Khulsan (general) [PMPE]: Campanian, Mongolia
collected by PMPE 1970-1971

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Myriapoda - Spirobolida
Diplopoda indet. Gervais 1844
2 specimens
    = Gobiulus sabulosus n. gen., n. sp. Dzik 1975
Dzik 1975
ZPAL MgMyr/6, 7
Reptilia - Avetheropoda
Aves indet. Linnaeus 1758
Sabath 1991 30 specimens
eggs; ZPAL MgOv-I/19, I/21a-c, I/25c-d
Gobipteryx minuta n. gen., n. sp. Elzanowski 1974
Elzanowski 1974 2 specimens
Reptilia - Ankylosauridae
Ankylosauridae indet. Brown 1908
1 specimen
sull wiith skeleton (6 measurements)
Reptilia - Protoceratopsidae
Protoceratopsidae indet. Granger and Gregory 1923
Sabath 1991 40 specimens
ZPAL MgOv-I/5-7, 8a-b, 9a-e, 10a, 12-13, 14a-b, 15a-d, 17, 20, 23-24, 26a-f,; P2, P3 group eggs
? Protoceratopsidae indet. Granger and Gregory 1923
Sabath 1991 2 specimens
ZPAL MgOv-I/10, I/25a-b
? Protoceratops kozlowskii n. sp. Maryanska and Osmólska 1975
Maryanska and Osmólska 1975 1 specimen
synonym of Bagaceratops rozhdestvenskyi
ZPAL MgD 1/117
Reptilia - Bagaceratopsidae
Protoceratops sp. Granger and Gregory 1923
Gradzinski and Jerzykiewicz 1972 2 individuals
    = Lamaceratops tereschenkoi n. gen., n. sp. Alifanov 2003
Alifanov 2003
Reptilia - Squamata - Paravaranidae
Paravaranus angustifrons n. gen., n. sp. Borsuk-Bialynicka 1984
Borsuk-Bialynicka 1984
ZPAL MgR I/67 (holotype), damaged skull
Reptilia - Squamata - Necrosauridae
Proplatynotia longirostrata n. gen., n. sp. Borsuk-Bialynicka 1984
Borsuk-Bialynicka 1984
ZPAL MgR I/68 (holotype), posteriorly damaged skull and mandible
Reptilia - Squamata
Gobiderma pulchra Borsuk-Bialynicka 1984
Borsuk-Bialynicka 1984
ZPAL MgR-I/54, skull fragment
Reptilia - Squamata - Varanidae
Saniwides mongoliensis n. gen., n. sp. Borsuk-Bialynicka 1984
Borsuk-Bialynicka 1984
ZPAL MgR-I/72 (holotype), incomplete and deformed skull and mandible
Telmasaurus grangeri Gilmore 1943
Borsuk-Bialynicka 1984
ZPAL MgR-I/65, skull lacking preorbital part
Reptilia - Squamata
Parviderma inexacta n. gen., n. sp. Borsuk-Bialynicka 1984
Borsuk-Bialynicka 1984
ZPAL MgR-I/43 (holotype), very damaged skull with left mandible
Reptilia - Squamata - Polyglyphanodontidae
Adamisaurus magnidentatus Sulimski 1972
Sulimski 1972 3 individuals
ZPAL MgR-I/13 (partial skull, vertebrae and hand), ZPAL MgR-1/35,42 (partial skulls)
Macrocephalosaurus chulsanensis n. sp. Sulimski 1975
Sulimski 1975 16 specimens
recombined as Gilmoreteius chulsanensis
ZPAL MgR-I/14–18, 20, 22–31
Macrocephalosaurus ? gilmorei Sulimski 1975
Maryanska and Osmólska 1975
recombined as Gilmoreteius gilmorei
Reptilia - Squamata
Polrussia mongoliensis n. gen., n. sp. Borsuk-Bialynicka and Alifanov 1991
Borsuk-Bialynicka and Alifanov 1991 1 individual
ZPAL MgR-I/119
Reptilia - Squamata - Phrynosomatidae
Desertiguana gobiensis n. gen., n. sp. Alifanov 2013
Alifanov 2013
