Sample A, Titusville Site (1967 excavation) (Pleistocene of the United States)

Where: Pennsylvania (41.6° N, 79.6° W: paleocoordinates 41.6° N, 79.6° W)

• coordinate stated in text

• hand sample-level geographic resolution

When: Titusville Till Formation, Late/Upper Pleistocene (0.1 - 0.0 Ma)

Environment/lithology: mire or swamp; peat

• Like Zone I, analogues to the Zone III fauna occur today in forest–tundra environments of North America where mean July temperatures range from 10 to 13C.

Size class: mesofossils

Collected in 1967

Collection methods: quarrying, sieve

• All specimens, including those examined by Totten and Coope in 1967, were mounted onto micropaleontological slides and reposited in the Quaternary Entomology Laboratory at North Dakota State University.

Primary reference: S. Cong, A. C. Ashworth, D. P. Schwert and S. M. Totten. 1996. Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania. Quaternary Research 45:216-225 [M. Clapham/J. Karr/J. Karr]more details

Purpose of describing collection: paleoecologic analysis

PaleoDB collection 137209: authorized by Matthew Clapham, entered by Jered Karr on 12.12.2012, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

 Coleoptera - Helophoridae
Helophorus tuberculatus Gyllenhal 1808 water scavenger beetle
 Coleoptera - Staphylinidae
Acidota quadrata Zetterstedt 1838 rove beetle
Olophrum consimile Gyllenhal 1810 rove beetle
Olophrum rotundicolle Sahlberg 1817 rove beetle
"Holoboreaphilus nordenskioeldi" = Holoboreaphilus nordenskioldi Maklin 1878 rove beetle
Pycnoglypta sp. Thomson 1858 winter rove beetle
Pycnoglypta aptera or Pycnoglypta lurida
Pycnoglypta aptera Campbell 1983 winter rove beetle
 Coleoptera - Carabidae
Elaphrus lapponicus Gyllenhal 1810 ground beetle
"Carabus maeander" = Carabus (Homoeocarabus) maeander Fischer von Waldheim 1822 ground beetle
"Pterostichus patruelis" = Pterostichus (Phonias) patruelis Dejean 1831 ground beetle
"Bembidion fortestriatum" = Bembidion (Trepanedoris) fortestriatum Motschulsky 1845 ground beetle
"Trechus crassiscapus" = Trechus (Trechus) crassiscapus Lindroth 1955 ground beetle
Dyschirius melancholicus Putzeys 1867 ground beetle
Patrobus septentrionis Dejean 1828 ground beetle