Puesto Estancia La Costa: Santacrucian, Argentina
collected 2003-2009

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Panameriungulata - Adianthidae
Adianthus bucatus n. gen., n. sp. Ameghino 1891
Cifelli and Soria 1983 1 specimen
MACN A1812, dentary i2-m2, neotype
Mammalia - Panameriungulata - Proterotheriidae
Tetramerorhinus mixtum n. sp. Ameghino 1894
Cassini et al. 2012
synonym of Proterotherium australe
MACN-A 8970–71, cranium, mandible, ulna, femur, tibia, type
Tetramerorhinus mixtum Ameghino 1894
MacPhee et al. 2021
synonym of Proterotherium australe
MACN A 8970
Diadiaphorus robustus n. sp. Ameghino 1894
Cassini et al. 2012
MACN-A 9200–08, cranium and mandible
Diadiaphorus majusculus Ameghino 1887
    = Diadiaphorus majusculus Ameghino 1887
Cassini et al. 2012
MACN-A 2711–12, cranium and mandible; MACN-A 9180–82, cranium, mandible
Tetramerorhinus cingulatum Ameghino 1891
MacPhee et al. 2021
MACN A8666
Licaphrium floweri Ameghino 1887
Thoatherium minusculum Ameghino 1887
Mammalia - Panameriungulata - Macraucheniidae
Theosodon gracilis Ameghino 1891
Cassini et al. 2012
MACN-A 9297, mandible ("Corriguen Aike")
Theosodon lydekkeri n. sp. Ameghino 1887
Cassini et al. 2012
MACN-A 2487–90, cranium and mandible (type); MACN-A 9269–88, cranium and mandible
Mammalia - Primates - Homunculidae
Homunculus patagonicus Ameghino 1891
    = Homunculus patagonicus n. gen., n. sp. Ameghino 1891
Kay et al. 2012
MACN-A 5757, mandible with worn right i1-m3, left i1, p2-m3, neotype; 5758, right femur; 5759, ulna; 5760, partial radius; 5761, distal humerus (lost); 5968, skull fragment with p2-m3; CORD-PZ 1130, partial skull
Mammalia - Rodentia
Neoreomys australis Ameghino 1887
Mammalia - Rodentia - Eocardiidae
Eocardia fissa Ameghino 1891
Kay et al. 2012
recombined as Dicardia fissa
Mammalia - Rodentia
Eocardia montana Ameghino 1887
Mammalia - Rodentia - Acaremyidae
Acaremys murinus Ameghino 1887
Arnal and Vucetich 2015 1 specimen
MACN A 4143 ("Corrigen Aike")
Mammalia - Rodentia
Acarechimys constans (Ameghino 1887)
Arnal et al. 2017
MPM-PV 4223 MPM-PV 15002
Mammalia - Rodentia - Octodontidae
Sciamys sp. Ameghino 1887
Kay et al. 2012
Mammalia - Rodentia
Spaniomys modestus Ameghino 1887
Mammalia - Rodentia - Neoepiblemidae
Perimys sp. Ameghino 1887
Mammalia - Rodentia
Steiromys duplicatus Ameghino 1887
Candela et al. 2012
MACN-A 10053–54, skull and right dentary; MACN-A 10055–78, maxillary, dentary, postcrania (both "Corriguen Kaik")
Steiromys detentus Ameghino 1887
Kay et al. 2012
Mammalia - Mylodontidae
Nematherium sp. Ameghino 1887
Kay et al. 2012
Mammalia - Megalonychidae
Eucholaeops informal sp. 1 (Amegino 1887)
original and current combination Eucholoeops
Eucholaeops informal sp. 2 (Amegino 1887)
original and current combination Eucholoeops
Eucholoeops fronto Ameghino 1891
Kay et al. 2012
Eucholoeops ingens Ameghino 1887
Kay et al. 2012
Hapalops informal sp. 1 Ameghino 1887
Hapalops informal sp. 2 Ameghino 1887
Hyperleptus sp. Ameghino 1891
Kay et al. 2012
Mammalia - Cingulata - Chlamyphoridae
Proeutatus informal sp. 1 Ameghino 1891
    = Proeutatus oenophorus Ameghino 1887
Kay et al. 2012
recombined as Proeutatus oenophorum
Stenotatus patagonicus (Ameghino 1887)
Kay et al. 2012
Prozaedyus sp. (Ameghino 1891)
    = Prozaedyus proximus Ameghino 1887
Kay et al. 2012
Mammalia - Cingulata
Peltephilus pumilus Ameghino 1887
Kay et al. 2012
Peltephilus strepens Ameghino 1887
Mammalia - Cingulata - Propalaehoplophoridae
Propalaehoplophorus incisivus
Vizcaíno et al. 2012
MACN-A 7655, skull; 7656, skull
Propalaehoplophorus minus Ameghino 1891
Vizcaíno et al. 2012
MACN-A 4757, mandible
