Soyana (PIN collection 117) (Permian of Russian Federation)

Where: Arkangelsk, Russian Federation (65.6° N, 43.7° E: paleocoordinates 33.2° N, 33.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Iva-Gora Beds Formation, Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified siliciclastic sediments

• Probable brackish lagoon with some access to sea

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by Yu. D. Zekkel in 1935

Primary reference: A. V. Martynov. 1938. Permian fossil Insects from the Arkhangelsk district. Part V. The family Euthygrammidae and its relationships (with the description of a new genus and family from Chekarda). Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7(3):69-80 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 104716: authorized by Matthew Clapham, entered by Jered Karr on 08.02.2011, edited by Matthew Clapham, Lindy Edwards and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Insecta
 Hypoperlida - Hypoperlidae
Hypoperla elegans24, Boreopsocus danksae30
Hypoperla elegans24 Martynov 1928 winged insect
PIN 117/968
Boreopsocus danksae30 Shcherbakov 1994 winged insect
 Hemiptera - Scytinopteridae
Scytinoptera reducta13
Scytinoptera reducta13 Martynov 1928 true bug
PIN 117/1667, 1666, 2022, 1963, 577
 Hemiptera - Archescytinidae
Archescytinidae indet.15, Archescytina fusca15, Archescytina gunderseni15
Archescytinidae indet.15 Tillyard 1926 true bug
PIN 117/1703
Archescytina fusca15 Becker-Migdisova 1960 true bug
PIN 117/1654(1685)
Archescytina gunderseni15 Becker-Migdisova 1960 true bug
PIN 117/1991, 1855, 1931, 1798, 1934
 Hemiptera - Boreoscytidae
"Boreoscyta nefasta n. sp." = Boreoscyta nefasta14, "Boreoscyta imperfecta n. sp." = Boreoscyta nefasta14, "Boreoscyta mirabilis n. gen. n. sp." = Boreoscyta nefasta14
"Boreoscyta nefasta n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
PIN 117/1854
"Boreoscyta imperfecta n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
PIN 117/1673
"Boreoscyta mirabilis n. gen. n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
no number given
 Hemiptera - Prosbolopseidae
Anaprosbole ivensis n. gen. n. sp.15, Scytoneura elliptica15, Ivaia procercopoides n. sp.15, Prosbolopsis ovalis15
Anaprosbole ivensis n. gen. n. sp.15 Becker-Migdisova 1960 true bug
PIN 117/2020
Scytoneura elliptica15 Martynov 1935 true bug
PIN 117/1659(83), 1711, 2040, 1817(2127),1644(2039), 2068, 1694, 1693, 1876, 2044
Ivaia procercopoides n. sp.15 Becker-Migdisova 1960 true bug
PIN 117/1914
Prosbolopsis ovalis15 Martynov 1935 true bug
PIN 117/1819
 Hemiptera - Prosbolidae
Permocicada simulatrix12, Permocicada nigronervosa12, Permocicada triramosa12, Permocicada elliptica12, Permocicada integra12, Permocicada idelensis12, Permocicadopsis angustata12, Permocicadopsis aberrans12, Sojanoneura "sp. 1 nov"12, Sojanoneura edemskii12, Sojanoneura bimaculata12, Sojanoneura elytrata12, Sojanoneura triangularis12, Sojanoneura kazanensis12, Sojanoneura tincta n. sp.12, Prosbole dilatata12, Prosbole zekkeli n. sp.12, Prosbole curvimarginata n. sp.12
Permocicada simulatrix12 Martynov 1935 true bug
PIN 117/2049
Permocicada nigronervosa12 Martynov 1928 true bug
PIN 117/2016(2019), 1702, 1648, (2046, 2070(2057) (var. longifurcata))
Permocicada triramosa12 Becker-Migdisova 1940 true bug
PIN 117/1709
