Shabarakh Usu (AMNH loc. 12049): Late/Upper Campanian, Mongolia
collected by Shackelford, Granger, Andrews 1922–1923, 1925

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Zalambdalestidae
Zalambdalestes lechei n. gen., n. sp. Gregory and Simpson 1926
Gregory and Simpson 1926 3 individuals
AMNH 21708 (holotype), 21707, 21704 (paratypes) partial skulls
Zalambdalestes grangeri n. sp. Simpson 1928
Simpson 1928 1 individual
AMNH 21709
Mammalia - Deltatheroida - Deltatheridiidae
Deltatheridium pretrituberculare n. gen., n. sp. Gregory and Simpson 1926
Gregory and Simpson 1926 1 individual
AMNH 21705
Hydotherium dobsoni n. gen., n. sp. Gregory and Simpson 1926
Gregory and Simpson 1926 1 individual
AMNH 21702
Mammalia - Deltatheroida
Deltatheroides cretacicus n. gen., n. sp. Gregory and Simpson 1926
Gregory and Simpson 1926 1 individual
AMNH 21700
Mammalia - Multituberculata
Multituberculata indet. (Cope 1884)
Andrews 1932 1 specimen
Djadochtatherium matthewi Simpson 1925
Gregory and Simpson 1926 1 individual
AMNH 21703 (right maxilla)
    = cf. Nemegtbaatar sp. Kielan-Jaworowska 1974
Rougier et al. 1997
Mammalia - Multituberculata - Djadochtatheriidae
Djadochtatherium matthewi n. gen., n. sp. Simpson 1925
Simpson 1925 1 individual
AMNH 20440
Reptilia - Testudines - Nanhsiungchelyidae
Testudines indet. Batsch 1788
Andrews 1932 1 specimen
AMNH 6658, 6659
    = Dermatemydidae indet. Baur 1888
Gilmore 1931
    = Nanhsiungchelyidae indet. Yeh 1966
Danilov and Syromyatnikova 2008
Reptilia - Varanidae
Telmasaurus grangeri n. gen., n. sp. Gilmore 1943
Gilmore 1943 2 individuals
AMNH 6645 (holotype), posterior dorsal part of skull, three dorsal vertebrae, parts of others, rib and other bone fragments; AMNH 6643, articulated hind limbs, incomplete feet and pelvis, sacrum, a few dorsal and caudal vertebrae, portions of one forelimb and foot
Reptilia - Priscagamidae
Mimeosaurus crassus n. gen., n. sp. Gilmore 1943
Gilmore 1943 1 individual
AMNH 6655 (holotype), left maxillary with dentition
Reptilia
Isodontosaurus gracilis n. gen., n. sp. Gilmore 1943
Gilmore 1943 1 individual
AMNH 6647 (holotype), posterior portions of both rami with teeth
Reptilia - Teiidae
Conicodontosaurus djadochtaensis n. gen., n. sp. Gilmore 1943
Andrews 1932 1 individual
AMNH 6519 (holotype), fragmentary skull, parts of both lower jaws and teeth
Reptilia - Polyglyphanodontidae
Macrocephalosaurus ferrugenous n. gen., n. sp. Gilmore 1943
Andrews 1932 1 individual
recombined as Gilmoreteius ferrugenous
AMNH 6520 (holotype), incomplete skull and part of the right ramus
Reptilia
Lepidosauria indet. (Haeckel 1866)
Gilmore 1943 3 specimens
"Sauria undetermined": AMNH 6648, small portions of both lower jaws, parts of maxillaries; AMNH 6521, portions of integument of tail assoc. with 2 caudals; AMNH 6656, articulated right ramus and maxillary with pleurodont dentition unlike other lizards described in the paper
Reptilia - Oviraptoridae
Fenestrosaurus philoceratops n. gen., n. sp. Osborn 1924
Osborn 1924 2 individuals
AMNH 6517, AMNH FARB 33092 (baby)
    = Oviraptor philoceratops n. gen., n. sp. Osborn 1924
Osborn 1924
Reptilia - Troodontidae
Ornithoides oshiensis n. gen., n. sp. Osborn 1924
Osborn 1924 1 individual
AMNH 6516
    = Saurornithoides mongoliensis n. gen., n. sp. Osborn 1924
Osborn 1924
Reptilia - Dromaeosauridae
Ovoraptor djadochtari n. gen., n. sp. Osborn 1924
Osborn 1924 1 individual
AMNH 6515
    = Velociraptor mongoliensis n. gen., n. sp. Osborn 1924
Osborn 1924
Reptilia - Elongatoolithidae
Protoceratops andrewsi Granger and Gregory 1923
Andrews 1932
eggs: AMNH 6508
    = Oviraptor sp. Osborn 1924
Dashzeveg et al. 2005
    = cf. Elongatoolithus sp. Zhao 1975
Simon et al. 2019
Reptilia - Ankylosauridae
Pinacosaurus grangeri n. gen., n. sp. Gilmore 1933
Gilmore 1933 1 individual
AMNH 6523
Reptilia - Protoceratopsidae
Protoceratops sp. Granger and Gregory 1923
Andrews 1932
Protoceratops andrewsi n. gen., n. sp. Granger and Gregory 1923
80 specimens
AMNH 6251
Reptilia - Saurischia
Saurischia indet. Seeley 1888
Andrews 1932 2 specimens
Reptilia - Paralligatoridae
Shamosuchus djadochtaensis n. gen., n. sp. Mook 1924
Andrews 1932 1 individual
AMNH FR 6412
see common names

Geography
Country:Mongolia State/province:Omnogov
Coordinates: 44.0° North, 104.0° East (view map)
Paleocoordinates:41.7° North, 93.0° East
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
Key time interval:Late/Upper Campanian
Age range of interval:83.60000 - 72.10000 m.y. ago
Stratigraphy
Formation:Djadokhta Member:Bayn Dzak
Stratigraphic resolution:group of beds
Lithology and environment
Primary lithology:red sandstone
Includes fossils?Y
Environment:eolian indet.