PIN 4487/0, left mandibular ramus without the occipital end
Reptilia - Squamata
Igua minuta n. gen., n. sp. Borsuk-Bialynicka and Alifanov 1991
Borsuk-Bialynicka and Alifanov 1991 1 individual
ZPAL MgR-I/60 (holotype skull)
Reptilia - Squamata - Priscagamidae
Borsuk-Bialynicka and Moody 1984 1 specimen
ZPAL MgR-III/69
    = Priscagama gobiensis Borsuk-Bialynicka and Moody 1984
Gao and Hou 1995
Reptilia - Squamata - Slavoiidae
Slavoia darevskii n. gen., n. sp. Sulimski 1984
Sulimski 1984 29 specimens
Reptilia - Squamata - Carusiidae
Shinisauroides latipalatum Borsuk-Bialynicka 1985
Borsuk-Bialynicka 1985 3 individuals
ZPAL MgR-I/56, 55, 57 (partial skulls)
Reptilia - Squamata
Globaura venusta Borsuk-Bialynicka 1988
Borsuk-Bialynicka 1988 10 individuals
ZPAL MgR-I/45, 46, 48, 49, 51, 55, 118, 47 (partial-complete large skulls), 50, 71 (small skulls)
Eoxanta lacertifrons Borsuk-Bialynicka 1988
Borsuk-Bialynicka 1988 2 individuals
ZPAL MgR-I/61, 74 (partial skulls)
Mammalia
Asioryctes nemegtensis Kielan-Jaworowska 1975
Kielan-Jaworowska 1975 1 specimen
Mammalia - Zalambdalestidae
Barunlestes butleri n. gen., n. sp. Kielan-Jaworowska 1975
Kielan-Jaworowska 1975 1 specimen
ZPAL MgM-I/77 (holotype partial skull and postcranium)
Mammalia
Deltatheridium pretrituberculare Gregory and Simpson 1926
Maryanska and Osmólska 1975
subspecies tardum
Mammalia - Multituberculata - Djadochtatheriidae
Djadochtatherium catopsaloides Kielan-Jaworowska 1974
Kielan-Jaworowska 1974 1 specimen
recombined as Catopsbaatar catopsaloides
fragment of lower jaw
Mammalia - Multituberculata
Chulsanbaatar vulgaris n. gen., n. sp. Kielan-Jaworowska 1974
Kielan-Jaworowska 1974 36 specimens
36 specimens, including holotype skull (ZPAL MgM-I/139)
Nemegtbaatar gobiensis Kielan-Jaworowska 1974
Kielan-Jaworowska 1974
Phaceloolithidae
Dinosauria indet. Owen 1842
eggs
    = Dendroolithidae indet. Zhao and Li 1988
Sabath 1991
ZPAL MgOv-I/16
    = Dendroolithus sp. Zhao and Li 1988
Mikhailov et al. 1994
see common names

Geography
Country:Mongolia State/province:Omnogov
Coordinates: 43.5° North, 101.1° East (view map)
Paleocoordinates:40.7° North, 90.7° East
Basis of coordinate:estimated from map
Altitude:1425 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
*Period:Late/Upper Cretaceous *Epoch:Senonian
*International age/stage:Middle Campanian
Key time interval:Campanian
Age range of interval:83.60000 - 72.10000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Baruungoyot
Stratigraphic resolution:group of beds
Stratigraphy comments: unspecified stratigraphic information
Lithology and environment
Primary lithology:"cross stratification",planar lamination sandstone
Secondary lithology:pebbly conglomeratic sandstone
Includes fossils?Y
Includes fossils?Y
Lithology description: "The sandstone beds are massive, horizontally stratified or cross-stratified."
Environment:eolian indet.