Permocicada elliptica12 Martynov 1935 true bug
PIN 117/2135
Permocicada integra12 Becker-Migdisova 1940 true bug
PIN 117/1705, 1662
Permocicada idelensis12 Zalessky 1929 true bug
PIN 117/1655, 1701(2063), 2060, 1696
Permocicadopsis angustata12 Martynov 1935 true bug
PIN 117/2131; 1665, 1679(2136) (var. typica); 1642(var. brevifurcillata); 1647(var. devia); 2132(var. remota)
Permocicadopsis aberrans12 Martynov 1935 true bug
PIN 117/1684, 1656, 1651
Sojanoneura "sp. 1 nov"12 Martynov 1928 true bug
PIN 117/2065
Sojanoneura edemskii12 Martynov 1928 true bug
PIN 117/1675(2051), 2037(1691), 1646(2059), 2134(2062)
Sojanoneura bimaculata12 Martynov 1935 true bug
PIN 117/2009, 1695, 2059, 1713, 2058
Sojanoneura elytrata12 Martynov 1928 true bug
PIN 117/1654(1641)
Sojanoneura triangularis12 Martynov 1935 true bug
PIN 117/1660, 1697(2014)
Sojanoneura kazanensis12 Zalessky 1929 true bug
PIN 117/2064(1652)
Sojanoneura tincta n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/2129
Prosbole dilatata12 Martynov 1935 true bug
PIN 117/2050
Prosbole zekkeli n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/2047(2139)
Prosbole curvimarginata n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/1687
 Dicondylia - Lophioneuridae
Lophiocypha incompleta n. sp.35
Lophiocypha incompleta n. sp.35 Vishnyakova 1981 winged insect
PIN 117/2442
 Dicondylia - Permosialidae
Permosialis quadriramosa n. sp.21, Permosialis latiformis n. sp.21, Permosialis bifasciata21, Permosialis fasciata n. sp.21, Permosialis perfecta n. sp.21, Permosialis defurcata n. sp.21
Permosialis quadriramosa n. sp.21 Martynova 1952 winged insect
PIN 117/1444(2652)
Permosialis latiformis n. sp.21 Martynova 1952 winged insect
PIN 117/1153, 2653
Permosialis bifasciata21 Martynov 1932 winged insect
PIN 117/1688(2656)
Permosialis fasciata n. sp.21 Martynova 1952 winged insect
PIN 117/2651
Permosialis perfecta n. sp.21 Martynova 1952 winged insect
PIN 117/1772
Permosialis defurcata n. sp.21 Martynova 1952 winged insect
PIN 117/1316
 Cnemidolestodea - Sylvabestiidae
Sojanopermula rasnitsyni n. sp.10, Kazanalicula reducta n. gen. n. sp.10
Sojanopermula rasnitsyni n. sp.10 Aristov and Storozhenko 2011 winged insect
PIN 117/301,195
Kazanalicula reducta n. gen. n. sp.10 Aristov and Storozhenko 2011 winged insect
PIN 117/467
 Cnemidolestodea - Protembiidae
Sojanoraphidia rossica n. gen. n. sp.21, Tshekardomina novokshonovi n. sp.7
Sojanoraphidia rossica n. gen. n. sp.21 Martynova 1952 winged insect
PIN 117/2549
Tshekardomina novokshonovi n. sp.7 Aristov 2015 winged insect
PIN 117/1528
 Cnemidolestodea - Pinideliidae
Idelopterum samoedum n. sp.7
Idelopterum samoedum n. sp.7 Aristov 2015 winged insect
PIN 117/43
 Caloneurodea - Caloneuridae
Euthygramma parallellum
Euthygramma parallellum Martynov 1928 winged insect
PIN 117/315, 316, 336, 357, 283, 160, 191, 101, 603, 182, 625
 Protorthoptera - Soyaniidae
Soyania logica8
Soyania logica8 Aristov and Rasnitsyn 2008 winged insect
PIN 177/1314,2759,2803
 Orthoptera - Pruvostitidae
Sylvoedischia curvovenosa n. sp.29
Sylvoedischia curvovenosa n. sp.29 Sharov 1968 grasshopper
PIN 117/426
 Orthoptera - Oedischiidae
Elcanoedischia major n. gen. n. sp.17, Elcanoedischia minor n. sp.17, "Pinegia martynovi n. sp." = Tcholmanvissia noinskii16
Elcanoedischia major n. gen. n. sp.17 Gorochov 1987 grasshopper
