PBDB Collection

San Just amber: Late/Upper Albian, Spain

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Arachnida - Araneae - Oonopidae

	Orchestina informal sp. 1 Smith 1882	Saupe et al. 2012	1 specimen
	CPT-4100
Arachnida - Araneae - Lagonomegopidae

	Spinomegops aragonensis n. sp. Pérez-de la Fuente et al. 2013	Pérez-de la Fuente et al. 2013	1 specimen
	CPT-4155
Arachnida - Araneae

	Araneoidea indet. Simon 1895
	spider silk
Arachnida - Acariformes - Trhypochthoniidae

	Trhypochthonius lopezvallei n. sp. Arillo et al. 2012	Arillo et al. 2012	1 specimen
	CPT-4161
Arachnida - Acariformes - Otocepheidae

	Cretaceobodes martinezae n. gen., n. sp. Arillo et al. 2010	Arillo et al. 2010	1 specimen
	CPT-SJ07-05
Arachnida - Acariformes - Cymbaeremaeidae

	Ametroproctus valeriae n. sp. Arillo et al. 2009	Arillo et al. 2009
Arachnida - Acariformes - Erythraeidae

	Leptus sp. Latreille 1796	Arillo et al. 2018	1 specimen
	CPT-4167
Insecta - Diptera - Limoniidae

	Helius turolensis n. sp. Kania‑Kłosok et al. 2021	Kania‑Kłosok et al. 2021	1 specimen
	CPT SJ-10-54 (2 measurements)
Insecta - Diptera - Ceratopogonidae

	Leptoconops zherikhini Szadziewski and Arillo 2003	Arillo et al. 2008	1 specimen
	recombined as Leptoconops (Leptoconops) zherikhini
	CPT-3345

	Protoculicoides skalskii Szadziewski and Arillo 1998	Arillo et al. 2008	1 specimen
	recombined as Gerontodacus skalskii
	CPT-970

	Protoculicoides hispanicus n. sp. Szadziewski and Arillo 2016	Szadziewski et al. 2016	8 specimens
	recombined as Atriculicoides hispanicus
	CPT NBSJ 09/12 A c, 09/12 A b (and others not in type series) (3 measurements)

	Protoculicoides sanjusti n. sp. Szadziewski and Arillo 2016	Szadziewski et al. 2016	2 specimens
	recombined as Atriculicoides sanjusti
	CPT NBSJ 09/12 A a, 09/12 A d (2 measurements)
Insecta - Diptera - Archizelmiridae

	Burmazelmira grimaldii n. sp. Arillo et al. 2018	Arillo et al. 2018	2 specimens
	CPT-3342, 4166
Insecta - Diptera - Dolichopodidae

	Microphorites sp. Hennig 1971		1 specimen
	(2 measurements)
	= Microphorites utrillensis n. sp. Arillo et al. 2008	Arillo et al. 2008
	CPT-963
Insecta - Diptera - Rhagionidae

	Litoleptis fossilis n. sp. Arillo et al. 2009	Arillo et al. 2009	1 specimen
	CPT-3344 (2 measurements)
Insecta - Thysanoptera - Stenurothripidae

	Hispanothrips utrillensis n. gen., n. sp. Peñalver and Nel 2010	Peñalver and Nel 2010	3 specimens
	CPT SJ-07-47, 48, 49
Insecta - Hemiptera

	Iberofoveopsis miguelesi n. gen., n. sp. Peñalver and Szwedo 2010	Peñalver and Szwedo 2010	1 specimen
	CPT-4132
Insecta - Psocodea - Archaeatropidae

	Archaeatropidae indet. Baz and Ortuño 2000	Álvarez-Parra et al. 2022	1 specimen
	SJNB2012-12-02

	Archaeatropos alavensis Baz and Ortuño 2000	Álvarez-Parra et al. 2022	2 specimens
	MAP-7812; SJ-10-50 SJ2
Insecta - Psocodea - Empheriidae

	Preempheria antiqua Baz and Ortuño 2001	Álvarez-Parra et al. 2022	1 specimen
	CPT-4117
Insecta - Mantodea

	Aragonimantis aenigma n. gen., n. sp. Delclòs et al. 2016	Delclòs et al. 2016	1 specimen
	CPT SJ-10-17
Insecta - Blattodea