Propalaehoplophorus informal sp. 1 Ameghino 1887
    = Propalaehoplophorus australis Moreno 1882
Kay et al. 2012
original and current combination Hoplophorus australis
Cochlops muricatus Ameghino 1889
Kay et al. 2012
Mammalia
Adiastaltus habilis n. gen., n. sp. Ameghino 1893
McDonald et al. 2008 2 specimens
MACN A-7772, humerus, type; A-7773, ulna
Mammalia - Astrapotheria - Astrapotheriidae
Astrapotherium magnum (Owen 1853)
Mammalia - Notoungulata
Hegetotherium mirabile Ameghino 1887
Mammalia - Notoungulata - Interatheriidae
Interatherium robustum Ameghino 1891
synonym of Interatherium extensus
Protypotherium claudum Ameghino 1889
Fernández et al. 2023
Protypotherium praerutilum Ameghino 1887
Cassini et al. 2012
MACN-A 3920–21, cranium, mandible ("Corriguen Aike")
Protypotherium attenuatum Ameghino 1887
    = Protypotherium attenuatum Ameghino 1887
Cassini et al. 2012
synonym of Protypotherium praerutilum
MACN-A 3991, mandible, paratype of _Protypotherium leptocephalum_
Protypotherium australe (Moreno 1882)
Mammalia - Notoungulata - Homalodotheriidae
Homalodotherium sp. Huxley 1870
    = Homalodotherium cunninghami Flower 1884
Kay et al. 2012
Mammalia - Notoungulata - Toxodontidae
Nesodon sp. Owen 1846
    = Nesodon imbricatus Owen 1846
Kay et al. 2012
Nesodon imbricatus Owen 1846
Cassini et al. 2012
MACN-A 5145, mandible; MACN-A 774–775, cranium
Adinotherium sp. Ameghino 1887
    = Adinotherium ovinum Owen 1846
Kay et al. 2012
Adinotherium ferum n. sp. Ameghino 1887
Cassini et al. 2012
synonym of Adinotherium ovinum
MACN-A 5352–53, cranium, mandible, type
Mammalia - Microbiotheria - Microbiotheriidae
Microbiotherium tehuelchum Ameghino 1887
Marshall 1982 2 specimens
MACN 8504, dentary m2-4; 8505, skull with left M1-4
Mammalia - Paucituberculata - Palaeothentidae
Palaeothentes minutus Ameghino 1887
Palaeothentes lemoinei Ameghino 1887
Mammalia - Sparassodonta
Perathereutes pungens Ameghino 1891
Kay et al. 2012
Cladosictis patagonica Ameghino 1887
Marshall 1981 9 specimens
MACN 5927, cranium; MACN 5928, dentary i1-p1, p3-m2; MACN 5929, vertebrae and other bones; MACN 6280, maxilla C-M4; MACN 6281, skull fragment; MACN 6282, vertebrae; MACN 6283, vertebrae; MACN 6284, proximal femur; MACN 6285, fibula
Cladosictis lateralis n. sp. Ameghino 1894
Marshall 1981 2 specimens
synonym of Cladosictis patagonica
MACN 5950, partial skull, holotype; MACN 5951, distal humerus, said to be same individual as 5950
Sipalocyon gracilis Ameghino 1887
Amphiproviverra crassa n. sp. Ameghino 1894
Marshall 1981 2 specimens
synonym of Sipalocyon gracilis
MACN 5957, dentary p2-m4; MACN 5958, maxilla P1-M4; cotypes
Lycopsis torresi Cabrera 1927
Prevosti et al. 2012
Anatherium oxyrhynchus n. sp. Ameghino 1894
Marshall 1979 1 specimen
synonym of Lycopsis torresi
MACN 5930, dentary p2-3, m2-4, holotype
Prothylacynus patagonicus Ameghino 1891
    = Prothylacynus patagonicus n. gen., n. sp. Ameghino 1891
Marshall 1979
Holotype (one associated individual): MACN 706, mandible left i1-m4, right i1-c; MACN 707, maxillary M1-4; MACN 708–720, postcrania;
Prothylacynus brachyrhynchus n. sp. Ameghino 1894
Marshall 1979 1 specimen
synonym of Prothylacynus patagonicus
MACN 5926, dentary left c, p2, m1-4, right p2, m2-4
Napodonictis thylacynoides n. gen., n. sp. Ameghino 1894
Marshall 1979 7 specimens
synonym of Prothylacynus patagonicus
MACN 5931, skull; MACN 5932, dentary with teeth; MACN 5933–5937, partial skeleton; all are one individual and part of the holotype
Borhyaena tuberata Ameghino 1887
    = Borhyaena tuberata Ameghino 1887
Prevosti et al. 2012
MACN-A 9343 ("Corriguen Aike")
Arctodictis munizi Mercerat 1891
Prevosti et al. 2012
