Ravin de la Pluie (Miocene of Greece)

Also known as RPL

Where: Greece (40.8° N, 22.8° E: paleocoordinates 40.4° N, 22.4° E)

• coordinate estimated from map

When: MN10 mammal zone, Nea Messimvria Formation, Vallesian (11.6 - 8.7 Ma)

• late Miocene (Vallesian Mammal Age, Zone MN 10, correlated with chron C4AR.1N, 9.069–9.149 Ma based on the combination of the evolutionary grade of the mammalian fauna and paleomagnetostratigraphy

• formation-level stratigraphic resolution

Environment/lithology: terrestrial; sandstone and gravel

• sands, gravels, loose or hard conglomerates and red clay

Size class: macrofossils

Collected by de Bonis, Melentis, Koufos

Collection methods: quarrying, ,

• LGPUT, Thessaloniki

Primary reference: G. Koufos. 2012. A new protictithere from the late Miocene hominoid locality Ravin de la Pluie of Axios Valley (Macedonia, Greece). Palaeontologische Zeitschrift 86:219-229 [E. Vlachos/E. Vlachos/M. Uhen]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 191070: authorized by Evangelos Vlachos, entered by Evangelos Vlachos on 08.01.2018

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Reptilia
 Squamata - Colubridae
"Colubrinae" indet.3 Oppel 1811 colubrid snake
A series of incomplete vertebrae embedded in matrix (RPl 302)
 Squamata - Elapidae
Naja cf. romani3 Hoffstetter 1939 cobra
Two precloacal vertebrae (RPl 300–301)
 Squamata - Varanidae
Varanus sp.3 Merrem 1820 monitor lizard
Two presacral vertebrae (RPl 297–298) (Fig. 2); a humerus (RPl 295) (Fig. 3a, b, d); a tibia (RPl 296) (Fig. 3e, f).
 Squamata - Anguidae
? Anguinae indet.3 Gray 1825 squamates
One caudal vertebra (RPl 299)
 Testudines - Testudinidae
Testudo hellenica n. sp.2 Garcia et al. 2020 turtle
LGPUT RPI-216, a nearly complete carapace with plastron
Mammalia
 Proboscidea - Deinotheriidae
Deinotherium giganteum Kaup 1829 deinothere
 Proboscidea - Choerolophodontidae
Choerolophodon pentelici1 Gaudry and Lartet 1856 proboscidean
 Carnivora - Mustelidae
Eomellivora wimani Zdansky 1924 ratel
 Carnivora - Hyaenidae
"Protictitherium aff. intermedium" = Protoviverrops intermedium Schmidt-Kittler 1976 hyaena
Protictitherium thessalonikensis n. sp. Koufos 2012 hyaena
Adcrocuta eximia1 Roth and Wagner 1854 hyaena
subspecies leptorhyncha
Plioviverrops orbignyi1 Gaudry and Lartet 1856 hyaena
Hyaenictis sp. Gaudry 1861 hyaena
 Carnivora - Felidae
Metailurus parvulus Hensel 1862 cat
 Perissodactyla - Equidae
Hipparion macedonicum1 Koufos 1984 hipparionine horse
 Perissodactyla - Rhinocerotidae
Rhinocerotidae indet.1 Gray 1821 rhinoceros
 Artiodactyla - Giraffidae
Bohlinia cf. attica1 Lydekker 1886 giraffe
Palaeogiraffa major Bonis and Bouvrain 2003 giraffe
Palaeotragus rouenii1 Gaudry 1861 giraffe
 Artiodactyla - Bovidae
 Theriamorpha - Erinaceidae
"Palerinaceus sp." = Amphechinus
"Palerinaceus sp." = Amphechinus Aymard 1850 hedgehog
 Rodentia - Muridae
Progonomys cathalai1 Schaub 1938 mouse
 Rodentia - Sciuridae
Spermophilinus sp. de Bruijn and Mein 1968 squirrel
 Primates - Hominidae
Ouranopithecus macedoniensis1 de Bonis and Melentis 1977 ape