PPP# 2172 - Escudo de Veraguas Island [Escudo de Veraguas Fm] (Pliocene of Panama)

Also known as NMB Loc. 18671; 95-05-11; mislabelled as PPP# 2127 in Landau et al (2012)

Where: Bocas, Panama (9.1° N, 81.6° W: paleocoordinates 9.0° N, 81.1° W)

• coordinate stated in text

• outcrop-level geographic resolution

When: Escudo de Veraguas Formation (Bocas del Toro Group), Piacenzian (3.6 - 2.6 Ma)

• STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), not specified to Early or Late Pliocene; direct date extrapolated from Cheetham et al. (1999); age range also provided by Bybell (1999). STRATIGRAPHIC POSITION: From a height of 78.5 m in section 10 of Collins et al. (1999).

• bed-level stratigraphic resolution

Environment/lithology: deep-water; poorly lithified, burrowed, bioturbated, fine-grained, argillaceous sandstone and poorly lithified, burrowed, bioturbated, argillaceous siltstone

• ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
• GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, pervasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.

Size class: macrofossils

Collected by Jung et al in 1995; reposited in the BMNH, NMB, USNM

Collection methods: bulk, sieve,

• COLLECTORS:coll. P. Jung, A. Heitz, J. Todd et al., 2/22/95. REPOSITORY: Naturhistorisches Museum Basel (NMB).

Primary reference: L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 135885: authorized by Austin Hendy, entered by Austin Hendy on 12.11.2012

Creative Commons license: CC BY (attribution)

