Dolnice (Miocene of Czech Republic)

Also known as Dölitz, Delitz

Where: Karlovy Vary, Czech Republic (50.1° N, 12.4° E: paleocoordinates 50.4° N, 10.9° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: MN 4 (16.9 - 16.0 Ma)

• The Dolnice locality belongs to the upper part of the so called "coal seam beds" and is located immediately above the main lignite seam with its grey and brown mudstones (so called "Kapuzin-Tone"). The "coal seam beds" are overlain by the muddy to clayey Cypris beds which are characterized by ostracode bearing freshwater limestone lenses (Fejfar, 1974).

• member-level stratigraphic resolution

Environment/lithology: fluvial-lacustrine; calcareous siltstone and silty, sandy marl

• The locality is situated in the Cheb Basin at the intersection of the SW-NE trending Krušné Hory Graben (Ohře Graben, German: Eger-Graben) and the NW-SE trending Cheb-Domazlice Graben (see Reicherter et al., 2008, pp. 1233-1286 in McCann (ed.) The Geology of Central Europe, vol. 2).

Size classes: macrofossils, mesofossils

• The squamate remains reported by Roček (1984) and Szyndlar (1987) are in fact all isolated bones. The only articulated specimen is a partial pelvis of Lacerta sp. (DP FNSP 449).

•"[...] vertebrae and tooth bearing bones are prevailing, while frontals, parietals and pterygoids are scarce [...], and majority of other bones is completely lacking" (Roček, 1984).

Collection methods: bulk, sieve,

• This collection comprises fossils which have been collected during systematic excavations in the second half of the 20th century. All of the material is housed in the Department of Paleontology of the Faculty of Natural Sciences, Charles University, Prague (DP-FNSP).

Primary reference: Z. Szyndlar. 1987. Snakes from the Lower Miocene Locality of Dolnice (Czechoslovakia). Journal of Vertebrate Paleontology 7(1):55-71 [J. Mueller/T. Liebrecht]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 138666: authorized by Johannes Mueller, entered by Torsten Liebrecht on 25.01.2013, edited by Terri Cleary

