et al. 1999
Salisbury et al. (1999) considered Dollo's assignment of this material to Goniopholis simus as incorrect, and suggested it represents a new species of Goniopholis
    = Anteophthalmosuchus hooleyi Salisbury and Naish 2011
Martin et al. 2016
IRSNB R47 and R290
Reptilia - Crocodylia
Crocodylia indet. (Owen 1842)
Norman 1987 2 individuals
Reptilia
Bernissartia fagesii n. gen., n. sp. Dollo 1883
Dollo 1883 1 individual
Two individuals
Entamoebidae
Entamoebites antiquus n. gen., n. sp. Poinar and Boucot 2006
Poinar and Boucot 2006 1 specimen
Oregon State University Y2 852 (2), from IRSNB
Cycadeoideopsida - Cycadeoideales
Conites minuta
Quinet 1969
Equisetopsida - Equisetales - Equisetaceae
Equisetites sp. Sternberg 1833
Quinet 1969
Ginkgoopsida - Pentoxylales - Pentoxylaceae
cf. Taeniopteris sp. Brongniart 1828
Quinet 1969
Ginkgoopsida - Caytoniales - Caytoniaceae
Sagenopteris mantelli
Quinet 1969
Ginkgoopsida - Calamopityales - Calamopityaceae
Adiantites sp. Bell 1960
Quinet 1969
unclassified
Stellatopollis hughesii Penny 1986
Yans et al. 2006
equivalent to Superret-croton
Pteridopsida
Onychiopsis mantelli
Quinet 1969
Cladophlebis dunkeri
Quinet 1969
Pteridopsida - Gleicheniales - Matoniaceae
Weichselia mantelli
Quinet 1969
Pteridopsida - Gleicheniales - Gleicheniaceae
Gleichenia sp. Smith
Woodward 1885
Pteridopsida - Polypodiales
Gleichenites sp. Goeppert 1836
Woodward 1885
unclassified
Pecopteris sp. Brongniart 1822
Woodward 1885
    = Pecopteris polymorpha
Dupont 1892
Pecopteris conybeari
Dupont 1892
Alethopteridae
Alethopteris sp. Sternberg 1825
Woodward 1885
    = Alethopteris elegans
Dupont 1892
unclassified
Lonchopteris sp. Brongniart 1828
Woodward 1885
    = Lonchopteris mantelli
Dupont 1892
"dominant species"
Sphenopteridae
Sphenopteris sp. Sternberg 1825
Woodward 1885
    = Sphenopteris roemeri
Dupont 1892
Sphenopteris fittoni
Quinet 1969
Sphenopteris delicatissima
Quinet 1969
unclassified
Algites sp. Hantzschel 1975
Quinet 1969
see common names

Geography
Country:Belgium State/province:Sauerland
Coordinates: 50.5° North, 3.6° East (view map)
Paleocoordinates:40.7° North, 13.7° East
Basis of coordinate:based on nearby landmark
Altitude:13 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Early/Lower Cretaceous
*International age/stage:Aptian - Albian
Key time interval:Barremian - Early/Lower Aptian
Age range of interval:129.40000 - 122.46000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Geological group:Wealden Formation:Sainte-Barbe Clays
Stratigraphic resolution:group of beds
Stratigraphy comments: 2 layers within mine; age middle Barremian to earliest Aptian
Lithology and environment
Primary lithology:black claystone
Secondary lithology: bituminous coal
Includes fossils?Y
Lithology description: ETE rock type adj: 0, ETE lithology comments: .