Taxonomic list

• COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
Bivalvia
 Arcida - Arcidae
Arca (Arca) sp.1 Linnaeus 1758 ark
Barbatia (Barbatia) sp.1 Gray 1842 clam
Anadara (Sectiarca) sp.1 Olsson 1961 ark
 Pectinida - Propeamussiidae
Cyclopecten sp.1 Verrill 1897 mud scallop
 Pectinida - Pectinoidae
Argopecten sp.1 Monterosato 1889 calico scallop
Spathochlamys sp.1 Waller 1993 scallop
 Carditida - Carditidae
Cardites sp.1 Link 1807 clam
 Pholadida - Corbulidae
"Bothrocorbula (Hexacorbula) sp." = Hexacorbula1, Caryocorbula sp.1, "Varicorbula sp." = Corbula (Varicorbula)1
"Bothrocorbula (Hexacorbula) sp." = Hexacorbula1 Olsson 1932 clam
Caryocorbula sp.1 Gardner 1926 clam
"Varicorbula sp." = Corbula (Varicorbula)1 Grant and Gale 1931 clam
 Cardiida - Chamidae
Chama sp.1 Linnaeus 1758 jewel box
 Cardiida - Veneridae
"Ventricolaria sp." = Globivenus1, Gouldia sp.1
"Ventricolaria sp." = Globivenus1 Coen 1934 venus clam
Gouldia sp.1 Adams 1847 venus clam
 Cardiida - Tellinidae
Eurytellina sp.1 Fischer 1887 tellin clam
 Cardiida - Semelidae
Abra sp.1 Lamarck 1818 abra
 Cardiida - Cardiidae
Laevicardium sp.1 Swainson 1840 cockle
 Nuculida - Nuculidae
"Leionucula sp." = Nucula (Leionucula)1 Quenstedt 1930 nut clam
Nucula (Lamellinucula) sp.1 Schenck 1944 nut clam
 Protobranchia - Nuculanidae
Costelloleda sp.1 Hertlein and Strong 1940 pointed nut clam
Saccella sp.1 Woodring 1925 pointed nut clam
Gastropoda
 Thecosomata - Cavoliniidae
Cavolinia sp.1 Avilgaard 1791 snail
Styliola sp.1 Gray 1850 snail
Clio sp.1 Linnaeus 1767 snail
Creseis sp.1 Rang 1828 snail
 Opisthobranchia - Bullidae
Bulla sp.1 Linnaeus 1758 snail
 Opisthobranchia - Retusidae
Volvulella (Volvulella) sp.1 Newton 1891 snail
 Opisthobranchia - Cylichnidae
Acteocina sp.1 Gray 1847 barrel-bubble
Cylichnella sp.1 Gabb 1873 snail
Scaphander sp.1 Monfort 1810 snail
 Opisthobranchia - Haminoeidae
"Atys (Aliculastrum) sp." = Aliculastrum1
"Atys (Aliculastrum) sp." = Aliculastrum1 Pilsbry 1896 snail
 Neogastropoda - Olividae
Olivella sp.1 Swainson 1831 olive snail
Olivella (Macgintiella) sp.1 Olsson 1956 olive snail
Olivella (Niteoliva) sp.1 Olsson 1956 olive snail
Olivella (Minioliva) sp.1 Olsson 1956 olive snail
 Neogastropoda -
"Ancilla (Eburna) sp." = Eburna1
"Ancilla (Eburna) sp." = Eburna1 Lamarck 1801 snail
 Neogastropoda - Columbellidae
Mitrella sp.1 Risso 1826 snail
Cotonopsis (Cotonopsis) sp.1 Olsson 1942 snail
Cosmioconcha sp.1 Dall 1913 snail
 Neogastropoda - Buccinidae
Metula sp.1 Adams and Adams 1853 true whelk
Antillophos (Antillophos) sp.1 Woodring 1928 true whelk
 Neogastropoda - Nassariidae
Nassarius sp.1 Duméril 1806 snail
 Neogastropoda - Fasciolariidae
Fusinus sp.1 Rafinesque 1815 snail
 Neogastropoda - Cancellariidae
Aphera cf. lindae2 Petuch 1987 snail
 Neogastropoda - Cystiscidae
Granulina sp.1 Jousseaume 1888 snail
Persicula sp.1 Schumacher 1817 snail
 Neogastropoda - Mitridae
Subcancilla sp.1 Olsson and Harbison 1953 mitre shell
 Neogastropoda - Marginellidae
Dentimargo sp.1 Cossmann 1899 margin shell
Prunum sp.1 Swainson 1840 margin shell
Volvarina sp.1 Hinds 1844 margin shell
 Neogastropoda - Volutidae
Voluta sp.1 Linnaeaus 1758 volute
 Neogastropoda - Muricidae
Siphonochelus (Pilsbrytyphis) sp.1 Woodring 1959 murex snail
Haustellum sp.1 Schumacher 1817 murex snail
 Neogastropoda - Mangeliidae
Vitricythara sp.1 Fargo 1953 snail
Saccharoturris sp.1 Woodring 1928 snail
Granoturris sp.1 Fargo 1953 snail
Ithycythara sp.1 Woodring 1928 snail
 Neogastropoda - Drilliidae
Leptadrillia sp.1 Woodring 1928 snail
Hindsiclava sp.1 Hertlein and Strong 1955 snail
Syntomodrillia sp.1 Woodring 1928 snail
 Neogastropoda - Turridae
Knefastia sp.1 Dall 1919 turrid
Miraclathurella sp.1 Woodring 1928 turrid
Lioglyphostoma sp.1 Woodring 1928 turrid
Globidrillia sp.1 Woodring 1928 turrid
Polystira sp.1 Woodring 1928 turrid
 Neogastropoda - Raphitomidae
Pleurotomella sp.1 Verrill 1872 snail
 Neogastropoda -
 Neogastropoda - Conidae
Conus sp.1 Linnaeus 1758 cone shell
 Neogastropoda - Terebridae
Strioterebrum sp.1 Sacco 1891 auger snail
 Stromboidea - Strombidae
Strombus (Strombus) sp.1 Linnaeus 1758 conch
 Vanikoroidea - Eulimidae
Niso sp.1 Risso 1826 snail
 Velutinoidea - Triviidae
Trivia sp.1 Broderip 1837 snail
 Naticoidea - Naticidae
Stigmaulax sp.1, "Natica (Naticarius) sp." = Naticarius1, Sinum sp.1, Polinices sp.1
Stigmaulax sp.1 Morch 1852 moon snail
"Natica (Naticarius) sp." = Naticarius1 Dumeril 1806 moon snail
Sinum sp.1 Röding 1798 moon snail
Polinices sp.1 Monfort 1810 moon snail
 Vermetoidea - Vermetidae
Serpulorbis sp.1 Sasso 1827 worm shell
 Xenophoroidea - Xenophoridae
Xenophora sp.1 Fischer von Waldheim 1807 carrier shell
Onustus sp.1 Swainson 1840 carrier shell
 Tonnoidea - Personidae
Distorsio clathrata1 Lamarck 1816 snail
 Rissooidea - Rissoidae
Zebina (Zebina) sp.1 Adams and Adams 1854 snail
Rissoina (Rissoina) sp.1 d'Orbigny 1840 snail
Folinia sp.1 Crosse 1868 snail
 Heterostropha - Acteonidae
Acteon sp.1 Montfort 1810 snail
 Heterostropha - Ringiculidae
 Pyramidelloidea - Pyramidellidae
Chemnitzia sp.1 d'Orbigny 1840 snail
"Pyrgiscus sp." = Turbonilla (Pyrgiscus)1 Philippi 1841 snail
Callolongchaeus sp.1 Dall 1903 snail
 Architectonicoidea - Architectonicidae
Architectonica sp.1 Röding 1798 sundial
 Fissurelloidea - Fissurellidae
Diodora sp.1 Gray 1821 snail
 Trochoidea - Trochidae
Calliostoma sp.1 Swainson 1840 top snail
Scaphopoda
 Dentaliida - Dentaliidae
Dentalium sp.1 Linnaeus 1758 tusk shell
 Gadilida - Gadilidae
Gadilopsis sp.1 Woodring 1925 tusk shell