San Gerardo de Limoncito: Hemphillian, Costa Rica

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Proboscidea - Gomphotheriidae
Rhynchotherium blicki (Frick 1933)
Valerio and Laurito 2008
CFM-2837, 3224, 3361, 3364 and 3365
Mammalia - Mylodontidae
Mylodontidae indet. Ameghino 1889
Laurito and Valerio 2012
Mammalia - Megatheriidae
Megatheriidae indet. (Gray 1821)
Laurito and Valerio 2012
    = Sibotherium ka n. gen., n. sp. Rincón et al. 2020
Rincón et al. 2020
CFM-994 (Type), CFM-3713, CFM-1785, CFM- 3058, CFM-2625, CFM-1302, CFM-1840, CFM-3795, CFM-1645, CFM-2498, CFM-998, CFM-1301, CFM-1239, CFM-1243, CFM-2607, CFM-3731, CFM-3634, 3000, 1532, 1530, 1447, 1858, CFM-1531, 2604, 2606, 1696; CFM-1654, ;CFM-3813, CFM-995,
Mammalia - Megalonychidae
Zacatzontli cotobrusensis n. sp. Valerio et al. 2022
Valerio et al. 2022
Holotype: CFM-2916; CFM-2917, first upper molariform; CFM-1000, molarifiorm. CFM-1000 and CFM-2917 were previously referred to as Pliometanastes cf. protistus
Pliometanastes cf. protistus Hirschfeld and Webb 1968
Laurito and Valerio 2012
CFM-2017, and 2645
Mammalia - Cingulata - Pampatheriidae
Scirrotherium antelucanus n. sp. Laurito and Valerio 2013
Laurito and Valerio 2013
Holotype CFM 2867
Mammalia - Camelidae
Hemiauchenia sp. Gervais and Ameghino 1880
Laurito and Valerio 2012
    = Hemiauchenia vera Matthew 1909
Laurito and Valerio 2016
recombined as Pleiolama vera
Mammalia - Tayassuidae
Tayassuidae indet. Palmer 1897
Laurito and Valerio 2012
Protherohyus brachydontus (Dalquest and Mooser 1980)
Valerio and Laurito 2020
CFM-2195
Prosthennops serus (Cope 1877)
Valerio and Laurito 2020
CFM-2915, CFM-1861, CFM-1862, CFM-1499, CFM-1652,CFM-1651,
Mammalia - Perissodactyla - Equidae
Dinohippus mexicanus (Lance 1950)
Laurito and Valerio 2010
original and current combination Pliohippus (Pliohippus) mexicanus
Protohippus gidleyi Hulbert 1988
Laurito and Valerio 2010
Calippus (Grammohippus) hondurensis (Olson and McGrew 1941)
Laurito and Valerio 2010
recombined as Calippus hondurensis
Reptilia
Gavialosuchus americana (Sellards 1915)
1 specimen
synonym of Thecachampsa sericodon
CFM-1793 (jaw fragments), CFM-1792 (jaw fragment) and CFM-960, 972, 1774, 1912, 1913, 1916 & 1923 (osteoderms)
Reptilia - Testudines - Emydidae
Emydinae indet. Rafinesque 1815
Laurito and Valerio 2012
Reptilia - Testudines - Trionychidae
Apalone sp. Rafinesque 1832
Laurito et al. 2005
CFM-1163-1166 and 1193, Five shell specimens
Actinopteri - Siluriformes - Ariidae
Arius sp. Cuvier and Valenciennes 1840
Laurito and Valerio 2012
Actinopteri - Perciformes - Sparidae
Sparus sp. Linnaeus 1758
Laurito and Valerio 2012
Chondrichthyes - Carcharhiniformes - Hemigaleidae
Hemipristis serra Agassiz 1835
Laurito and Valerio 2012
Chondrichthyes - Carcharhiniformes - Carcharhinidae
Isogomphodon acuarius
Laurito and Valerio 2012
Isogomphodon caunellensis
Laurito and Valerio 2012
Carcharhinus longimanus
Laurito and Valerio 2012
Carcharhinus priscus Agassiz 1843
Laurito and Valerio 2012
Chondrichthyes - Carcharhiniformes - Sphyrnidae
Sphyrna arambourgi
Laurito and Valerio 2012
Chondrichthyes - Lamniformes - Lamnidae
Isurus desorii (Agassiz 1843)
Laurito and Valerio 2012
synonym of Isurus oxyrinchus
Chondrichthyes - Myliobatiformes - Aetobatidae
Aetobatus arcuatus (Agassiz 1843)
Laurito and Valerio 2012
synonym of Aetobatus irregularis
Chondrichthyes - Myliobatiformes - Dasyatidae
Dasyatis cavernosa Probst 1877
Laurito and Valerio 2012
Chondrichthyes - Myliobatiformes - Myliobatidae
Rhinoptera sp. Cuvier 1829
Laurito and Valerio 2012
Myliobatis sp. Cuvier 1816
Laurito and Valerio 2012
see common names