Taphonomy
Modes of preservation:body
Degree of concentration:dispersed
Size of fossils:macrofossils
Spatial orientation:life position
Preservation of anatomical detail:excellent
Articulated whole bodies:many
Associated major elements:some
Disassociated major elements:some
Fragmentation:occasional
Spatial resolution:autochthonous
Collection methods and comments
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:AMNH
Collectors:Shackelford, Granger, Andrews Collection dates:1922–1923, 1925
Collection method comments: Central Asiatic Expedition
Metadata
Also known as:Protoceratops type, Flaming Cliffs, Bayn Dzak, Kwei-wa-ting trail, Bayan Zag
Database number:46152
Authorizer:M. Carrano, P. Barrett, P. Mannion Enterer:M. Carrano, P. Mannion, T. Cleary
Modifier:M. Carrano Research group:vertebrate
Created:2005-01-05 13:26:18 Last modified:2023-10-12 15:02:12
Access level:the public Released:2005-01-05 13:26:18
Creative Commons license:CC BY
Reference information

Primary reference:
14566.ETE W. Granger and W. K. Gregory. 1923. Protoceratops andrewsi, a pre-ceratopsian dinosaur from Mongolia, with an appendix on the structural relationships of the Protoceratops beds. American Museum Novitates 72:1-9 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

38112 V. R. Alifanov. 2000. The fossil record of Cretaceous lizards from Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 368-389 [R. Benson/R. Benson/M. Carrano]
63491 R. C. Andrews. 1923. Where the dinosaur hid its eggs. The Illustrated London News 4417:1097, 1106-1111 [M. Carrano/M. Carrano]
13060ETE R. C. Andrews. 1932. The New Conquest of Central Asia: A Narrative of the Explorations of the Central Asiatic Expeditions ni Mongolia and China, 1921-1930. Natural History of Central Asia 1:1-678 [M. Carrano/M. Carrano/M. Carrano]
54198 Anonymous. 1923. A forerunner of the horned dinosaurs. Natural History 23:192 [M. Carrano/M. Carrano/M. Carrano]
64040 V. M. Arbour and P. J. Currie. 2016. Systematics, phylogeny and palaeobiogeography of the ankylosaurid dinosaurs. Journal of Systematic Palaeontology 14(5):385-444 [G. Lloyd/G. Lloyd/M. Carrano]
15004ETE R. Barsbold. 1983. Khishchnye dinosavry mela Mongoliy [Carnivorous dinosaurs from the Cretaceous of Mongolia]. Transactions of the Joint Soviet-Mongolian Paleontological Expedition 19:1-117 [M. Carrano/M. Carrano]
69232 E. H. Colbert. 2000. Asiatic dinosaur rush. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 211-234 [M. Carrano/M. Carrano]
18347ETE P. J. Currie. 2000. Theropods from the Cretaceous of Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 434-455 [M. Carrano/M. Carrano/M. Carrano]
41748 I. G. Danilov and E. V. Syromyatnikova. 2008. New materials on turtles of the family Nanhsiungchelyidae from the Cretaceous of Uzbekistan and Mongolia, with a review of the Nanhsiungchelyid record in Asia. Proceedings of the Zoological Institute RAS 312(1/2):3-25 [R. Benson/R. Benson/P. Mannion]
15459ETE D. Dashzeveg, L. Dingus, D.B. Loope, C.C. Swisher, III, T. Dulam and M.R. Sweeney. 2005. New stratigraphic subdivision, depositional environment, and age estimate for the Upper Cretaceous Djadokhta Formation, southern Ulan Nur Basin, Mongolia. American Museum Novitates (3498)1-31 [M. Carrano/K. Maguire/G. Varnham]
34192ETE C. W. Gilmore. 1931. Fossil turtles of Mongolia. Bulletin of the American Museum of Natural History 59:213-257 [M. Carrano/M. Carrano]