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Squamata -
Squamata "indet. I"6 Oppel 1811 squamates
maxilla (DP FNSP 213)
Squamata "indet. III"6 Oppel 1811 squamates
premaxilla (DP FNSP 196)
 Squamata - Lacertidae
Lacertidae indet.6 Bonaparte 1831 squamates
dentary fragment (DP FNSP 3773)
Lacerta sp.6 Linnaeus 1758 squamates
37 dentaries (DP FNSP 1, 3, 5, 7, 11, 14-17, 25, 27, 28, 30, 330, 50, 51, 63, 73, 77, 82, 83, 94, 214, 236, 265, 514,3772, 3775, 3780, 3787, 3800, 3824, 3849, 3851, 3857, 3895, 4368), 13 maxillae (DP FNSP 24, 43, 52, 74, 81, 98, 124, 1473, 3795, 3843, 3852, 3860), 1 pterygoid (DP FNSP 4320), 1 pelvis (DP FNSP 449)
? Lacerta sp.6 Linnaeus 1758 squamates
specimens listed as Lacerta sp. but assigned tentatively to this taxon due to bad preservation: 35 dentaries (DP FNSP 12, 21, 23, 29, 38, 44, 47, 58, 62, 65, 68, 76, 88-90, 222, 242, 244, 245, 296, 518, 3778, 3784, 3789, 3790, 3810, 3822, 3827, 3828, 3830, 3834, 3835, 3876, 4305, 4309), 7 maxillae (DP FNSP 32, 70, 72, 3818, 3855, 3884, 4307)
Lacerta cf. viridis2 Laurenti 1768 European green lizard
Pb 02054, fused frontals; Pb 02055, fragment of left frontal; Pb 02024, fragment of posterior right portion of parietal
Miolacerta tenuis n. gen. n. sp.6 Roček 1984 squamates
Holotype: left dentary (DP FNSP 3785); paratypes: 2 dentaries (DP FNSP 3791, 4350); 14 referred specimens, mainly tooth bearing bones (DP FNSP 54, 55, 69, 75, 78, 130, 240, 243, 3811, 3847, 3850, 3868, 3873, 3910):
Amblyolacerta dolnicensis n. gen. n. sp.6 Roček 1984 squamates
Holotype: right maxilla (DP FNSP 4820); paratypes: 2 dentaries (DP FNSP 3809, 3837), 1 maxilla (DP FNSP 46); 3 referred specimens (DP FNSP 1485*, 3864, 4328; *"rather doubtful")
 Squamata - Anguidae
Anguidae indet.6 Gray 1825 squamates
trunk vertebrae: 8 of "morphotype II" (DP FNSP 1286, 1287, 1299, 1313, 1315, 1418, 1454, 3961), 9 of "morphotype III" (DP FNSP 1308, 1415, 1487, 3934, 3943, 3948, 4006, 4013, 4015), 2 of "morphotype IV" (DP FNSP 3951, 4017), 1 of "morphotype VI" (DP FNSP 1310); caudal vertebrae: 9 of "morphotype I" (DP FNSP 1302, 1333, 1424, 1446, 3919, 3925, 3972, 3977, 3989), 12 of "morphotype II" (DP FNSP 1241, 1303, 1305, 1335, 1363, 1449, 1452, 1453, 3918, 3952, 3995, 3996), 2 of "morphotype III" (DP FNSP 1251, 3939), 2 of "morphotype IV" (DP FNSP 1244, 1289), 3 of "morphotype V" (DP FNSP 354, and probably also 3933, 3950), 1 of "morphotype VI" (DP FNSP 4002), 4 of "morphotype VII" (DP FNSP 1212, 1215, 1439, 3923), 1 of "morphotype VIII" (DP FNSP 3945)
Anguidae "indet. I"6 Gray 1825 squamates
14 dentigerous bones (DP FNSP 4, 45, 67, 1395, 3801, 3820, 3823, 3858, 3866, 4308, 4312, "and probably also" 35, 3786, 3816)
Anguidae "indet. II"6 Gray 1825 squamates
1 maxilla (DP FNSP 93)
cf. Xestops sp.6 Cope 1873 squamates
8 dentaries (DP FNSP 18, 39, 48, 60, 3781, 3783, 3794, 3859), 9 maxillae (DP FNSP 129, 212, 219, 3813, 3819, 3821, 3831, 3914, 4306), 6 premaxillae (DP FNSP 194, 195, 345, 1552, 3814, 3878), 7 fragments of dentigerous bone (DP FNSP 37*, 96*, 224*,277, 1396*, 3829*, 3841*; * = "with certain doubts")
Pseudopus sp.6 Merrem 1820 legless lizard
2 bones with marginal dentition (DP FNSP 218, 896)
Pseudopus ahnikoviensis3 Klembara 2012 legless lizard