Geography
Country:Costa Rica State/province:Puntarenas
Coordinates: 8.9° North, 83.1° West (view map)
Paleocoordinates:8.9° North, 83.1° West
Basis of coordinate:stated in text
Geographic resolution:small collection
Time
Period:Neogene
10 m.y. bin:Cenozoic 6
Key time interval:Hemphillian
Age range of interval:9.40000 - 4.70000 m.y. ago
Age estimate:maximum 5.8 ± 0.6 Ma (Sr isotope)
Stratigraphy
Formation:Curré
Stratigraphic resolution:group of beds
Stratigraphy comments: Age: Late Hemphillian (Hh3) (Late Miocene), Curré Formation, San Gerardo de Limoncito, Costa Rica. 87Sr/86Sr 5.80 Ma +/- 0.60/-0.80

The rock unit that contains the fossiliferous remains corresponds to the Curré Formation. Henningsen (1965) defines this formation as a series of tuffaceous sandstones with intercalations of conglomerates, siltstones and occasionally shales, dark green and brownish green when altered. Mora (1979), describes two subunits according to their sedimentological characteris- tics: The subunit of volcaniclastic conglomerates of gray-green color, overlain by shales that settled in a transitional environ- ment between delta and beach. The subunit of black shales, interspersed with strata of sandstones and fine conglomerates, was deposited in a paralic environment of estuary or swamp type. They are very chloritized and basically consist of subvolcanic products.
The paleontological material recovered is housed at the Geology Section of the Natural History Department at The National Museum of Costa Rica, and comes from a level of fine conglomerate that is located towards the roof of a sequence of blue clays, which in turn is overlain by a sequence of medium to fine sandstones, which confirms its association with the superior subunit or subunit of shales (sensu Mora, 1979). Granados and Aguilar (1983), based on the presence of molluscs, obtained a Middle-Late Miocene age for the Curré Formation.

Lithology and environment
Primary lithology: conglomerate
Lithology description: Subaquatic fan delta: The lithofacies of conglomerates is what characterizes this depositional environment and is composed of blue clay intraclasts, coming from the lower level which it overlies at an erosional discordance. The clasts have diameters that vary between 5 and 13 cm; they are well rounded, little or not spherical; some have point contact, but 90% of them float in a matrix of glauconitic clay. The clasts are arranged with their long axis parallel to the layer of stratification, and it is possible to observe some degree of imbrication. In this sequence are frequent bone and dental remains of various vertebrates, both terrestrial and marine. The Trypanites ichnofacies, in this case, is inferred indirectly from the blue clay intraclasts that are usually perforated by bivalves of the family Pholadidae, from which it is still possible to find remains of the valves within the fossil traces of Gastrochaenolites sp.
Environment:deltaic indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Metadata
Database number:144596
Authorizer:P. Mannion, C. Jaramillo, E. Vlachos Enterer:P. Mannion, G. Varnham, E. Vlachos, J. Carrillo, M. Kouvari
Modifier:G. Varnham Research group:vertebrate
Created:2013-05-13 23:56:42 Last modified:2022-03-15 10:25:38
Access level:the public Released:2013-05-13 23:56:42
Creative Commons license:CC BY
Reference information