Environment:lacustrine - small
Geology comments: ETE sed env 1: lacustrine_sm
Taphonomy
Modes of preservation:body,trace,coprolite
Lagerst�tten type:concentrate
Degree of concentration:-multiple events
Size of fossils:macrofossils,mesofossils,microfossils
Spatial orientation:life position
Preservation of anatomical detail:excellent
Abundance in sediment:common
Articulated whole bodies:many
Associated major elements:many
Fragmentation:occasional
Temporal resolution:time-averaged
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:selective quarrying,chemical,mechanical,field collection
Reason for describing collection:taphonomic analysis
Collectors:Fagès, Sohier, Sonnet, De Pauw Collection dates:1878-1881
Collection method comments: discovered March 1878 by coal miners and engineers, including Mr. Ballez, Mr. Latinis
Taxonomic list comments:coprolites also present, presumably from a carnivore
Metadata
Also known as:ETE Locality 1519, Cran du Midi, Cran du Nord
Database number:22645
Authorizer:A. Behrensmeyer, J. Alroy, P. Mannion, M. Clapham, R. Benson, M. Carrano Enterer:A. Behrensmeyer, J. Tennant, M. Clapham, J. Karr, J. Alroy, M. Carrano, R. Benson, P. Mannion
Modifier:M. Carrano Research group:vertebrate
Created:2001-03-27 01:00:00 Last modified:2019-03-15 15:59:02
Access level:the public Released:2001-03-27 01:00:00
Creative Commons license:CC BY
Reference information

Primary reference:

52560. P.-J. Van Beneden. 1878. Sur la découverte de reptiles fossiles gigantesques dans le charbonnage de Bernissart près de Péruwelz [On the discovery of gigantic fossil reptiles in the Bernissart coal mine near Péruwelz]. Bulletin de l'Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, série 2 45:578-579 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

63616 Anonymous. 1897. Les iguanodons de Bernissart [The Iguanodons of Bernissart]. Le Mouvement Géographique 14(314):3-4 [M. Carrano/M. Carrano]
63615 Anonymous. 1899. La nature et l’age du gisement des iguanodons de Bernissart [The nature and age of the Iguanodon locality of Bernissart]. Le Mouvement Géographique 16(1):2-3 [M. Carrano/M. Carrano]
63593 C.-E.g. Bertrand. 1903. Les coprolithes de Bernissart. Permière partie. Les coprolithes qui ont été attribués aux Iguanodons [The coprolites of Bernissart. First part. The coprolites that have been attributed to Iguanodon]. Mémoires du Musée Royal d’Histoire Naturelle de Belgique 1(4):1-154 [M. Carrano/M. Carrano]
19188ETE G. A. Boulenger. 1881. Sur l'arc pelvien chez les dinosauriens de Bernissart [On the pelvic arch in the dinosaurs of Bernissart]. Bulletins de L'Académie royale de Belgique, 3eme série 1(5):1-11 [M. Carrano/M. Carrano]
52262 E. Buffetaut and G. Wouters. 1978. Le centenaire des iguanodons de Bernissart. La Recherche 9(88):390-392 [M. Carrano/A. Murch]
53309 E. D. Cope. 1880. Iguanodons. The American Naturalist 14:612 [M. Carrano/M. Carrano]
63288 J. Cornet and G. Schmitz. 1899. Note sur les puits naturels du terrain Houiller du Hainaut et le gisement des Iguanodons de Bernissart [Note on the natural wells of the coal field of Hainaut and the Bernissart Iguanodon locality]. Bulletin de la Société Belge de Géologie, de Paléontologie et d’Hydrologie 12:301-318 [M. Carrano/M. Carrano/M. Carrano]
63289 L. F. De Pauw. 1902. Contribution a l’étude de l’Iguanodon bernissartensis [Contribution to the study of iguanodon bernissartensis]. Mémoires et Publications de la Société des Sciences, des Arts et des Lettres du Hainaut 6(4):85-95 [M. Carrano/M. Carrano]
63188 L. Dollo. 1883. Quatrième note sur les dinosauriens de Bernissart [Fourth note on the dinosaurs of Bernissart]. Bulletin du Musée Royal d'Histoire Naturelle de Belgique 2:223-252 [M. Carrano/M. Carrano]
38887 L. Dollo. 1883. Première note sur les crocodiliens de Bernissart [First note on the crocodilians of Bernissart]. Bulletin du Musée Royal d'Histoire Naturelle de Belgique 2:309-338 [M. Carrano/M. Carrano/M. Carrano]