Mammalia - Meridiolestida
Necrolestes patagonensis Ameghino 1891
Kay et al. 2012
Reptilia - Theropoda - Phorusrhacidae
Psilopterus lemoinei (Moreno and Mercerat 1891)
Alvarenga and Holfing 2003
Phorusrhacinae indet. Ameghino 1899
Degrange et al. 2019
MPEF-PV 11355: thoracic vertebra
Phorusrhacos longissimus (Ameghino 1887)
Reptilia - Theropoda - Falconidae
Thegornis debilis Ameghino 1894
Degrange et al. 2012 1 specimen
BMNH-A 601, distal right tarsometatarsus
see common names

Geography
Country:Argentina State/province:Santa Cruz
Coordinates: 51.2° South, 69.1° West (view map)
Paleocoordinates:51.8° South, 63.4° West
Basis of coordinate:stated in text
Altitude:0 meters
Geographic resolution:local area
Time
Period:Neogene Epoch:Miocene
Stage:Burdigalian 10 m.y. bin:Cenozoic 5
Key time interval:Santacrucian
Age range of interval:17.40000 - 16.20000 m.y. ago
Stratigraphy
Formation:Santa Cruz Member:Estancia La Costa
Stratigraphic resolution:formation
Stratigraphy comments: Dates from three levels in the lower one-third of the Santa Cruz fossiliferous beds at Monte Observación, rangebetween 16.42 Ma and 16.18 Ma and between 16.56 Ma and 16.28 Ma at Monte León. The dates are not wholly concordant but permit the assignment of the bulk of the coastal Santacrucian faunas to the late–early Miocene
Lithology and environment
Primary lithology:green sandy claystone
Secondary lithology:tuffaceous silty sandstone
Includes fossils?Y
Includes fossils?Y
Lithology description: tuffaceous silty sands, andgreenish silty clays.
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:paleoecologic analysis
Collection dates:2003-2009
Metadata
Also known as:Corriguen Aike, Corriguen Kaik, Corriken Aike
Database number:142533
Authorizer:C. Jaramillo, J. Zijlstra, P. Mannion Enterer:J. Carrillo, J. Zijlstra, G. Varnham, M. Kouvari
Modifier:M. Kouvari Research group:vertebrate
Collections that are a subset of this one:176602, 176603, 176636, 176643
Created:2013-04-13 01:37:34 Last modified:2020-09-21 09:10:43
Access level:database members Released:2014-04-13 01:37:34
Creative Commons license:CC BY
Reference information

Primary reference:

46189. S. F. Vizcaíno, M. S. Bargo, R. F. Kay, R. A. Fariña, M. Di Giacomo, J. M. G. Perry, F. J. Prevosti, N. Toledo, G. H. Cassini and J. C. Fernicola. 2010. A baseline paleoecological study of the Santa Cruz Formation (late-early Miocene) at the Atlantic coast of Patagonia, Argentina. Palaeogeography, Palaeoclimatology, Palaeoecology 292(3-4):507-519 [C. Jaramillo/J. Carrillo/P. Wagner]

Secondary references:

58179 M. A. Abello, E. Ortiz-Jaureguizar, and A. M. Candela. 2012. Paleoecology of the Paucituberculata and Microbiotheria (Mammalia, Marsupialia) from the late Early Miocene of Argentina. In S. F. Vizcaíno, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 156-172 [J. Zijlstra/J. Zijlstra]
35669 H. M. F. Alvarenga and E. Holfing. 2003. Systematic revision of the Phorusrhacidae (Avies: Ralliformes). Papeis Avulsos de Zoologia 43(4):55-91 [M. Uhen/M. Uhen]
58126 M. Arnal and M. Vucetich. 2015. Revision of the fossil rodent Acaremys Ameghino, 1887 (Hystricognathi, Octodontoidea, Acaremyidae) from the Miocene of Patagonia (Argentina) and the description of a new acaremyid. Historical Biology 27(1):42-59 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
72683 M. Arnal, M. G. Vucetich, D. A. Croft, M. S. Bargo, J. C. Fernícola and S. F. Viscaino. 2017. Systematic revision and evolutionary history of Acarechimys Patterson in Kraglievich 1965 (Rodentia, Caviomorpha, Octodontoidea). Ameghiniana 54(3):307-330 [P. Mannion/M. Kouvari]
