Los Mansuetos (Miocene of Spain)

Also known as LM

Where: Aragon, Spain (40.3° N, 2.6° W: paleocoordinates 40.5° N, 3.2° W)

• coordinate stated in text

When: Alfambra Formation, Turolian (8.7 - 5.3 Ma)

• Middle Turolian, MN12. late Miocene. LM is above LM2 in the stratigraphic column.


• 'A thick succession of red clays with some sand and conglomerate beds of the Peral Formation is followed by the Alfambra formation. The base of the Alfambra formation is a 2.5m thick bed of limestone mottled with gypsum. This is followed by 7.5m of brown-reddish clays and 7m of limy marl with abundant gypsum, 3m of calcareous gypsum and 0.6m of limestone, 15m of reddish sandy clay with some sand and conglomerate beds, 12.8m of gypsum and limestone with some clayey and marly intervals. One of these intervals corresponds to the Los Mansuetos 2 locality, it is a 0.3m thick lens of marl with pebbles of Neogene rocks, bone fragments and molluscs. Then follows a marly interval of 5.5m with on top a 40cm thick level of unsorted Neogene rock pebbles, bone fragments, debris of molluscs and plant remains, representing the classical locality of Los Mansuetos (LM). The section continues with about 43m of gypsum, limestones and poorly exposed clays.'

Size classes: macrofossils, mesofossils

Collection methods: bulk, chemical, sieve,

• material held in the collections of Geologisch Instituut, Rijksuniversiteit Utrecht.

Primary reference: H. de Bruijn and P. Mein. 1968. On the mammalian fauna of the Hipparion-Beds in the Calatayud-Teruel Basin (Prov. Zaragoza, Spain) Part V. The Sciurinae. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen. Series B: Palaeontology, Geology, Physics, Chemistry, Anthropology 71(1):73-90 [J. Alroy/E. Leckey/L. van den Hoek Ostende]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 35339: authorized by Alan Turner, entered by Hannah O'Regan on 02.12.2003, edited by Lars van den Hoek Ostende

Creative Commons license: CC BY (attribution)

Taxonomic list

• Los Mansuetos is the type site for the Turolian, and contains both small and large mammal remains.
 Rodentia - Muridae
Parapodemus gaudryi5, "Parapodemus adroveri n. sp." = Occitanomys (Rhodomys) adroveri5, "Valerymys turoliensis n. sp." = Huerzelerimys turoliensis5
Parapodemus gaudryi5 Dames 1883 mouse
Parapodemus gaudryi barbarae n. subsp.
"Parapodemus adroveri n. sp." = Occitanomys (Rhodomys) adroveri5 Thaler 1966 mouse
"Valerymys turoliensis n. sp." = Huerzelerimys turoliensis5 Michaux 1969 mouse
 Rodentia - Cricetidae
Ruscinomys schaubi5, "Kowalskia fahlbuschi" = Neocricetodon fahlbuschi5
"Kowalskia fahlbuschi" = Neocricetodon fahlbuschi5 Bachmayer and Wilson 1970 mouse
 Rodentia - Sciuridae
 Rodentia - Gliridae
Eliomys truci5 Mein and Michaux 1970 dormouse
 Rodentia - Castoridae
 Lipotyphla - Talpidae
"Dibolia turolense" = Archaeodesmana turolense3
"Dibolia turolense" = Archaeodesmana turolense3 Rümke 1985 mole
 Carnivora - Mustelidae
Baranogale adroveri2, "Enhydriodon lluecai" = Paludolutra lluecai1
Baranogale adroveri2 mustelid carnivore
"Enhydriodon lluecai" = Paludolutra lluecai1 Villalta Comella and Crusafont Pairó 1945 otter
 Carnivora -
Adcrocuta eximia2 Roth and Wagner 1854 carnivoran
 Carnivora - Canidae
Canis cipio2 canine
 Carnivora - Felidae
Metailurus parvulus2 Hensel 1862 cat
1 lower left P4.
Machairodus giganteus2 Wagner 1848 cat
upper teeth: 1 canine, P4 and P3 fragments. Lower teeth: 1 canine, 1 P3 fragment.
 Carnivora - Hyaenidae
Lycyaena sp.2 Hensel 1863 hyaena