	Aragonitermes teruelensis n. gen., n. sp. Engel and Delclòs 2010	Engel and Delclòs 2010	1 specimen
	CPT 4065 (1 measurement)
Insecta - Hymenoptera - Spathiopterygidae

	Diameneura marveni n. gen., n. sp. Santer and Álvarez-Parra 2021	Santer et al. 2021	1 specimen
	CPT-4095 (SJ-07-41) (1 measurement)

	Mymaropsis turolensis n. gen., n. sp. Engel and Ortega-Blanco 2013	Engel et al. 2013	2 specimens
	CPT 4077; CPT-4097 (SJ-07-43) in Santer et al., 2021 (4 measurements)
Insecta - Hymenoptera - Serphitidae

	Serphites silban n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	CPT-4059
Insecta - Hymenoptera - Gallorommatidae

	Galloromma turolensis n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	recombined as Cretaceomma turolensis
	CPT 4139 (1 measurement)
Insecta - Hymenoptera - Alavarommatidae

	Alavaromma n. gen. orchamum Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	CPT 4140
Insecta - Hymenoptera - Braconidae

	Utrillabracon electropteron n. gen., n. sp. Álvarez-Parra and Engel 2022	Álvarez-Parra et al. 2022	1 specimen
	MAP-7819 (SJE2012 49-04) (1 measurement)
Insecta - Hymenoptera - Evaniidae

	Cretevania montoyai n. sp. Peñalver et al. 2010	Peñalver et al. 2010	1 specimen
	CPT-957

	Cretevania sp. Rasnitsyn 1975		1 specimen
	= Cretevania alcalai n. sp. Peñalver et al. 2010	Peñalver et al. 2010
	CPT-960

	Cretevania rubusensis n. sp. Peñalver et al. 2010	Peñalver et al. 2010	1 specimen
	CPT-3335
Insecta - Hymenoptera - Stigmaphronidae

	Burmaphron cf. jentilak Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	CPT-4075
Insecta - Neuroptera

	Neuroptera indet. Linnaeus 1758	Pérez-de la Fuente et al. 2020	1 specimen
	SJNB 2012-04 ("Mantispoid or dilaroid larva")
Insecta - Neuroptera - Berothidae

	? Berothidae indet. Handlirsch 1906	Pérez-de la Fuente et al. 2020	1 specimen
	CPT SJ-10-25
Insecta - Neuroptera - Mantispidae

	Aragomantispa lacerata n. gen., n. sp. Pérez-de la Fuente and Peñalver 2019	Pérez-de la Fuente and Peñalver 2019	1 specimen
	CPT SJ-10-22 (1 measurement)
Insecta - Coleoptera - Ptinidae

	Actenobius magneoculus n. sp. Peris et al. 2015	Peris et al. 2015	1 specimen
	CPT SJ-10-18 (2 measurements)
Insecta - Coleoptera - Salpingidae

	Arra legalovi n. gen., n. sp. Peris et al. 2014	Peris et al. 2014	1 specimen
	CPT-4106 (2 measurements)
Insecta - Coleoptera - Cucujidae

	Cucujidae indet. (Latreille 1802)
	Cucujipes
see common names

Geography

Country:	Spain	State/province:	Aragón
Coordinates:	40.8° North, 0.8° West (view map)
Paleocoordinates:	28.2° North, 2.4° East
Basis of coordinate:	based on nearby landmark
Geographic resolution:	outcrop

Time

Period:	Cretaceous	Epoch:	Early/Lower Cretaceous
Stage:	Albian	10 m.y. bin:	Cretaceous 4
Key time interval:	Late/Upper Albian
Age range of interval:	106.20000 - 100.50000 m.y. ago

Stratigraphy

Formation:	Escucha	Member:	La Orden
Stratigraphic resolution:	member
Stratigraphy comments: Upper Albian (E. Barron, pers. com. 2015; based on an accurate palynological study

Lithology and environment

Primary lithology:	amber
Environment:	terrestrial indet.

Taphonomy

Modes of preservation:	body,soft parts,amber
Size of fossils:	mesofossils,microfossils

Collection methods and comments

Collection methods:	selective quarrying
Reason for describing collection:	taxonomic analysis
Collection method comments: Repository: Fundación Conjunto Paleontológico de Teruel-Dinópolis (CPT)

Metadata

Database number:	91626
Authorizer:	J. Alroy, M. Clapham	Enterer:	J. Alroy, M. Clapham
Modifier:	M. Clapham	Research group:	paleoentomology
Created:	2009-10-07 21:13:22	Last modified:	2018-02-24 17:56:03
Access level:	the public	Released:	2009-10-07 21:13:22
Creative Commons license:	CC BY

Reference information

Primary reference:

18002.