Taphonomy
Modes of preservation:body,mold/impression
Size of fossils:macrofossils
Preservation of anatomical detail:good
Articulated whole bodies:some
Associated major elements:some
Disassociated major elements:some
Disassociated minor elements:some
Fragmentation:occasional
Temporal resolution:time-averaged
Spatial resolution:autochthonous
Collection methods and comments
Collection methods:selective quarrying,surface (float),surface (in situ),mechanical,field collection
Reason for describing collection:general faunal/floral analysis
Collectors:PMPE Collection dates:1970-1971
Metadata
Also known as:Polish-Mongolian Palaeontological Expedition, East Nemegt
Database number:13776
Authorizer:M. Carrano, P. Barrett, P. Mannion, R. Benson Enterer:M. Carrano, T. Cleary, J. Tennant, R. Benson
Modifier:M. Carrano Research group:vertebrate
Created:2002-04-25 13:26:22 Last modified:2019-06-25 12:58:13
Access level:the public Released:2002-04-25 13:26:22
Creative Commons license:CC BY
Reference information

Primary reference:

9899. Z. Kielan-Jaworowska and R. Barsbold. 1972. Narrative of the Polish-Mongolian Palaeontological Expeditions 1967-1971. Palaeontologia Polonica 27:5-136 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

38112 V. R. Alifanov. 2000. The fossil record of Cretaceous lizards from Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 368-389 [R. Benson/R. Benson/M. Carrano]
11744ETE V. R. Alifanov. 2003. Two new dinosaurs of the infraorder Neoceratopsia (Ornithischia) from the Upper Cretaceous of the Nemegt Depression, Mongolian People's Republic. Paleontological Journal 37(5):524-534 [M. Carrano/M. Carrano/M. Carrano]
47932 V. R. Alifanov. 2013. Desertiguana gobiensis gen. et sp. nov., a new lizard (Phrynosomatidae, Iguanomorpha) from the Upper Cretaceous of Mongolia. Paleontological Journal 47(4):417-424 [P. Mannion/J. Tennant]
10439 M. Borsuk-Bialynicka. 1984. Anguimorphans and related lizards from the Late Cretaceous of the Gobi Desert, Mongolia. Palaeontologia Polonica 46:5-105 [M. Carrano/M. Carrano/M. Carrano]
14413ETE M. Borsuk-Bialynicka. 1985. Carolinidae, a new family of xenosaurid-like lizards from the Late Cretaceous of Mongolia. Acta Palaeontologica Polonica 30(3-4):151-176 [M. Carrano/M. Carrano/M. Carrano]
37983 M. Borsuk-Bialynicka. 1988. Globaura venusta gen. et sp. n. and Eoxanta lacertifrons gen. et sp. n. - non-teiid lacertoids from the Late Cretaceous of Mongolia. Acta Palaeontologica Polonica 33(3):211-248 [R. Benson/R. Benson]
38030 M. Borsuk-Bialynicka and V. Alifanov. 1991. First Asiatic 'iguanid' lizards in the Late Cretaceous of Mongolia. Acta Palaeontologica Polonica 36(3):325-342 [R. Benson/R. Benson]
14417ETE M. Borsuk-Bialynicka and S. M. Moody. 1984. Priscagaminae, a new subfamily of the Agamidae (Sauria) from the Late Cretaceous of the Gobi Desert. Acta Palaeontologica Polonica 29:51-81 [M. Carrano/M. Carrano/M. Carrano]
14221ETE K. Carpenter and K. Alf. 1994. Global distribution of dinosaur eggs, nests, and babies. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge 13-30 [M. Carrano/M. Carrano/M. Carrano]
10458 J. Dzik. 1975. Spiroboloid millipeds from the Late Cretaceous of the Gobi Desert, Mongolia. Palaeontologia Polonica 33:17-23 [M. Carrano/M. Carrano/M. Clapham]
9759 A. Elzanowski. 1974. Preliminary note on the palaeognathous bird from the Upper Cretaceous of Mongolia. Palaeontologia Polonica 29:103-109 [J. Alroy/E. Leckey/M. Carrano]
26124 A. Elzanowski. 1976. Palaeognathous bird from the Cretaceous of Central Asia. Nature 264:51-53 [J. Alroy/J. Alroy]