58225 A. M. Candela, L. L. Rasia, and M. E. Pérez. 2012. Paleobiology of Santacrucian caviomorph rodents: a morphofunctional approach. In S. F. Vizcaíno, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 287-305 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
58224 G. H. Cassini, E. Cerdeño, A. L. Villafañe and N. A. Muñoz. 2012. Paleobiology of Santacrucian native ungulates (Meridiungulata: Astrapotheria, Litoptera and Notoungulata). In S. F. Vizcaíno, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 243-286 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
58245 R. L. Cifelli and M. F. Soria. 1983. Systematics of the Adianthidae (Litopterna, Mammalia). American Museum Novitates 2771:1-25 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
72945 F. J. Degrange, D. Eddy, P. Puerta and J. Clarke. 2019. New skull remains of Phorusrhacos longissimus (Aves, Cariamiformes) from the Miocene of Argentina: implications for the morphology of Phorusrhacidae. Journal of Paleontology 93(6):1221-1233 [P. Mannion/G. Varnham]
58178 F. J. Degrange, J. I. Noriega, and J. I. Areta. 2012. Diversity and paleobiology of the Santacrucian birds. In S. F. Vizcaíno, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 138-155 [J. Zijlstra/J. Zijlstra]
85306 M. Fernández, J. C. Fernicola, and E. Cerdeño. 2023. Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early–Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae. Zoological Journal of the Linnean Society zlad043:1-28 [J. Carrillo/K. Pino]
58226 R. F. Kay, J. M. G. Perry, M. Malinzak, K. L. Allen, E. C. Kirk, J. M. Plavcan, and J. G. Fleagle. 2012. Paleobiology of Santacrucian primates. In S. F. Vizcaíno, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 306-330 [J. Zijlstra/J. Zijlstra]
58227 R. F. Kay, S. F. Vizcaíno, and M. S. Bargo. 2012. A review of the paleoenvironment and paleoecology of the Miocene Santa Cruz Formation. In S. F. Vizcaíno, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 331-365 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
81723 R. D. E. MacPhee, S. Hernández del Pino, A. Kramarz, A. M. Forasiepi, M. Bond and R. B. Sulser. 2021. Cranial morphology and phylogenetic relationships of Trigonostylops wortmani, an Eocene South American native ungulate. Bulletin of the American Museum of Natural History 449(1):1-183 [P. Mannion/G. Varnham]
57760 L. Marshall. 1982. Systematics of the South American marsupial family Microbiotheriidae. Fieldiana Geology 10:1-75 [J. Zijlstra/J. Zijlstra]
58324 L. G. Marshall. 1979. Review of the Prothylacyninae, an extinct subfamily of South American "dog-like" marsupials. Fieldiana Geology 3:1-50 [J. Zijlstra/J. Zijlstra]
58306 L. G. Marshall. 1981. Review of the Hathlyacyninae, an extinct subfamily of South American "dog-like" marsupials. Fieldiana Geology 7:1-120 [J. Zijlstra/J. Zijlstra]
58233 H. G. McDonald, S. F. Vizcaíno, and M. S. Bargo. 2008. Skeletal anatomy and the fossil history of the Vermilingua. In S. F. Vizcaíno, W. J. Loughry (eds.), The Biology of the Xenarthra 64-78 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
58202 F. J. Prevosti, A. M. Forasiepi, M. D. Ercoli and G. F. Turazzini. 2012. Paleoecology of the mammalian carnivores (Metatheria, Sparassodonta) of the Santa Cruz Formation (late Early Miocene). In S. F. Vizcaíno, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 173-193 [J. Zijlstra/J. Zijlstra/J. Zijlstra]
58206 S. F. Vizcaíno, J. C. Fernicola, and M. S. Bargo. 2012. Paleobiology of Santacrucian glyptodonts and armadillos (Xenarthra, Cingulata). In S. F. Vizcaí­no, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 194-215 [J. Zijlstra/J. Zijlstra/J. Zijlstra]