E. Peñalver, D. A. Grimaldi, and X. Delclos. 2006. Early Cretaceous spider web with its prey. Science 312:1761 [M. Uhen/M. Uhen/J. Alroy]

Secondary references:

81784		S. Álvarez-Parra, E. Peñalver, A. Nel and X. Delclòs. 2022. New barklice (Psocodea, Trogiomorpha) from Lower Cretaceous Spanish amber. Papers in Palaeontology 8(e1436) [M. Clapham/M. Clapham]
81786		S. Álvarez-Parra, E. Peñalver, X. Delclòs and M. S. Engel. 2022. A braconid wasp (Hymenoptera, Braconidae) from the Lower Cretaceous amber of San Just, eastern Iberian Peninsula. ZooKeys 1103:65-78 [M. Clapham/M. Clapham]
64834		A. Arillo, V. Blagoderov, and E. Peñalver. 2018. Early Cretaceous parasitism in amber: A new species of Burmazelmira fly (Diptera: Archizelmiridae) parasitized by a Leptus sp. mite (Acari, Erythraeidae). Cretaceous Research 86:24-32 [M. Clapham/M. Clapham]
48312		A. Arillo, E. Peñalver, and X. Delclòs. 2008. Microphorites (Diptera: Dolichopodidae) from the Lower Cretaceous amber of San Just (Spain), and the co-occurrence of two ceratopogonid species in Spanish amber deposits. Zootaxa 1920:29-40 [M. Clapham/M. Clapham]
47999		A. Arillo, E. Peñalver, and V. García-Gimeno. 2009. First fossil Litoleptis (Diptera: Spaniidae) from the Lower Cretaceous amber of San Just (Teruel Province, Spain). Zootaxa 2026:33-39 [M. Clapham/M. Clapham]
30884		A. Arillo, L. S. Subías, and U. Shtanchaeva. 2009. A new fossil species of oribatid mite, Ametroproctus valeriae sp. nov. (Acariformes, Oribatida, Ametroproctidae), from the Lower Cretaceous amber of San Just, Teruel Province, Spain. Cretaceous Research 30(2):322-324 [J. Alroy/J. Alroy]
53410		A. Arillo, L. S. Subías, and U. Shtanchaeva. 2010. A new genus and species of oribatid mite, Cretaceobodes martinezae gen. et sp. nov., from the Lower Cretaceous amber of San Just (Teruel Province, Spain) (Acariformes, Oribatida, Otocepheidae). Paleontological Journal 44:287-290 [M. Clapham/M. Clapham]
53409		A. Arillo, L. S. Subías, and U. Shtanchaeva. 2012. A new species of fossil oribatid mite (Acariformes, Oribatida, Trhypochthoniidae) from the Lower Cretaceous amber of San Just (Teruel Province, Spain). Systematic & Applied Acarology 17:106-112 [M. Clapham/M. Clapham/M. Clapham]
57448		X. Delclòs, E. Peñalver, A. Arillo, M. S. Engel, A. Nel, D. Azar, and A. Ross. 2016. New mantises (Insecta: Mantodea) in Cretaceous ambers from Lebanon, Spain, and Myanmar. Cretaceous Research 60:91-108 [M. Clapham/M. Clapham/M. Clapham]
39358		M. S. Engel and X. Delclòs. 2010. Primitive termites in Cretaceous amber from Spain and Canada (Isoptera). Journal of the Kansas Entomological Society 83:111-128 [M. Clapham/M. Clapham]
47622		M. S. Engel, J. Ortega-Blanco, C. Soriano, D. A. Grimaldi, and X. Delclòs. 2013. A new lineage of enigmatic diaprioid wasps in Cretaceous amber (Hymenoptera: Diaprioidea). American Museum Novitates 3771:1-23 [M. Clapham/M. Clapham]
77190		I. Kania‑Kłosok, W. Krzemiński, and A. Arillo. 2021. Two new long‑rostrum cranefly species from the Cretaceous Iberian amber (Diptera, Limoniidae, Helius). Scientific Reports 11(12851) [M. Clapham/M. Clapham]
39574		J. Ortega-Blanco, X. Delclòs, and M. S. Engel. 2011. Diverse stigmaphronid wasps in Early Cretaceous amber from Spain (Hymenoptera: Ceraphronoidea: Stigmaphronidae). Cretaceous Research 32:762-773 [M. Clapham/M. Clapham/M. Clapham]
