Rians (Eocene of France)

Where: Var, France (43.6° N, 5.8° E: paleocoordinates 39.1° N, 3.0° E)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: MP7 mammal zone, Lignites de Soissonais Formation, Early/Lower Eocene (55.8 - 48.6 Ma)

• early Sparnacian

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size class: mesofossils

Reposited in the MNHN

Primary reference: D. E. Russell, M. Godinot, P. Louis and D. E. Savage. 1979. Apatotheria (Mammalia) de l'Eocène inférieur de France et de Belgique. Bulletin du Muséum National d'Histoire Naturelle, section C: Science de la Terre 1(3):203-243 [M. Carrano/M. Carrano/J. Alroy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 35538: authorized by Matthew Carrano, entered by Matthew Carrano on 19.12.2003, edited by John Alroy, K. Christopher Beard, Philip Mannion, David Nicholson and Terri Cleary

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Testudines -
Francellia salouagmirae n. gen. n. sp.5
Francellia salouagmirae n. gen. n. sp.5 turtle
MNHN RI 451 carapace and plastron
 Testudines - Podocnemidae
Neochelys arenarum n. sp.6 Broin 1977 sideneck turtle
Type MNHN.F.RI 6
 Squamata - Anguidae
Melanosaurini indet.1 Sullivan 1979 lizard
incomplete parietal
 Squamata - Amphisbaenidae
Amphisbaenidae indet.1 Gray 1865 worm lizard
four dorsal vertebrae
Mammalia
 Cimolesta - Palaeoryctidae
Palaeoryctidae indet.3 Winge 1917 eutherian
RI 343 and 355
 Leptictida - Leptictidae
? Leptictidae indet.3 Gill 1872 eutherian
RI 169
 Primates -
Donrussellia provincialis n. sp.4 Godinot 1978 tarsier
RI 170 - type
 Primates - Adapidae
Adapidae indet.3 Trouessart 1879 primate
RI 278 and 333
Pelycodus aff. eppsi3 Cooper 1932 primate
RI 179, 328 and 331
 Primates - Notharctidae
cf. Agerinia sp.4 Crusafont-Pairó and Golpe-Posse 1973 primate
 Primates - Paromomyidae
"Phenacolemur cf. fuscus" = Arcius fuscus3
"Phenacolemur cf. fuscus" = Arcius fuscus3 Russell et al. 1967 primate
RI 225
 Rodentia - Ischyromyidae
Microparamys cf. chandoni3 Hartenberger 1971 rodent
RI 315 and 366
Paramys cf. pourcyensis3 Michaux 1964 rodent
RI 199, 185 and 201
? Ailuravinae indet.3 Michaux 1968 rodent
RI 325
 Condylarthra - Hyopsodontidae
Hyopsodus itinerans n. sp.2 Godinot 1978 condylarth
RI 279 - type
 Perissodactyla -
? Paschatherium sp.3 Russell 1963 odd-toed ungulate
RI 421, 312 and 250
Paschatherium russelli n. sp.2 Godinot 1978 odd-toed ungulate
RI 153 - type
Paschatherium cf. dolloi3 Teilhard de Chardin 1927 odd-toed ungulate
RI 154, 155, 408, 330, 263, 231, 234, 259, 268, 397, 295, 299, 354, 417, 270, 288, 340, 380, 412, 156, 233, 386, 391, 241, ? 292, 374 and 416
"Hyracotherium aff. cuniculus" = Cymbalophus cuniculus3 Owen 1842 odd-toed ungulate
RI 136-139, 145, 183 and 308
 Tribosphenida - Nyctitheriidae
? Nyctitheriidae indet.3 Simpson 1928 placental
GOT 680-1
cf. Leptacodon sp.3 Matthew and Granger 1921 placental
RI 218, 219 and 382
 Chiroptera - Archaeonycteridae
cf. Archaeonycteris sp.3 Revilliod 1917 bat
RI 349, 361 and 334
 Carnivora - Miacidae
? Miacidae "sp. 1"3 Cope 1880 carnivoran
? Miacidae "sp. 2"3 Cope 1880 carnivoran
RI 364
 Hyaenodonta - Hyaenodontidae
Galecyon gallus n. sp.8, Didelphodus cf. absarokae3, "Proviverra eisenmanni n. sp." = Eoproviverra eisenmanni3
Galecyon gallus n. sp.8 Solé et al. 2013 eutherian
RI 275 and RI 276
Didelphodus cf. absarokae3 Cope 1881 eutherian
RI 202, 205 and 402
"Proviverra eisenmanni n. sp." = Eoproviverra eisenmanni3 Godinot 1981 eutherian
MNHN.F.RI400 - type; referred specimens: MNHN.F.RI203, 204, 362 and 401
 Theriiformes - Peradectidae
Peradectes louisi3 Crochet 1979 metatherian
RI 342, 420, 221, 226, 244 and 252
 Marsupialia - Herpetotheriidae
Peratherium matronense3 Crochet 1979 marsupial
R1 392 and 378
Amphiperatherium aff. brabantense3 Crochet 1979 marsupial
RI 379 and 411
Amphiperatherium cf. maximum3 Crochet 1979 marsupial
RI 220, 296 and 368
Amphiperatherium goethei3 Crochet 1979 marsupial
RI 385
 Artiodactyla - Diacodexeidae
? Diacodexis sp.3 Cope 1882 even-toed ungulate
RI 403
Diacodexis gazini n. sp.2 Godinot 1978 even-toed ungulate
RI 164 - type
 Sarcobora - Apatemyidae
Apatemys sp. Marsh 1872 insectivoran
RI 303, 424, 239 and 284
 Multituberculata - Neoplagiaulacidae
Ectypodus riansensis n. sp.7 Marandat et al. 2012 multituberculate
RI 273, 348, 363, 393, 413, 414, 376, 394, 418