Cleveland-Lloyd Dinosaur Quarry: Tithonian, Utah
collected by Hintze & York; Stokes, Warner, Body & Jepsen; Madsen & Stokes 1927, 1929–1931, 1939–1941, 1960–1967, 1975-1990, 2001–2002, 2012

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Stellatochara obovata
Foster et al. 2016
Charophyta indet. Migula 1890
Bilbey 1993
    = Latochara latitruncata Peck 1937
Foster et al. 2016
Charales - Characeae
Aclistochara bransoni Peck 1937
Foster et al. 2016
Gastropoda - Heterostropha - Valvatidae
Valvata sp. Müller 1774
Foster et al. 2016
Mollusca indet. Linnaeus 1758
Bilbey 1993
"freshwater mollusks"
    = Amplovalvata sp. Yen 1952
Foster et al. 2016
Gastropoda - Viviparidae
Viviparus sp. Montfort 1810
Foster et al. 2016
Reptilia
Reptilia indet. Laurenti 1768
Stokes 1945
Reptilia - Dinosauria
Dinosauria indet. Owen 1842
Hirsch et al. 1989 1 specimen
eggshell
Reptilia - Camarasauridae
Camarasaurus sp. Cope 1877
Ikejiri 2005 1 individual
RTMP 83.35.03, to 83.35.06
Camarasaurus supremus Cope 1877
Chure et al. 2000 3 individuals
UUVP 1555, 1859, 1860, 1984, 3568, 3609, 4273, 4286, 4315, 5309, 6341, 10070, 11626, 10795, 22940
    = Camarasaurus cf. lentus Marsh 1889
Smith 1997
    = Camarasaurus ? lentus Marsh 1889
Ikejiri 2005
Camarasaurus lentus (Marsh 1889)
Ikejiri 2005 1 specimen
CEU 1694
Reptilia
Brontosaurus sp. Marsh 1879
Hou 1979 1 individual
    = ? Apatosaurus sp. Marsh 1877
Madsen and Stokes 1972
Apatosaurus sp. Marsh 1877
Foster and Peterson 2016 1 specimen
posterior cervical vertebra
Diplodocus sp. Marsh 1878
Hou 1979 1 individual
    = ? Diplodocus sp. Marsh 1878
Madsen and Stokes 1972
    = Barosaurus sp. Marsh 1890
Foster et al. 2016
Reptilia - Nodosauridae
? Acanthopholidae indet. (Nopcsa 1902)
Madsen and Stokes 1972
synonym of Nodosauridae
"unknown sp. (probably assignable to Acanthopholidae)"
Reptilia - Stegosauridae
Stegosaurus ? sp. Marsh 1877
Madsen and Stokes 1972 1 individual
    = Stegosaurus sp. Marsh 1877
Madsen and Stokes 1978
Stegosaurus sp. Marsh 1877
Hou 1979 4 individuals
    = Stegosaurus stenops Marsh 1887
Madsen and Stokes 1972
    = Stegosaurus cf. stenops Marsh 1887
Smith 1997
Reptilia
Camptosaurus sp. Marsh 1885
Hou 1979 5 individuals
    = Camptosaurus cf. medius Marsh 1894
Madsen and Stokes 1978
    = Camptosaurus cf. browni Gilmore 1909
Smith 1997
synonym of Camptosaurus dispar
Reptilia - Prismatoolithidae
Prismatoolithus coloradensis Hirsch 1994
Hirsch 1994 1 specimen
recombined as Preprismatoolithus coloradensis
11584/BYU-VP 12019/HEC 464, 488
Reptilia
Stokesosaurus clevelandi Madsen 1974
Madsen 1976 1 specimen
UUVP 2999/UMNH 7821
    = Tanycolagreus topwilsoni Carpenter et al. 2005
Carpenter et al. 2005
Coelurus sp. Marsh 1879
Madsen and Stokes 1972 2 specimens
"(n. sp.?)"
    = Coelurus fragilis Marsh 1879
Carpenter et al. 2005
UUVP 11743, 7795
Stokesosaurus clevelandi n. gen., n. sp. Madsen 1974
Madsen 1974 1 individual
Stokesosaurus sp. Madsen 1974
Chure and Madsen 1993 1 specimen
UUVP 2455
    = ? Stokesosaurus clevelandi Madsen 1974
Chure and Madsen 1998
Reptilia - Piatnitzkysauridae
Marshosaurus bicentesimus n. gen., n. sp. Madsen 1976
Madsen 1976 2 individuals
Reptilia
Ceratosaurus sp. Marsh 1884
Hou 1979 1 individual
(4 measurements)
    = Ceratosaurus dentisulcatus n. sp. Madsen and Welles 2000
Madsen and Welles 2000
synonym of Ceratosaurus nasicornis
UMNH VP 5278
Antrodemus sp. Leidy 1873
Hou 1979 46 individuals
UMNH, YPM-PU, MHNG GEPI V2567, MNHN
    = Allosaurus fragilis Marsh 1877
Stokes 1961
Reptilia - Goniopholididae
Crocodylia indet. (Owen 1842)
Stokes 1945 1 individual
    = Goniopholis sp. Owen 1842
Madsen and Stokes 1972
Reptilia - Pleurosternidae
Glyptops sp. Marsh 1890
Miller et al. 1996 2 individuals
see common names