39579		J. Ortega-Blanco, X. Delclòs, E. Peñalver and M. S. Engel. 2011. Serphitid wasps in Early Cretaceous amber from Spain (Hymenoptera: Serphitidae). Cretaceous Research 32:143-154 [M. Clapham/M. Clapham]
36176		J. Ortega-Blanco, E. Peñalver, X. Declòs and M. S. Engel. 2011. False fairy wasps in Early Cretaceous amber from Spain (Hymenoptera: Mymarommatoidea). Palaeontology 54(3):511-523 [M. Clapham/M. Clapham]
39651		E. Peñalver and P. Nel. 2010. Hispanothrips from Early Cretaceous Spanish amber, a new genus of the resurrected family Stenurothripidae (Insecta: Thysanoptera). Annales de la Société Entomologique de France 46:138-147 [M. Clapham/M. Clapham/M. Clapham]
44262		E. Peñalver, J. Ortega-Blanco, and X. Delclòs. 2010. Mesozoic Evaniidae (Insecta: Hymenoptera) in Spanish amber: reanalysis of the phylogeny of the Evanioidea. Acta Geologica Sinica 84:809-827 [M. Clapham/M. Clapham]
48003		E. Peñalver and J. Szwedo. 2010. Perforissidae (Hemiptera: Fulgoroidea) from the Lower Cretaceous San Just amber (Eastern Spain). Alavesia 3:97-103 [M. Clapham/M. Clapham]
69955		R. Pérez-de la Fuente, M. S. Engel, X. Delclòs and E. Peñalver. 2020. Straight-jawed lacewing larvae (Neuroptera) from Lower Cretaceous Spanish amber, with an account on the known amber diversity of neuropterid immatures. Cretaceous Research 106(104200) [M. Clapham/M. Clapham/M. Clapham]
70231		R. Pérez-de la Fuente and E. Peñalver. 2019. A mantidfy in Cretaceous Spanish amber provides insights into the evolution of integumentary specialisations on the raptorial foreleg. Scientific Reports 9(13248):1-16 [M. Clapham/M. Clapham/M. Clapham]
47526		R. Pérez-de la Fuente, E. E. Saupe, and P. A. Selden. 2013. New lagonomegopid spiders (Araneae: †Lagonomegopidae) from Early Cretaceous Spanish amber. Journal of Systematic Palaeontology 11:531-553 [M. Clapham/M. Clapham]
51873		D. Peris, S. R. Davis, M. S. Engel and X. Delclòs. 2014. An evolutionary history embedded in amber: reflection of the Mesozoic shift in weevil-dominated (Coleoptera: Curculionoidea) faunas. Zoological Journal of the Linnean Society 171:534-553 [M. Clapham/M. Clapham]
53709		D. Peris, T. K. Philips, and X. Delclòs. 2015. Ptinid beetles from the Cretaceous gymnosperm-dominated forests. Cretaceous Research 52:440-452 [M. Clapham/M. Clapham]
79612		M. Santer, S. Álvarez-Parra, A. Nel, E. Peñalver, and X. Delclòs. 2021. New insights into the enigmatic Cretaceous family Spathiopterygidae (Hymenoptera: Diaprioidea). Cretaceous Research 133(105128) [M. Clapham/M. Clapham]
47396		E. E. Saupe, R. Pérez-de la Fuente, P. A. Selden, X. Delclòs, P. Tafforeau and C. Soriano. 2012. New Orchestina Simon, 1882 (Araneae: Oonopidae) from Cretaceous ambers of Spain and France: first spiders described using phase-contrast X-ray synchrotron microtomography. Palaeontology 55:127-143 [M. Clapham/M. Clapham]
56527		R. Szadziewski, A. Arillo, A. Urbanek and E. Sontag. 2016. Biting midges of the extinct genus Protoculicoides Boesel from Lower Cretaceous amber of San Just, Spain and new synonymy in recently described fossil genera (Diptera: Ceratopogonidae). Cretaceous Research 58:1-9 [M. Clapham/M. Clapham]