parietal (DP FNSP 1016, paratype, identified as Ophisaurus aff. spinari by Klembara, 1981), frontal (DP FNSP 1017, paratype, identified as Ophisaurus sp. by Klembara, 1981), 2 dentaries (DP FNSP 1030 and 1031, paratypes, identified as Pseudopus sp. by Klembara, 1981)
Ophisaurus "sp. I"6 Daudin 1803 glass lizard
dentary (DP FNSP 3844)
Ophisaurus "sp. II"6 Daudin 1803 glass lizard
3 dentaries (DP FNSP 84*, 1472, 3846; * = "probably")
cf. Ophisaurus sp.6 Daudin 1803 glass lizard
49 trunk vertebrae of "morphotype I" (DP FNSP 1187, 1209, 1210, 1234, 1240, 1242, 1246, 1292, 1293, 1306, 1309, 1314, 1409, 1410, 1412, 1416, 1417, 1420, 1423, 1425, 1426, 1429, 1442, 1445, 1455, 1456, 1460, 3928, 3941, 3942, 3944, 3946, 3949, 3955, 3958, 3960, 3963, 3965, 3968, 3973, 3992, 3997, 4001, 4007, 4010, 4015, 4019, 4024, 4027) and 4 trunk vertebrae of "morphotype V" (DP FNSP 1214, 1450, 3937, 4016)
"Anguis robustus n. sp." = Ophisaurus robustus4 Klembara 1979 glass lizard
Specimens listed as O. robustus in this paper. DP FNSP 1018 (holotype), parietal; 1020, prev. ID'd as O. spinari
Ophisaurus feifari n. sp.4 Klembara 1979 glass lizard
DP FNSP 1013 (holotype), parietal; FNSP 1501, parietal; 1023, parietal (previously ID'd as O. spinari)
Ophisaurus cf. spinari6 Klembara 1979 glass lizard
14 dentaries (DP FNSP 79, 80, 99, 121-123, 128, 246, 290, 3774, 3788, 3874, 3896, 3911); 6 maxillae (DP FNSP 9, 64, 117, 120, 125, 127), 4 pterygoids (DP FNSP 1512, 1518, 1550, 1558)
? Ophisaurus cf. spinari6 Klembara 1979 glass lizard
"perhaps" Ophisaurus cf. spinari: 3 maxillae (DP FNSP 19, 41, 3871), 1 pterygoid (DP FNSP 1551)
Ophisaurus spinari4 Klembara 1979 glass lizard
Parietals DP FNSP 1014, DP FNSP 1019, DP FNSP 1021, 1022, DP FNSP 1025, DP FNSP 1028
Ophisaurus holeci4 Klembara 2015 glass lizard
Parietals DP FNSP 1026, ?DP FNSP 1027, ?DP FNSP 1029
 Squamata - Chamaeleonidae
Chamaeleo caroliquarti n. sp.5 Moody and Roček 1980 chameleon
DP FNSP 101 (holotype), left dentary; 102, midsection of right dentary; 103, left maxillary; 104, right maxillary; 105, section of max with 2 teeth; 106, midsection left max; 107, anteriormost section of right dentary w 7 teeth; 108, ant section of right dentary w 14 teeth; 109, section of left max incl palatine process; 110, ant part of right dentary; 111, midsection left dentary; 112, posterior section left dentary. Add'l specimens from ref 43973: 5 fragments of unidentified dentigerous bone (DP FNSP 113, 279, 512, 3867, 3869); 1 partial right maxilla (DP FNSP 4316), 1 postorbital (DP FNSP 226)
Chamaeleo andrusovi n. sp.1 Čerňanský 2010 chameleon
Pb 02056 (holotype), nearly complete parietal but lacking the posterior portion; Right jugal (Pb 02057); left jugal (Pb 02058); left prefrontal (Pb 02059); postfrontal portion of the left postorbitofrontal (Pb 02050)
 Squamata - Viperidae
Viperidae indet. Oppel 1811 viper
4 vertebrae (DP FNSP 1158, 3929, 4032, 4529)
 Squamata - Boidae
Bavarioboa hermi7, "Bransateryx septentrionalis n. sp." = Falseryx petersbuchi
Bavarioboa hermi7 Szyndlar and Schleich 1993 boa
14 precaudal vertebrae (DP FNSP 1147, 1148, 1157-1159, 1167, 1489, 1490, 3921, 3922, 4005, 4084, 4533, 5270)
"Bransateryx septentrionalis n. sp." = Falseryx petersbuchi Szyndlar and Rage 2003 boa
a single posterior caudal vertebra (DP FNSP 4555 = holotype) and 6 further vertebrae (DP FNSP 3916, 3990, 4003, 4528, 4556, 4557)