12056ETE C. W. Gilmore. 1933. Two new dinosaurian reptiles from Mongolia with notes on some fragmentary specimens. American Museum Novitates 679:1-20 [M. Carrano/M. Carrano/M. Carrano]
12062ETE C. W. Gilmore. 1943. Fossil lizards of Mongolia. Bulletin of the American Museum of Natural History 81(4):361-384 [M. Carrano/M. Carrano/R. Benson]
56333 W. Granger. 1936. The story of the dinosaur eggs. Natural History 88:21-25 [M. Carrano/M. Carrano]
15701ETE W. K. Gregory and G. G. Simpson. 1926. Cretaceous mammal skulls from Mongolia. American Museum Novitates 225:1-20 [M. Carrano/M. Carrano]
63572 Z. Kielan-Jaworowska. 1984. A caccia di dinosauri nel deserto di Gobi [Hunting dinosaurs in the Gobi Desert]. In G. M. Ronzoni, M. L. Greggio, & T. B. Guarinoni (eds.), Sulle Orme dei Dinosauri 87-109 [M. Carrano/M. Carrano]
12314ETE R. S. Lull. 1933. A revision of the Ceratopsia or horned dinosaurs. Memoirs of the Peabody Museum of Natural History 3(3):1-175 [M. Carrano/M. Carrano/M. Carrano]
86065 M. A. Norell, A. M. Balanoff, D. E. Barta and G. M. Erickson. 2018. A second specimen of Citipati osmolskae associated with a nest of eggs from Ukhaa Tolgod, Omnogov Aimag, Mongolia. American Museum Novitates 3899:1-44 [M. Carrano/M. Carrano]
13096 H. F. Osborn. 1924. Three new Theropoda, Protoceratops zone, central Mongolia. American Museum Novitates 144:1-12 [M. Carrano/M. Carrano/M. Carrano]
13076ETE H. F. Osborn. 1924. The discovery of an unknown continent. Natural History 24(2):133-149 [M. Carrano/M. Carrano/M. Carrano]
62839 H. F. Osborn. 1930. Ancient vertebrate life of central Asia: discoveries of the central Asiatic expeditions of the Museum of Natural History in the years 1921–1929. Centenaire de la Société géologique de France. Livre jubilaire 1830-1930 2:519-543 [M. Carrano/M. Carrano]
2986 G. W. Rougier, M. J. Novacek, and D. Dashzeveg. 1997. A new multituberculate from the Late Cretaceous locality Ukhaa Tolgod, Mongolia. Considerations on multituberculate interrelationships. American Museum Novitates 3191:1-26 [J. Alroy/J. Alroy/R. Benson]
62429 L. Schwarz, F. Fehse, G. Müller, F. Andersson, and F. Sieck. 1961. Untersuchungen an Dinosaurier-Eischalen von Aix en Provence und der Mongolei (Shabarakh Usu) [Studies on dinosaur eggshells from Aix en Provence and Mongolia (Shabarakh Usu)]. Zeitschrift für Wissenschaftliche Zoologie Abteilung A 165(3-4):344-379 [M. Carrano/M. Carrano/M. Carrano]
70244 D. J. Simon, D. J. Varricchio, Z. Jin and S. F. Robison. 2019. Microstructural overlap of Macroelongatoolithus eggs from Asia and North America expands the occurrence of colossal oviraptorosaurs. Journal of Vertebrate Paleontology 38(6):e1553046:1-16 [M. Carrano/M. Carrano/M. Carrano]
38232 G. G. Simpson. 1925. A Mesozoic mammal skull from Mongolia. American Museum Novitates 201:1-12 [R. Benson/R. Benson]
38230 G. G. Simpson. 1928. Further notes on Mongolian Cretaceous mammals. American Museum Novitates 329:1-14 [R. Benson/R. Benson]
10460 A. Sulimski. 1975. Macrocephalosauridae and Polyglyphanodontidae (Sauria) from the Late Cretaceous of Mongolia. Palaeontologia Polonica 33:25-102 [M. Carrano/M. Carrano/M. Carrano]
18351ETE T. A. Tumanova. 2000. Armoured dinosaurs from the Cretaceous of Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 517-532 [M. Carrano/M. Carrano/M. Carrano]
59215 V. Van Straelen. 1928. Les oeufs de reptiles fossiles [The eggs of fossil reptiles]. Palaeobiologica 1:295-312 [M. Carrano/M. Carrano]