PBDB

Shanwang, Linqu County, collection K0 (Shandong Geological Museum) (Miocene of China)

Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)

• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone

• Two minor volcanic faulted basins, the Linqu and Shanwang Basins, are present in the study area. The former lies near the county town of Linqu, and the latter in the Shanwang National Geopark, east of this town. The faulted basins are the result of intense Mesozoic-Cenozoic tectonic extension and rifting; the development of these basins was closely related in time with the nearby NE-SW and NW-SE trending faults in the Yishu Fault Zone.

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Shandong Geological Museum. This collection contains specimens labeled with three-digit numbers prefaced by K0.

Primary reference: J. F. Zhang. 1989. Miocene insects from Shanwang of Shandong, China and their bearing on palaeoenvironment. Proceedings of International Symposium on Pacific Neogene Continental and Marine Events 149-156 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 138944: authorized by Matthew Clapham, entered by Matthew Clapham on 03.02.2013

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Hemiptera - Cicadidae	"Meimuna miocenica n. sp." = Cryptotympana miocenica², "Meimuna incasa n. sp." = Cryptotympana miocenica¹ "Meimuna miocenica n. sp." = Cryptotympana miocenica² Zhang and Zhang 1990 cicada K0003 "Meimuna incasa n. sp." = Cryptotympana miocenica¹ Zhang and Zhang 1990 cicada K0431
Hemiptera - Coreidae	Plinachtus fossilis n. sp.², Cloresmus ambimodestus n. sp.² Plinachtus fossilis n. sp.² Zhang and Zhang 1990 leaf-footed bug K0112 Cloresmus ambimodestus n. sp.² Zhang and Zhang 1990 leaf-footed bug K0083
Hemiptera - Lygaeidae	Lygaeus daulus n. sp.¹ Lygaeus daulus n. sp.¹ Zhang et al. 1994 milkweed bug K0207
Hemiptera - Pentatomidae	Carbula aff. crassiventris² Carbula aff. crassiventris² Dallas 1849 stink bug K0008
Hemiptera - Reduviidae	Reduvius piceus n. sp.¹, Reduvius shandongianus n. sp.², Reduvius diatomus n. sp.² Reduvius piceus n. sp.¹ Zhang et al. 1994 assassin bug K0331 Reduvius shandongianus n. sp.² Zhang and Zhang 1990 assassin bug K0152, 0129 Reduvius diatomus n. sp.² Zhang and Zhang 1990 assassin bug K0136
Hemiptera - Nepidae	Ranatra dormientis¹ Ranatra dormientis¹ Zhang et al. 1994 true bug K0378
Hemiptera - Belostomatidae	Sphaerodema microcephalum n. sp.¹ Sphaerodema microcephalum n. sp.¹ Zhang et al. 1994 giant water bug K0359
Blattodea - Blattoidae	Periplaneta sphodra n. sp.¹ Periplaneta sphodra n. sp.¹ Zhang et al. 1994 cockroach K0387
Hymenoptera - Vespidae	Harmorhynchium reflexum n. gen. n. sp.¹, Provespa nobilis n. sp.¹, Vespa ciliata n. sp.¹ Harmorhynchium reflexum n. gen. n. sp.¹ Zhang et al. 1994 potter wasp K0367 Provespa nobilis n. sp.¹ Zhang et al. 1994 wasp K0380 Vespa ciliata n. sp.¹ Zhang et al. 1994 wasp K0371