65082 L. Dollo. 1883. Troisième note sur les dinosauriens de Bernissart [Third note on the dinosaurs of Bernissart]. Bulletin du Musée Royal d'Histoire Naturelle de Belgique 2:85-120 [M. Carrano/M. Carrano/M. Carrano]
46929 L. Dollo. 1884. Les découvertes de Bernissart [The discoveries of Bernissart]. Annales des Sciences Géologiques 16(6):1-14 [M. Carrano/M. Carrano/M. Carrano]
55910 L. Dollo. 1885. Les Iguanodons de Bernissart [The Iguanodons of Bernissart]. Revue des Questions Scientifiques 18:5-55 [M. Carrano/M. Carrano]
42005 L. Dollo. 1885. Premiere note sur les cheloniens de Bernissart. Bulletin de la Musee Royale d'Histoire naturelle de Belgique 3:63-79 [R. Benson/R. Benson/M. Carrano]
63189 L. Dollo. 1887. Les ligaments ossifiés des Iguanodons [The ossified ligaments of Iguanodon]. Revue des Questions Scientifiques 21:305-307 [M. Carrano/M. Carrano]
29875 L. Dollo. 1909. The fossil vertebrates of Belgium. Annals of the New York Academy of Science 19(4):99-119 [M. Uhen/M. Uhen]
30984 M. L. Dollo. 1884. Note sur le batracien de Bernissart. Bulletin du Musée royal d'histoire naturelle de Belgique 3(1):85-93 [J. Alroy/J. Alroy/J. Alroy]
51771 L. F. du Pauw. 1902. Notes sur les Fouilles du Charbonnage de Bernissart. Découverte, Solidification et Montage des Iguanodons [Notes on the excavations of the Bernissart coal mine. Discovery, solidification, and mounting of the Iguanodons] 1-25 [M. Carrano/M. Carrano]
63198 É Dupont. 1878. Sur la découverte d’ossements d’Iguanodon, de poissons et de végétaux dans la fosse Sainte-Barbe du charbonnage de Bernissart [On the discovery of bones of Iguanodon, fishes, and plants in the Sainte-Barbe pit of the Bernissart coal mine]. Bulletins de l'Académie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, série 2 46(9-10):387-408 [M. Carrano/M. Carrano]
63200 É Dupont. 1883. Observations sur une nouvelle note de M. P.-J. Van Beneden, concernant la découverte des ossements de Bernissart [Observations on a new note from Mr. P.-J. Van Beneden, concerning the discovery of the bones of Bernissart]. Bulletins de l'Académie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, série 3 6(11):541-549 [M. Carrano/M. Carrano]
63199 É Dupont. 1883. Observations sur une note récente de M. P.-J. Van Beneden, concernant la découverte des ossements de Bernissart [Observations on a recent note by Mr. P.-J. Van Beneden concerning the discovery of the bones of Bernissart]. Bulletins de l'Académie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, série 3 6(8):138-143 [M. Carrano/M. Carrano]
63201 É Dupont. 1892. Le gisement des Iguanodons de Bernissart [The locality of the Iguanodons of Bernissart]. Bulletin de la Société Belge de Géologie, de Paléontologie et d’Hydrologie 6:86-92 [M. Carrano/M. Carrano]
46182 A. Gaudry. 1890. Les Enchainements du Monde Animal dans les Temps Geologiques. Fossiles Secondaires 1-322 [M. Carrano/M. Carrano]
35280ETE P. Godefroit and F. Knoll. 2003. Late Triassic dinosaur teeth from southern Belgium. Comptes Rendus Palevol 2(1):3-11 [M. Carrano/M. Carrano]
58956 J. Gosselet. 1879. Découverte d'ossements d'Iguanodon à Bernissart. Analyse d'une lecture faite par M. Ed. Dupont, à l'Académie de Belgique [Discovery of bones of Iguanodon at Bernissart. Analysis of a lecture made by Mr. Ed. Dupont at the Belgian Academy]. Annales de la Société Géologique du Nord 6:61-64 [M. Carrano/M. Carrano]
56026 W. Gross. 1934. Die Typen des mikroskopischen Knochenbaues bei fossilen Stegocephalen und Reptilien [The types of microscopic bone structure in fossil stegocephalians and reptiles]. Zeitschrift für Anatomie und Entwicklungsgeschichte 103:731-764 [M. Carrano/M. Carrano]