 Squamata -
Colubroidea indet. Oppel 1811 snake
5 caudal vertebrae (DP FNSP 1211, 1427, 1444, 1447, 3971)
 Squamata - Colubridae
Colubridae indet. Oppel 1811 colubrid snake
11 precaudal vertebrae (DP FNSP 1155, 1374, 1432, 3938, 3959, 3964, 3966, 4200, 4231, 4530, 5207)
Palaeonatrix lehmani Rage and Roček 1983 colubrid snake
7 precaudal vertebrae (DP FNSP 933, 1408, 1434, 1437, 4096, 4553, 4554); the author does not say whether this material comes (at least in part) from the same stratigraphic level as the type material of Dolniceophis lehmani (see PBDB coll. 68711) or not
Texasophis bohemiacus n. sp. Szyndlar 1987 colubrid snake
3 trunk vertebrae (holotype: DP FNSP 1238; referred specimens: DP FNSP 1239, 3927)
? Texasophis bohemiacus Szyndlar 1987 colubrid snake
2 trunk vertebrae (DP FNSP 3947, 4025)
Coluber dolnicensis n. sp. Szyndlar 1987 racer
2 trunk vertebrae (holotype: DP NFSP 4558; referred specimen 4559), 1 compound bone (DP FNSP 310)
Neonatrix nova n. sp. Szyndlar 1987 colubrid snake
63 precaudal vertebrae (holotype: DP FNSP 5197; referred specimens: DP FNSP 931, 932, 934-937, 1171, 1237, 1245, 1253, 1311, 1312, 1414, 1419, 1430, 1433, 1440, 1443, 1451, 1458, 3962, 3967, 3979, 3983, 3987, 3988, 4026, 4029, 4030, 4075, 4107, 4201, 4511, 4526, 4532, 5156-5159, 5162, 5164, 5169, 5171-5173, 5175, 5176, 5182, 5186-5188, 5192, 5194-5196, 5198, 5200, 5202, 5204, 5214, 5221, 5224)
 Squamata - Crotalidae
Vipera antiqua n. sp. Szyndlar 1987 pit viper
6 precaudal vertebrae (holotype: DP FNSP 4538; referred specimens: DP FNSP 938, 1114, 1213, 4014, 4057)
Vipera platyspondyla n. sp. Szyndlar 1987 pit viper
8 precaudal vertebrae (holotype: DP FNSP 940; referred specimens: 939, 1421, 1431, 1435, 3926, 4004, 4008)
 Squamata -
Scolecophidia indet. snake
a single broken vertebra (DP FNSP 1316)
 Squamata - Amphisbaenidae
"Omoiotyphlops gracilis n. sp." = Blanus gracilis6, "? Blanus gracilis" = Blanus gracilis8
"Omoiotyphlops gracilis n. sp." = Blanus gracilis6 Roček 1984 worm lizard
Holotype: right dentary (DP FNSP 3771); paratypes: 2 left dentaries (DP FNSP 217, 3739) and 3 trunk vertebrae (DP FNSP 1317, 1336, 1364); referred material: 35 vertebrae (DP FNSP 1235, 1247, 1249, 1252, 1254, 1255, 1288, 1291, 1294-1296, 1298, 1300-1302, 1304, 1319, 1334, 1337, 1389, 1401, 1404, 1405, 1428, 1448, 1457, 3920, 3953, 3976, 3985, 3993, 3994, 3999)
"? Blanus gracilis" = Blanus gracilis8 Roček 1984 worm lizard
premaxilla (DP FNSP 317)
 Squamata - Scincidae
cf. Scincidae indet.6 Gray 1825 skink
2 maxillae (DP FNSP 59 "and perhaps" 61)
 Squamata - Cordylidae
Palaeocordylus bohemicus n. gen. n. sp.6 Roček 1984 spinytail lizard
holotype: left dentary (DP FNSP 97); paratypes: 8 dentaries (DP FNSP 20, 57, 100, 119, 216, 294, 3798, 3804), 3 maxillae (DP FNSP 71, 86, 3802), 1 premaxilla (DP FNSP 114); referred specimens: 13 dentaries (DP FNSP 22, 49, 56, 85, 87, 3776, 3803, 3806, 3836, 3839, 3842, 38, 45, 3854), 2 maxillae (DP FNSP 3796, 3805)
? Palaeocordylus bohemicus6 Roček 1984 spinytail lizard
specimens listed as P. bohemicus but "referred to this form with certain doubts" due to bad preservation: 9 dentaries (DP FNSP 40, 92, 268, 3779, 3815, 3840, 3848, 3856, 4350), 3 maxillae (DP FNSP 126, 232, 3792), 1 unidentified tooth bearing bone (DP FNSP 91)