Hymenoptera - Tiphiidae	Tiphia dimidiata n. sp.¹ Tiphia dimidiata n. sp.¹ Zhang et al. 1994 wasp K0372
Hymenoptera - Formicidae	Formica flavifemoralis n. sp.¹, Camponotus pictus n. sp.¹, Dolichoderus luridivenosus n. sp.¹, Liometopum lubricum n. sp.¹, Alloiomma differentialis n. sp.¹ Formica flavifemoralis n. sp.¹ Zhang et al. 1994 ant K0267 Camponotus pictus n. sp.¹ Zhang et al. 1994 ant K0174 Dolichoderus luridivenosus n. sp.¹ Zhang et al. 1994 ant K0231 Liometopum lubricum n. sp.¹ Zhang et al. 1994 ant K0295 Alloiomma differentialis n. sp.¹ Zhang et al. 1994 ant K0288
Hymenoptera - Megachilidae	Anthidium basalticum n. sp. Anthidium basalticum n. sp. Zhang 1989 mason bee K0138
Hymenoptera - Apidae	Xylocopa obata n. sp.¹, Bombus anacolus n. sp.¹ Xylocopa obata n. sp.¹ Zhang et al. 1994 large carpenter bee K0390 Bombus anacolus n. sp.¹ Zhang et al. 1994 bumble bee K0347
Hymenoptera - Ichneumonidae	Miolyta volcania n. gen. n. sp. Miolyta volcania n. gen. n. sp. Zhang 1989 ichneumon wasp K0147
Hymenoptera - Cimbicidae	Abia cf. lonicerae¹, "Clavellaria shandongensis" = Abia shandongensis¹, Sinocimbex silacea n. sp.¹ Abia cf. lonicerae¹ Linnaeus 1758 cimbicid sawfly K0274 "Clavellaria shandongensis" = Abia shandongensis¹ Hong and Wang 1985 cimbicid sawfly K0644 Sinocimbex silacea n. sp.¹ Zhang et al. 1994 cimbicid sawfly K0271
Hymenoptera - Tenthredinidae	Tenthredo petrae n. sp.³, Tenthredo miocenica n. sp.³, Driocampus shanwanganus n. gen. n. sp.³ Tenthredo petrae n. sp.³ Zhang and Zhang 1990 wood wasp K0115 Tenthredo miocenica n. sp.³ Zhang and Zhang 1990 wood wasp K0130 Driocampus shanwanganus n. gen. n. sp.³ Zhang and Zhang 1990 wood wasp K0057
Coleoptera - Buprestidae	Illolampra ampulla n. gen. n. sp.¹, Illolampra phlegma n. sp.¹, Eolampra gorgia n. sp.¹ Illolampra ampulla n. gen. n. sp.¹ Zhang et al. 1994 jewel beetle K0051 Illolampra phlegma n. sp.¹ Zhang et al. 1994 jewel beetle K0361 Eolampra gorgia n. sp.¹ Zhang et al. 1994 jewel beetle K0215
Coleoptera - Dascillidae	Lyprodascillus peritelus n. sp.¹, Dascillus musculus n. sp.¹ Lyprodascillus peritelus n. sp.¹ Zhang et al. 1994 soft bodied plant beetle K0208 Dascillus musculus n. sp.¹ Zhang et al. 1994 soft bodied plant beetle K0343
Coleoptera - Cantharidae	"Curticantharis trapezialis n. sp." = Themus (Themus) trapezialis¹ "Curticantharis trapezialis n. sp." = Themus (Themus) trapezialis¹ Zhang et al. 1994 soldier beetle K0349
Coleoptera - Elateridae	Corymbites euprepes n. sp.¹ Corymbites euprepes n. sp.¹ Zhang et al. 1994 click beetle K0253
Coleoptera - Cleridae	Titanoclerus shanwangianus n. gen. n. sp. Titanoclerus shanwangianus n. gen. n. sp. Zhang 1989 checkered beetle K0126