57073 S. F. Harmer. 1896. On the casts of Iguanodon bernissartensis, Boulenger, recently presented to the Museum of Zoology by H.M. the King of the Belgians. Proceedings of the Cambridge Philosophical Society 9(4):202-203 [M. Carrano/M. Carrano]
63047 L. Joleaud. 1922. Les reptiles fossiles [Fossil reptiles]. Association Française pour l'Avancement des Sciences. Conférences. Compte Rendu de la 45e Session 49-66 [M. Carrano/M. Carrano]
62919 J. Joubert. 1908. Le Diplodocus de l’ère secondaire [Diplodocus from the Secondary Era]. Revue de l’Anjou 57(4):267-282 [M. Carrano/M. Carrano]
38095 H.-V. Karl, J. Milan, and G. Tichy. 2012. New shell remains of Peltochelys duchastelli (Dollo, 1884) from Western Europe and the rise of trionychids (Testudines: Trionychoidea). Studia Palaeocheloniologica 4:87-106 [R. Benson/R. Benson/J. Tennant]
46234 A. Lameere and G. Severin. 1897. Les insectes de Bernissart. Annales de la Société Entomologique de Belgique 41:35-38 [M. Clapham/J. Karr/M. Clapham]
60585 J. E. Martin, M. Delfino, and T. Smith. 2016. Osteology and affinities of Dollo's goniopholidid (Mesoeucrocodylia) from the Early Cretaceous of Bernissart, Belgium. Journal of Vertebrate Paleontology 36(6):e1222534:1-27 [P. Mannion/P. Mannion/P. Mannion]
55547 H. N. Moseley. 1883. The Bernissart Iguanodon. Nature 28(723):439-443 [M. Carrano/M. Carrano]
62807 D. B. Norman. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the Lower Cretaceous of Bernissart (Belgium). Institut Royal des Sciences Naturelles de Belgique Mémoire 178:1-103 [M. Carrano/M. Carrano/M. Carrano]
5691ETE 1447D. B. Norman. 1987. A mass accumulation of vertebrates from the Lower Cretaceous of Nehden (Sauerland), West Germany. Proceedings of the Royal Society, London B 230:215-255 [A. Behrensmeyer/A. Behrensmeyer/M. Kosnik]
32089 G. S. Paul. 2008. A revised taxonomy of the iguanodont dinosaur genera and species. Cretaceous Research 29(2):192-216 [R. Butler/R. Butler/M. Carrano]
42022 A. Pérez-García. 2011. Revisión de la serie tipo de Peltochelys duchastelli (Chelonii, Trionychoidea) del Cretácio Inferior de Bernissart (Bélgica) [Review of the type series of Peltochelys duchastelii (Chelonii, Trionychoidea) from the Lower Cretaceous of Bernissart (Belgium)]. In A. Pérez-García, F. Gascó, J. M. Gasulla, F. Escaso (eds.), Viajando a Mundos Pretéritos 249-260 [R. Benson/R. Benson]
42442 A. Pérez-García. 2012. The European Lower Cretaceous Chitracephalus dumonii (Testudines: Cryptodira) and the diversity of a poorly known lineage of turtles. Acta Palaeontologica Polonica 57(3):575-588 [R. Benson/R. Benson/P. Mannion]
55013 G. Poinar and A. J. Boucot. 2006. Evidence of intestinal parasites of dinosaurs. Parasitology 133:245-249 [M. Clapham/M. Clapham]
55637 G.-E. Quinet. 1969. Bernissart...il y a 125.000.000 d'Années. Le Règne des Iguanodons [Bernissart...125,000,000 Years Ago. The Reign of the Iguanodon] 1-86 [M. Carrano/M. Carrano]
65779 J. Ristevski, M. T. Young, M. B. Andrade and A. K. Hastings. 2018. A new species of Anteophthalmosuchus (Crocodylomorpha, Goniopholididae) from the Lower Cretaceous of the Isle of Wight, United Kingdom, and a review of the genus. Cretaceous Research 84:340-383 [P. Mannion/P. Mannion]
39157 S. W. Salisbury, P. M. A. Willis, S. Peitz and P. M. Sander. 1999. The crocodilian Goniopholis simus from the Lower Cretaceous of north-western Germany. Special Papers in Palaeontology 60:121-148 [P. Mannion/P. Mannion]
64081 A. L. L. Seitz. 1907. Vergleichende Studien über den mikroskopischen Knochenbau fossiler und rezenter Reptilien und dessen Bedeutung für das Wachstum und Umbildung des Knochengewebes im allgemeinen [Comparative studies on the microscopic bone structure of fossil and recent reptiles and its importance for the growth and remodeling of bone tissue in general]. Nova Acta. Abhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Deutschen Akademie der Naturforscher 87(2):228-370 [M. Carrano/M. Carrano]