10467 A. Elzanowski. 1977. Skulls of Gobipteryx (Aves) from the Upper Cretaceous of Mongolia. Palaeontologia Polonica 37:153-165 [M. Carrano/M. Carrano/M. Carrano]
197345% 54920K. Gao and M. A. Norell. 2000. Taxonomic composition and systematics of Late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert. Bulletin of the American Museum of Natural History 249:1-118 [J. Alroy/E. Leckey/R. Benson]
37480 K. -Q. Gao and L. -H. Hou. 1995. Iguanians from the Upper Cretaceous Djadochta Formation, Gobi Desert, China. Journal of Vertebrate Paleontology 15(1):57-78 [R. Benson/R. Benson]
10453 R. Gradzinski and T. Jerzykiewicz. 1972. Additional geographical and geological data from the Polish-Mongolian Palaeontological Expeditions. Palaeontologia Polonica 27:17-306 [M. Carrano/M. Carrano/M. Carrano]
62116 R. Gradzinski and T. Jerzykiewicz. 1974. Sedimentation of the Barun Goyot Formation. Palaeontologica Polonica 30:111-146 [M. Uhen/F. Villegas-Garin]
59136 R. Gradzinski, Z. Kielan-Jaworowska, and T. Maryanska. 1977. Upper Cretaceous Djadokhta, Barun Goyot and Nemegt formations of Mongolia, including remarks on previous subdivisions. Acta Geologica Polonica 27(3):281-318 [M. Carrano/M. Carrano]
51934 T. Jerzykiewicz. 1998. Okavango Oasis, Kalahari Desert: a contemporary analogue for the Late Cretaceous vertebrate habitat of the Gobi Basin, Mongolia. Geoscience Canada 25(1):15-26 [M. Carrano/A. Murch]
77615% 37520Z. Kielan-Jaworowska. 1974. Multituberculate succession in the Late Cretaceous of the Gobi Desert (Mongolia). Palaeontologia Polonica 30:23-44 [J. Alroy/E. Leckey/E. Leckey]
10442 Z. Kielan-Jaworowska. 1975. Preliminary description of two new eutherian genera from the Late Cretaceous of Mongolia. Palaeontologia Polonica 33:5-15 [M. Carrano/M. Carrano/M. Carrano]
10462 Z. Kielan-Jaworowska. 1978. Evolution of the therian mammals in the Late Cretaceous of Asia. Part III. Postcranial skeleton in Zalambdalestidae. Palaeontologia Polonica 58:5-41 [M. Carrano/M. Carrano/M. Carrano]
39263 Z. Kielan-Jaworowska and B. A. Trofimov. 1980. Cranial morphology of the Cretaceous eutherian mammal Barunlestes. Acta Palaeontologica Polonica 25(2):167-185 [R. Benson/R. Benson]
18352ETE E. N. Kurochkin. 2000. Mesozoic birds of Mongolia and the former USSR. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 533-559 [M. Carrano/M. Carrano/M. Carrano]
9909 T. Maryanska. 1977. Ankylosauridae (Dinosauria) from Mongolia. Palaeontologia Polonica 37:85-151 [M. Carrano/M. Carrano/M. Carrano]
60485% 6700T. Maryanska and H. Osmólska. 1975. Protoceratopsidae (Dinosauria) of Asia. Palaeontologica Polonica 33:133-181 [M. Carrano/M. Carrano/M. Carrano]
14217ETE K. Mikhailov, K. Sabath, and S. Kurzanov. 1994. Eggs and nests from the Cretaceous of Mongolia. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge 88-115 [M. Carrano/M. Carrano/M. Carrano]
10452 M. Mlynarski. 1972. Zangerlia testudinimorpha n. gen., n. sp., a primitive land tortoise from the Upper Cretaceous of Mongolia. Palaeontologia Polonica 27:85-92 [M. Carrano/M. Carrano/M. Carrano]
41004 K. Sabath. 1991. Upper Cretaceous amniotic eggs from the Gobi Desert. Acta Palaeontologica Polonica 36(2):151-192 [M. Carrano/M. Carrano]
10451 A. Sulimski. 1972. Adamisaurus magnidentatus n. gen., n. sp. (Sauria) from the Upper Cretaceous of Mongolia. Palaeontologia Polonica 27:33-40 [M. Carrano/M. Carrano/M. Carrano]
10460 A. Sulimski. 1975. Macrocephalosauridae and Polyglyphanodontidae (Sauria) from the Late Cretaceous of Mongolia. Palaeontologia Polonica 33:25-102 [M. Carrano/M. Carrano/M. Carrano]
10444 A. Sulimski. 1984. A new Cretaceous scincomorph lizard from Mongolia. Palaeontologia Polonica 46:143-155 [M. Carrano/M. Carrano/M. Carrano]