Coleoptera - Cerambycidae	Cerambycidae indet.¹, Mesocacia strumulosa n. sp.¹, Zoodes torridus n. sp.¹, Glischra aestivalis n. gen. n. sp.¹, Stromatium antiquum n. sp.¹, Eutetrapha terenia n. sp.¹, Mesosa longiuscula n. sp.¹, Podozoodes campalus n. gen. n. sp.¹, Paraphilus rejectus n. sp.¹, Perimesosa dissita n. sp.¹, Hornia penisularis n. gen. n. sp.¹ Cerambycidae indet.¹ Latreille 1802 longhorned beetle K0277 Mesocacia strumulosa n. sp.¹ Zhang et al. 1994 longhorned beetle K0229 Zoodes torridus n. sp.¹ Zhang et al. 1994 longhorned beetle K0366 Glischra aestivalis n. gen. n. sp.¹ Zhang et al. 1994 longhorned beetle K0224, 0259 Stromatium antiquum n. sp.¹ Zhang et al. 1994 longhorned beetle K0004 Eutetrapha terenia n. sp.¹ Zhang et al. 1994 longhorned beetle K0297 Mesosa longiuscula n. sp.¹ Zhang et al. 1994 longhorned beetle K0206 Podozoodes campalus n. gen. n. sp.¹ Zhang et al. 1994 longhorned beetle K0293 Paraphilus rejectus n. sp.¹ Zhang et al. 1994 longhorned beetle K0355 Perimesosa dissita n. sp.¹ Zhang et al. 1994 longhorned beetle K0249 Hornia penisularis n. gen. n. sp.¹ Zhang et al. 1994 longhorned beetle K0081
Coleoptera - Scarabaeidae	Adoretus rhinus n. sp.¹, Holotrichia cressona n. sp.¹, Petulantis yimengensis n. gen. n. sp.¹, Anomala furva¹, Anomala brachytarsia n. sp.¹, Anomala ursa n. sp.¹, Anomala endoxa n. sp.¹, Anomala orcina n. sp.¹, "Dolichopoda extumida n. gen. n. sp." = Zhangsunia extumida¹ Adoretus rhinus n. sp.¹ Zhang et al. 1994 scarab beetle K0168 Holotrichia cressona n. sp.¹ Zhang et al. 1994 scarab beetle K0056/0085 Petulantis yimengensis n. gen. n. sp.¹ Zhang et al. 1994 scarab beetle K0286 Anomala furva¹ Hong 1983 scarab beetle K0049 Anomala brachytarsia n. sp.¹ Zhang et al. 1994 scarab beetle K0220 Anomala ursa n. sp.¹ Zhang et al. 1994 scarab beetle K0377 Anomala endoxa n. sp.¹ Zhang et al. 1994 scarab beetle K0305 Anomala orcina n. sp.¹ Zhang et al. 1994 scarab beetle K0187 "Dolichopoda extumida n. gen. n. sp." = Zhangsunia extumida¹ Zhang et al. 1994 scarab beetle K0299
Coleoptera - Carabidae	Rhopalochlaenius phalanthus n. gen. n. sp.¹ Rhopalochlaenius phalanthus n. gen. n. sp.¹ Zhang et al. 1994 ground beetle K0200
Diptera - Asilidae	Promachus paucinervis n. sp.¹ Promachus paucinervis n. sp.¹ Zhang et al. 1994 robber fly K0292
Diptera - Bibionidae	Plecia capitata n. sp.¹ Plecia capitata n. sp.¹ Zhang et al. 1994 love bug K0266
Diptera - Tipulidae	Tipula corollata n. sp.¹ Tipula corollata n. sp.¹ Zhang et al. 1994 crane fly K0381
Odonata - Aeshnidae	"Mediaeschna lucida" = Epiaeschna lucida¹ "Mediaeschna lucida" = Epiaeschna lucida¹ Zhang 1989 darner dragonfly K0001
Arachnida
Araneae - Araneidae	"Testudinaria papposa n. gen. n. sp." = Testudinaroides papposa¹ "Testudinaria papposa n. gen. n. sp." = Testudinaroides papposa¹ Zhang et al. 1994 orb-weaving spider K0099

¹Zhang et al. 1994; ²Zhang and Zhang 1990; ³Zhang and Zhang 1990