Geography
Country:United States State/province:Utah County:Emery
Coordinates: 39.3° North, 110.7° West (view map)
Paleocoordinates:36.0° North, 57.6° West
Basis of coordinate:stated in text
Altitude:5760 feet
Geographic resolution:outcrop
Time
Period:Jurassic Epoch:Late/Upper Jurassic
Stage:Tithonian 10 m.y. bin:Jurassic 6
Key time interval:Tithonian
Age range of interval:152.10000 - 145.00000 m.y. ago
Stratigraphy
Formation:Morrison Member:Brushy Basin
Stratigraphic resolution:bed
Stratigraphy comments: lower part of member, ca. 38 m above basal contact
Lithology and environment
Primary lithology:nodular,intraclastic,gray calcareous mudstone
Secondary lithology: lime mudstone
Includes fossils?Y
Lithology description: "a drab gray calcareous claystone...capped by a fossiliferous micritic limestone"; "a calcarous mudstone that...also includes abundant diagenetic limestone nodules and clay clasts."
Environment:pond
Geology comments: channel-top swale or oxbow pond in poorly drained floodplain/delta; overlain by lacustrine facies
Taphonomy
Modes of preservation:body
Lagerst�tten type:concentrate
Degree of concentration:-multiple events
Size of fossils:macrofossils,mesofossils,microfossils
Spatial orientation:random
Preservation of anatomical detail:good
Abundance in sediment:abundant
Articulated whole bodies:none
Associated major elements:none
Disassociated major elements:many
Disassociated minor elements:many
Size sorting:poor
Fragmentation:occasional
Bioerosion:occasional
Feeding/predation traces:arthropod boring
Temporal resolution:time-averaged
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:core,selective quarrying,mechanical,field collection,survey of museum collection
Collection size:5000 specimens
Reason for describing collection:taxonomic analysis
Collectors:Hintze & York; Stokes, Warner, Body & Jepsen; Madsen & Stokes Collection dates:1927, 1929–1931, 1939–1941, 1960–1967, 1975-1990, 2001–2002, 2012
Metadata
Also known as:CLDQ, Cleveland Quarry, Malcolm Lloyd Jr. Quarry, UT-7
Database number:46439
Authorizer:M. Carrano Enterer:M. Carrano, K. Maguire
Modifier:M. Carrano Research group:vertebrate
Created:2005-01-13 08:24:39 Last modified:2023-11-21 16:35:18
Access level:the public Released:2005-01-13 08:24:39
Creative Commons license:CC BY
Reference information

Primary reference:

62377. W. L. Stokes. 1944. Jurassic dinosaurs from Emery County, Utah. Proceedings of the Utah Academy of Sciences, Arts and Letters 21:11 [M. Carrano/M. Carrano]

Secondary references:

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53052 J. H. Madsen Jr. and W. L. Stokes. 1978. University of Utah Dinosaur Project: a final report, 1971–1977. Encyclia 54(2):68-70 [M. Carrano/M. Carrano]
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62297 W. L. Stokes. 1985. The Cleveland-Lloyd Dinosaur Quarry: Window to the Past 1985-0-576:1-27 [M. Carrano/M. Carrano]
52112 W. L. Stokes and J. H. Madsen, Jr. 1962. Progress report on excavations at Cleveland-Lloyd Dinosaur Quarry, Emery County, Utah. Proceedings of the Utah Academy of Sciences, Arts and Letters 39:200 [M. Carrano/A. Murch]
52113 W. L. Stokes and J. H. Madsen, Jr. 1967. Status and contributions of the University of Utah Cooperative Dinosaur Project (1967). Proceedings of the Utah Academy of Sciences, Arts and Letters 44(1):385-386 [M. Carrano/A. Murch]
58662 C. Suarez and M. Suarez. 2004. Use of facies, taphonomy, and rare earth element geochemistry analyses at the Cleveland-Lloyd Dinosaur Quarry: tools for bone bed interpretations. Journal of Vertebrate Paleontology 24(3, suppl.):119A [M. Carrano/M. Carrano]
76532 A. L. Titus, K. Knoll, J. J. W. Sertich, D. Yamamura, C. A. Suarez, I. J. Glasspool, J. E. Ginouves, A. K. Lukacic, and E. M. Roberts. 2021. Geology and taphonomy of a unique tyrannosaurid bonebed from the upper Campanian Kaiparowits Formation of southern Utah: implications for tyrannosaurid gregariousness. PeerJ 9:e11013:1-50 [P. Mannion/P. Mannion/M. Carrano]
62962 S. P. Welles. 1972. Dinosaurs come to the Academy. Pacific Discovery 25(5):3-11 [M. Carrano/M. Carrano]
55126 O. Wings. 2015. The rarity of gastroliths in sauropod dinosaurs – a case study in the Late Jurassic Morrison Formation, western USA. Fossil Record 18(1):1-16 [M. Carrano/M. Carrano]