Boca del Sapo, Universidad Nacional del Comahue: Santonian, Argentina
collected by O. de Ferrariis, J. O. Calvo 1984-1987

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia
Ornithothoraces indet. Chiappe and Calvo 1994
Chiappe 2007 8 specimens
MUCPv-284, 305–306, 350–355 (eggs and embryos)
Reptilia - Avisauridae
Enantiornithes indet. (Walker 1981)
Chiappe 1991 1 individual
MUCHPv-142
    = Neuquenornis volans n. gen., n. sp. Chiappe and Calvo 1994
Chiappe and Calvo 1994
Reptilia
Patagopteryx deferrariisi n. gen., n. sp. Alvarenga and Bonaparte 1992
Alvarenga and Bonaparte 1992 8 individuals
MACN-N 03, 10-11, 14, 48, 207
Reptilia - Alvarezsauridae
Alvarezsaurus calvoi n. gen., n. sp. Bonaparte 1991
Bonaparte 1991 1 individual
MUCPv-54 (1 measurement)
Reptilia
Paleochelco occultato n. gen., n. sp. Martinelli et al. 2021
Martinelli et al. 2021
MACN-Pv-N 120 - holotype
see common names

Geography
Country:Argentina State/province:Neuquén
Coordinates: 39.0° South, 68.2° West (view map)
Paleocoordinates:43.5° South, 49.2° West
Basis of coordinate:estimated from map
Altitude:260 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Santonian 10 m.y. bin:Cretaceous 6
*Period:Late/Upper Cretaceous *Epoch:Senonian
*International age/stage:Coniacian - Campanian
Key time interval:Santonian
Age range of interval:86.30000 - 83.60000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Geological group:Neuquén Formation:Bajo de la Carpa
Stratigraphic resolution:group of beds
Stratigraphy comments: Rio Colorado Subgroup
Lithology and environment
Primary lithology:fine,medium,red,white poorly lithified sandstone
Includes fossils?Y
Lithology description: "predominantly composed of medium to fine, friable, whitish gray to pink sandstones, generally made of clean quartz with very little clay."
Environment:fluvial-deltaic indet.
Geology comments: Derived from Chiappe and Calvo (1994)
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:good
Articulated whole bodies:many
Associated major elements:many
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:selective quarrying,mechanical,field collection
Reason for describing collection:taxonomic analysis
Collectors:O. de Ferrariis, J. O. Calvo Collection dates:1984-1987
Collection method comments: Chiappe (1991) notes that "The Bajo de la Carpa Member yielded several other remains of continental vertebrates such as notosuchian crocodiles, dynilisid ophidians, and small theropod dinosaurs (Bonaparte, 1987)". Alvarezsaurus found by J. Calvo in May 1987.
Chiappe (1991) also notes that the collection includes a nearly complete skeleton and partial skeleton of a then unpublished cursorial flightless bird (later named Patagopteryx deferrariisi by Alvarenga and Bonaparte, 1992) and a complete, articulated skeleton of a then unpublished enantiornithine bird (later named Neuquenornis volans by Chiappe and Calvo, 1994).
Taxonomic list comments:Birds named in this list are from later publications (Alvarenga and Bonaparte, 1992, and Chiappe and Calvo, 1994)
Metadata
Database number:13305
Authorizer:J. Hunter, M. Carrano, P. Mannion Enterer:J. Hunter, P. Mannion, M. Carrano
Modifier:M. Carrano Research group:vertebrate
Created:2002-02-20 11:47:30 Last modified:2013-09-12 10:44:04
Access level:the public Released:2002-02-20 11:47:30
Creative Commons license:CC BY
Reference information

Primary reference:
4423.5% 19300L. M. Chiappe. 1991. Cretaceous birds of Latin America. Cretaceous Research 12(1):55-63 [J. Hunter/J. Hunter/J. Hunter]

Secondary references:

6047 H. M. F. Alvarenga and J. F. Bonaparte. 1992. A new flightless landbird from the Cretaceous of Patagonia. Los Angeles County Museum of Natural History, Science Series 36:51-64 [J. Hunter/J. Hunter/J. Hunter]
12794 J. F. Bonaparte. 1991. Los vertebrados fósiles de la Formación Rio Colorado, de la Ciudad de Neuquén y Cercanías, Cretácico Superior, Argentina [The vertebrate fossils of the Rio Colorado Formation, from the city of Neuquén and surrounding areas, Upper Cretaceous, Argentina]. Revista del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigación de las Ciencias Naturales: Paleontología 4(3):17-123 [J. Head/J. Head/M. Carrano]
16547ETE J. F. Bonaparte. 1996. Cretaceous tetrapods of Argentina. Münchner Geowissenschaften Abhandlungen 30:73-130 [M. Carrano/M. Carrano/M. Carrano]
45878 L. M. Chiappe. 1991. Cretaceous avian remains from Patagonia shed new light on the early radiation of birds. Alcheringa 15(4):333-338 [P. Mannion/J. Tennant/M. Carrano]
12947ETE L. M. Chiappe. 1996. Early avian evolution on the southern hemisphere: the fossil record of birds in the Mesozoic of Gondwana. Memoirs of the Queensland Museum 39(3):533-554 [M. Carrano/M. Carrano/M. Carrano]
29151ETE L. M. Chiappe. 2002. Osteology of the flightless Patagopteryx deferrariisi from the Late Cretaceous of Patagonia (Argentina). In L. M. Chiappe and L. M. Witmer (eds.), Mesozoic Birds: Above the Heads of Dinosaurs. University of California Press, Berkeley 281-361 [M. Carrano/M. Carrano]
48178 L. M. Chiappe. 2007. Aves. In Z. B. Gasparini, L. Sangado & R. A. Coria (eds.), Patagonian Mesozoic Reptiles 257-270 [M. Carrano/M. Carrano]
6046 L. M. Chiappe and J. O. Calvo. 1994. Neuquenornis volans, a new Late Cretaceous bird (Enantiornithes, Avisauridae) from Patagonia, Argentina. Journal of Vertebrate Paleontology 14(2):230-246 [J. Hunter/J. Hunter/M. Carrano]
84750 A. G. Martinelli, F. L. Agnolín, and M. D. Ezcurra. 2021. Unexpected new lizard from the Late Cretaceous of southern South America sheds light on Gondwanan squamate diversity. Revista del Museo Argentino de Ciencias Naturales, nueva serie 23(1):57-80 [P. Mannion/P. Mannion]