Tugrikin Shire I [HMNS-GIN]: Late/Upper Campanian, Mongolia
collected by HMNS-GIN 1992-1995, 1998, 2000, 2007

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia
Mammalia indet. (Linnaeus 1758)
Watabe and Suzuki 2000
mammals
Reptilia
Lacertilia indet. (Owen 1842)
Suzuki and Watabe 2000
lizards, incl. skeleton field no. 980725 TS Lizard ORKHON
? Lacertilia indet. (Owen 1842)
Watabe and Tsogtbaatar 2004 1 individual
skull and postcrania; 000711 TS SZK LIZARD?
Reptilia - Ankylosauridae
Ankylosauridae indet. Brown 1908
Suzuki and Watabe 2000 1 specimen
MPC-D 100/1337
Pinacosaurus grangeri Gilmore 1933
Chiappe et al. 2007
Reptilia - Protoceratopsidae
Protoceratops sp. Granger and Gregory 1923
Matsumoto 1997 4 individuals
2 skeletons, 930902-TS-I-1, 2, 8, 940801 TS-I SZK, 070619 TS TSUI, 070619 TS SZK, 070620 TS TSGT
    = Protoceratops andrewsi Granger and Gregory 1923
Matsumoto et al. 2000
MPC-D 100/530, 533, 534, 526
    = Protoceratops cf. andrewsi Granger and Gregory 1923
Fastovsky et al. 2011
Reptilia - Hadrosauridae
Hadrosauridae indet. Cope 1869
Watabe and Tsogtbaatar 2004 1 individual
field no. 000714-TS-I (baby skeleton)
Reptilia - Prismatoolithidae
Prismatoolithidae indet. Hirsch 1994
Suzuki and Watabe 2000
Reptilia - Troodontidae
Velociraptor sp. Osborn 1924
Suzuki and Watabe 2000 1 individual
940801 TS-I WTB
    = Troodontidae indet. Gilmore 1924
Tsogtbaatar and Chinzorig 2010
Reptilia - Dromaeosauridae
Velociraptor sp. Osborn 1924
Suzuki and Watabe 2000 1 individual
field no. 980725-TS-I Velocira NAR
    = Velociraptor mongoliensis Osborn 1924
Chiappe et al. 2007
MPC-D 100/54
Reptilia - Ornithomimidae
Ornithomimidae indet. Marsh 1890
Watabe and Suzuki 2000 1 specimen
940801 TS-I SZK
    = Aepyornithomimus tugrikinensis n. gen., n. sp. Chinzorig et al. 2017
Chinzorig et al. 2017
MPC-D 100/130
Reptilia - Elongatoolithidae
Elongatoolithidae indet. Zhao 1975
Suzuki and Watabe 2000
see common names

Geography
Country:Mongolia State/province:Omnogov
Coordinates: 44.2° North, 103.3° East (view map)
Paleocoordinates:41.8° North, 92.3° East
Basis of coordinate:stated in text
Altitude:1043 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
Key time interval:Late/Upper Campanian
Age range of interval:83.50000 - 70.60000 m.y. ago
Stratigraphy
Formation:Djadokhta Member:Tugrugyin
Stratigraphic resolution:group of beds
Lithology and environment
Primary lithology:"cross stratification",burrows,fine,red,white poorly lithified sandstone
Includes fossils?Y
Lithology description: "yellowish-white sandstones"; "Tugrik sandstones are compositionally and texturally sub-mature. They are generally poorly consolidated, moderately well-sorted, fine-grained, sub-rounded to well-rounded, and present a 'frosted' appearance. Colors vary from pink (induced by minor amounts of hematite), to white, reflecting the absence of any hematitie or other coloring agent."
Environment:dune
Geology comments: eolian dune, erg
Taphonomy
Modes of preservation:body
Degree of concentration:dispersed
Size of fossils:macrofossils,mesofossils
Spatial orientation:preferred
Preservation of anatomical detail:excellent
Abundance in sediment:abundant
Articulated whole bodies:many
Associated major elements:many
Disassociated major elements:some
Disassociated minor elements:some
Fragmentation:occasional
Feeding/predation traces:tooth marks,arthropod boring
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:surface (float),surface (in situ),mechanical,field collection,observed (not collected)
Reason for describing collection:general faunal/floral analysis
Collectors:HMNS-GIN Collection dates:1992-1995, 1998, 2000, 2007
Metadata
Also known as:Tögrögiin Shiree, Tugrig Shire, Toogreeg Shire
Database number:55256
Authorizer:M. Carrano, R. Benson Enterer:M. Carrano, R. Benson
Modifier:M. Carrano Research group:vertebrate
Created:2005-10-03 15:00:50 Last modified:2022-02-23 15:31:33
Access level:the public Released:2005-10-03 15:00:50
Creative Commons license:CC BY
Reference information

Primary reference:

14738.ETE M. Watabe and D. B. Weishampel. 1994. Results of Hayashibara Museum of Natural Sciences–Institute of Geology, Academy of Sciences of Mongolia Joint Paleontological Expedition to the Gobi Desert in 1993. Journal of Vertebrate Paleontology 14(3, suppl.):51A [M. Carrano/M. Carrano]

Secondary references:

37931 V. R. Alifanov. 1989. [New priscagamas (Lacertilia) from the Upper Cretaceous of Mongolia and their systematic position in the Iguania]. Paleontologiceskij Zhurnal 4:73-87 [R. Benson/R. Benson]
28193ETE L. M. Chiappe, S. Suzuki, G. J. Dyke, M. Watabe, K. Tsogtbaatar and R. Barsbold. 2007. A new enantiornithine bird from the Late Cretaceous of the Gobi Desert. Journal of Systematic Palaeontology 5(2):193-208 [M. Carrano/M. Carrano]
65627 T. Chinzorig, Y. Kobayashi, K. Tsogtbaatar, P. J. Currie, M. Watabe and R. Barsbold. 2017. First ornithomimid (Theropoda, Ornithomimosauria) from the Upper Cretaceous Djadokhta Formation of Tögrögiin Shiree, Mongolia. Scientific Reports 7(1):5835:1-14 [M. Carrano/M. Carrano]
15459ETE D. Dashzeveg, L. Dingus, D.B. Loope, C.C. Swisher, III, T. Dulam and M.R. Sweeney. 2005. New stratigraphic subdivision, depositional environment, and age estimate for the Upper Cretaceous Djadokhta Formation, southern Ulan Nur Basin, Mongolia. American Museum Novitates (3498)1-31 [M. Carrano/K. Maguire/G. Varnham]
76481 D. E. Fastovsky. 1997. Short summary of the geologyof some selected dinosaur-bearing localities in the western Gobi Desert, Mongolia. Abstract of Report Meeting, Mongolia-Japan Joint Paleontological Expedition 6 [M. Carrano/M. Carrano]
50165 D. E. Fastovsky, D. B. Weishampel, M. Watabe, R. Barsbold, K. Tsogtbaatar and P. Narmandakh. 2011. A nest of Protoceratops andrewsi (Dinosauria, Ornithischia). Journal of Paleontology 85(6):1035-1041 [M. Carrano/M. Carrano]
53635 D. W. E. Hone, A. A. Farke, M. Watabe, S. Shigeru, and K. Tsogtbaatar. 2014. A new mass mortality of juvenile Protoceratops and size-segregated aggregation behaviour in juvenile non-avian dinosaurs. PLoS ONE 9(11):e113306:1-26 [M. Carrano/M. Carrano]
37171 K.-i. Ishii. 2000. Brief introduction of Hayashibara Museum of Natural Sciences and Mongolian Paleontological Center Joint Paleontological Expedition in the Gobi Desert. Hayashibara Museum of Natural Sciences Research Bulletin 1:6-7 [M. Carrano/M. Carrano]
38300 Z. Kielan-Jaworowska and J. H. Hurum. 1997. Djadochtatheria - a new suborder of multituberculate mammals. Acta Palaeontologica Polonica 42(2):201-242 [R. Benson/R. Benson]
76474 Y. Matsumoto. 1997. A reconstruction of the stance of Protoceratops based on the observation of the articulated skeletons. Abstract of Report Meeting, Mongolia-Japan Joint Paleontological Expedition 13 [M. Carrano/M. Carrano]
82301 Y. Matsumoto, R. Hashimoto, and T. Sonoda. 2000. Report of preparation works for Mongolian specimens in HMNS from March 1994 to December 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:113-127 [M. Carrano/M. Carrano]
41448 M. Saneyoshi, M. Watabe, S. Suzuki and K. Tsogtbaatar. 2011. Trace fossils on dinosaur bones from Upper Cretaceous eolian deposits in Mongolia: taphonomic interpretation of paleoecosystems in ancient desert environments. Palaeogeography, Palaeoclimatology, Palaeoecology 311(1):38-47 [M. Carrano/M. Carrano]
42293 S. Suzuki and M. Watabe. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:83-98 [R. Benson/R. Benson/M. Carrano]
42153 S. Suzuki and M. Watabe. 2000. Report on the preliminary joint field excursion to the Gobi desert, 1992. Hayashibara Museum of Natural Sciences Research Bulletin 1:13-16 [R. Benson/R. Benson]
35934ETE K. Tsogtbaatar. 2004. Fossil specimens prepared in Mongolian Paleontological Center 1993-2001. Hayashibara Museum of Natural Sciences Research Bulletin 2:123-128 [M. Carrano/M. Carrano]
80151 K. Tsogtbaatar and T. Chinzorig. 2010. Fossil specimens prepared in Mongolian Paleontological Center: 2002–2008. Hayashibara Museum of Natural Sciences Research Bulletin 3:155-166 [M. Carrano/M. Carrano]
17039ETE M. Watabe, S. Suzuki, K. Tsogtbaatar and R. Barsbold. 1995. Results of the Hayashibara Museum of Natural Sciences and Geological Institute, Academy of Science of Mongolia joint palaeontological expedition in the Gobi Desert: 1994 and 1995. Journal of Vertebrate Paleontology 15(3, suppl.):59A [M. Carrano/M. Carrano]
14764ETE M. Watabe, S. Suzuki, K. Tsogtbaatar, R. Barsbold, and D. B. Weishampel. 1996. Hatchlings of the dinosaur Protoceratops (Ornithischia, Ceratopsia) from the Upper Cretaceous locality Tugrikin Shire, Gobi Desert, Mongolia. Journal of Vertebrate Paleontology 16(3, suppl.):71A [M. Carrano/M. Carrano]
42289 M. Watabe and S. Suzuki. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1993. Hayashibara Museum of Natural Sciences Research Bulletin 1:17-29 [R. Benson/R. Benson/M. Carrano]
42154 M. Watabe and S. Suzuki. 2000. Cretaceous fossil localities and a list of fossils collected by the Hayashibara Museum of Natural Sciences and Mongolian Paleontological Center Joint Paleontological Expedition (JMJPE) from 1993 through 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:99-108 [R. Benson/R. Benson]
42294 M. Watabe and K. Tsogtbaatar. 2004. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 2000. Hayashibara Museum of Natural Sciences Research Bulletin 2:45-67 [R. Benson/R. Benson/M. Carrano]
42291 W. Watabe and S. Suzuki. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1994. Hayashibara Museum of Natural Sciences Research Bulletin 1:30-44 [R. Benson/R. Benson/M. Carrano]