Primary reference:

46789. C. A. Laurito and A. L. Valerio. 2008. The first record of Gavialosuchus americanus Sellards (1915)† (Eusuchia: Crocodylidae, Tomistominae) for the Late Tertiary of Costa Rica and Central America. Revista Geológica de América Central 39:107-115 [P. Mannion/P. Mannion]

Secondary references:

64966 C. A. Laurito and A. L. Valerio. 2010. Los caballos fósiles de la Formación Curré, Cantón de Coto Brus, Costa Rica. 1-131 [C. Jaramillo/J. Carrillo]
64950 C. A. Laurito and A. L. Valerio. 2012. Paleobiogeografía del arribo de mamíferos suramericanos al sur de América Central de previo al Gran Intercambio Biótico Americano: un vistazo al GABI en América Central. Revista Geológica de América Central 46:123-144 [C. Jaramillo/J. Carrillo]
64951 C. A. Laurito and A. L. Valerio. 2012. Primer registro fósil de Pliometanastes sp. (Mammalia, Xenarthra, Megalonychidae) para el Mioceno superior de Costa Rica, América Central. Una nueva pista en la compresión del pre-GABI. Revista Geológica de América Central 47:95-108 [C. Jaramillo/J. Carrillo]
64952 C. A. Laurito and A. L. Valerio. 2013. Scirrotherium antelucanus, una nueva especie de Pampatheriidae (Mammalia, Xenarthra, Cingulata) del Mioceno superior de Costa Rica, América Central. Revista Geológica de América Central 49:45-62 [C. Jaramillo/J. Carrillo]
64965 C. A. Laurito and A. L. Valerio. 2016. Camellos Laminos del Mioceno tardío (Henfiliano temprano) de la Formación Curré, San Gerardo de Limoncito, Cantón de Coto Brus, Provincia de Puntarenas, Costa Rica. Revista Geológica de América Central 54:7-55 [C. Jaramillo/J. Carrillo]
64124 C. M. Laurito, A. L. Valerio, L. D. Gomez, J. I. Mead, E. A. Perez and L. G. Perez. 2005. A Trionychidae (Reptilia: Testudines, Cryptodira) from the Pliocene of Costa Rica, Southern Central America. Revista Geologica de America Central 32:7-11 [E. Vlachos/E. Vlachos]
73190 A. D. Rincón, A. L. Valerio, and C. Laurito. 2020. First fossil record of a Megatheriidae-Megatheriinae in the Early Hemphillian (Late Miocene) from San Gerardo de Limoncito, Curré Formation, Costa Rica. Revista Geológica de América Central 62:1-24 [P. Mannion/M. Kouvari]
64931 A. L. Valerio and C. A. Laurito. 2008. Dental remains of immature individuals of Rhynchotherium blicki (Frick 1933) of San Gerardo de Limoncito, Coto, Brus, Costa Rica. Revista Geológica de América Central 39:87-91 [C. Jaramillo/J. Carrillo]
73192 A. L. Valerio and C. A. Laurito. 2014. Nueva evidencia de un estadio juvenil de Pliometanastes protistus y su relación con ?Pliometanastes galushai Hirschfeld & Webb, 1968 (Xernarthra, Megalonychidae) para el Mioceno Superior de Costa Rica. Revista Geológica de América Central 51:159-163 [P. Mannion/M. Kouvari]
73193 A. L. Valerio and C. A. Laurito. 2020. Primer registro de pecaríes fósiles (Artiodactyla, Tayassuidae) para el Mioceno Superior (Hh3: Hemphilliano tardío) de Costa Rica, América. Revista Geológica de América Central 62:25-47 [P. Mannion/M. Kouvari]
80759 A. L. Valerio, C. Laurito, H. G. McDonald and A. D. Rincón. 2022. Megalonychid Sloths from the Early Late Hemphillian (Late Miocene), Curré Formation, San Gerardo de Limoncito, Costa Rica. Revista Geológica de América Central 66:1-15 [P. Mannion/G. Varnham]