65100 W. E. Swinton. 1934. A Guide to the Fossil Birds, Reptiles, and Amphibians in the Department of Geology and Paleontology in the British Museum (Natural History), Cromwell Road, London, S.W. 7 xii-87 [M. Carrano/M. Carrano]
28930ETE R. A. Thulborn. 1991. Morphology, preservation and palaeobiological significance of dinosaur coprolites. Palaeogeography, Palaeoclimatology, Palaeoecology 83:341-366 [M. Carrano/M. Carrano]
52561 P.-J. Van Beneden. 1881. Sur l'arc pelvien chez les Dinosauriens de Bernissart [On the pelvic arch in the dinosaurs of Bernissart]. Bulletin de l'Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, série 3 1:600-608 [M. Carrano/M. Carrano]
52558 P.-J. Van Beneden. 1883. Seconde communication sur la découverte de l'Iguanodon de Bernissart [Second communication on the discovery of the Iguanodon of Bernissart]. Bulletin de l'Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, série 3 6:219-226 [M. Carrano/M. Carrano]
52559 P.-J. Van Beneden. 1883. Sur ce qu'il faut entendre par le mot DÉCOUVERTE, à propos des Iguanodons de Bernissart [On what is meant by the word DISCOVERY, regarding the Iguanodons of Bernissart]. Bulletin de l'Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, série 3 6:25-27 [M. Carrano/M. Carrano]
68171 E. Van Den Broeck. 1900. Les dépôts à Iguanodons de Bernissart et leur transfert dans l’étage Purbeckien ou Aquilonien du Jurassique supérieur. Exposé comprenant une revue de la faune des vertébrés du Purbeckien et du Wealdien dans le sud-est de l’Angleterre [The Iguanodon deposits of Bernissart and their transfer to the Purbeckian or Aquilonian stage of the Upper Jurassic. Presentation including a review of the vertebrate fauna of the Purbeckian and Wealden of southeast England ]. Mémoires de la Société Belge de Géologie de Paléontologie et d’Hydrologie 14:39-112 [M. Carrano/M. Carrano]
36225ETE E. Van den Broeck. 1902. Nouvelles observations relatives au gisement des Iguanodons de Bernissart (deuxième partie). Étude par sondage des dépôts de recouvrement de la région des puits naturels de Bernissart. Le Cran du Nord et le sondage horizontal de 1864, au Cran du Midi [New observations regarding the Iguanodon locality of Bernissart (second part). Survey study of the covering deposits from the region of the natural wells of Bernissart. The North Notch and the horizontal survey of 1864, at the South Notch]. Procès-Verbaux de la Sociéte Belge de Géologie de Paléontologie et d'Hydrologie, deuxième série 13:175-181 [M. Carrano/M. Carrano]
36224ETE E. Van den Broeck. 1902. Nouvelles observations relatives au gisement des Iguanodons de Bernissart (première partie). Étude critique sur les coupes et figures du gisement de Bernissart dressées et publiées par M. Éd. Dupont [New observations regarding the Iguanodon locality of Bernissart (first part). Critical study of the sections and figures of the Bernissart locality drawn up and published by Mr. Éd. Dupont]. Procès-Verbaux de la Sociéte Belge de Géologie de Paléontologie et d'Hydrologie, deuxième série 13:6-11 [M. Carrano/M. Carrano/M. Carrano]
61518 D. B. Weishampel and J. B. Weishampel. 1983. Annotated localities of ornithopod dinosaurs: implications to Mesozoic paleobiogeography. The Mosasaur 1:43-87 [M. Carrano/M. Carrano]
17192ETE H. Woodward. 1885. On Iguanodon mantelli, Meyer. Geological Magazine, decade 3 2(1):10-15 [M. Carrano/M. Carrano]
17190ETE H. Woodward. 1895. Note on the reconstruction of Iguanodon in the British Museum (Natural History), Cromwell Road. Geological Magazine, decade 4 2(8):289-292 [M. Carrano/M. Carrano]
18657ETE J. Yans, J. Dejax, D. Pons, L. Taverne, and P. Bultynck. 2006. The iguanodons of Bernissart (Belgium) are middle Barremian to earliest Aptian in age. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Science de la Terre 76:91-95 [M. Carrano/M. Carrano]