PIN 117/463
Elcanoedischia minor n. sp.17 Gorochov 1987 grasshopper
PIN 117/645+150, 702
"Pinegia martynovi n. sp." = Tcholmanvissia noinskii16 Zalessky 1929 grasshopper
PIN 117/258+259
 Orthoptera - Permelcanidae
Permelcana sojanense n. gen. n. sp.28
Permelcana sojanense n. gen. n. sp.28 Sharov 1962 grasshopper
PIN 117/346, 708
 Paoliida - Paoliidae
Stenaropodites sojanensis34, Stenaropodites mistshenkoi n. sp.34, Sojanidelia fluctuosa34, Sojanidelia parvula34, Sojanidelia lineata34, Sojanidelia longula n. sp.34, Sojanidelia makarkini34
Stenaropodites sojanensis34 Storozhenko 1992 winged insect
PIN 117/362(372), 374, 722, 794, 979(798), 814, 817, 835(840), 846, 860, 866, 2645, 103,104, 111, 356, 384(486), 508(9), 786, 888
Stenaropodites mistshenkoi n. sp.34 Storozhenko 1992 winged insect
PIN 117/801(4), 793, 353(836), 803
Sojanidelia fluctuosa34 Storozhenko 1992 winged insect
PIN 117/874
Sojanidelia parvula34 Storozhenko 1992 winged insect
PIN 117/2642
Sojanidelia lineata34 Storozhenko 1992 winged insect
PIN 117/816, 817, 861, 872
Sojanidelia longula n. sp.34 Storozhenko 1992 winged insect
PIN 117/814(869)
Sojanidelia makarkini34 Storozhenko 1992 winged insect
PIN 117/368, 1133
 Paoliida - Anthracoptilidae
Strephoneura pallida n. sp.25, Strephoneura vasta n. sp.25
Strephoneura pallida n. sp.25 Novokshonov and Aristov 2004 winged insect
PIN 117/376
Strephoneura vasta n. sp.25 Novokshonov and Aristov 2004 winged insect
PIN 117/90
 Grylloblattodea - Euremiscidae
Euremisca kazanica n. sp.2
Euremisca kazanica n. sp.2 Aristov 2009 ice crawler
PIN 117/1140
 Grylloblattodea - Camptoneuritidae
Camptoneurites soyanensis n. sp.11
Camptoneurites soyanensis n. sp.11 Aristov et al. 2010 ice crawler
PIN 117/244,807,871,2660
 Glosselytrodea - Jurinidae
Jurina marginata18
Jurina marginata18 Martynov 1938 winged insect
PIN 117/584
 Coleoptera - Permocupedidae
Permocupes semenovi26
Permocupes semenovi26 Martynov 1932 beetle
PIN 117/399(408)
 Mecoptera - Permochoristidae
Petromantis sp.23, ? Petromantis sp.23, "Petromantis robusta" = Petromantis robusta20, "Petromantis variolosa" = Petromantis robusta20, "Petromantis rossica" = Petromantis rossica20, "Petromantis borealis" = Petromantis rossica20, "Permochorista maculipennis" = Petromantis maculipennis20, "Permochorista venosa" = Petromantis maculipennis20, "Petrochorista dubia" = Petromantis dubia20, Petrochorista ivae n. sp.20, Petrochorista sojanensis20, "Agetochorista letopalae" = Agetopanorpa letopalae20, "Agetochorista ornata" = Agetopanorpa ornata20, "Agetochorista splendida" = Agetopanorpa ornata20, "Agetochorista dubia" = Agetopanorpa dubia20, "Agetochorista fasciata" = Agetopanorpa fasciata20
Petromantis sp.23 Handlirsch 1904 scorpionfly
PIN 117/2176, 1629-1851
? Petromantis sp.23 Handlirsch 1904 scorpionfly
PIN 117/276
"Petromantis robusta" = Petromantis robusta20 Martynov 1932 scorpionfly
PIN 117/1446
"Petromantis variolosa" = Petromantis robusta20 Martynov 1932 scorpionfly
PIN 117/1420
"Petromantis rossica" = Petromantis rossica20 Handlirsch 1904 scorpionfly
PIN 117/1321, 1336, 1358, 1361, 1392, 1397, 1406, 1439, 2133
"Petromantis borealis" = Petromantis rossica20 Handlirsch 1904 scorpionfly
PIN 117/1328, 1344, 1351, 1364, 1389, 1394, 1405, 1412, 1416, 1418, 1441, 2724, 2731
"Permochorista maculipennis" = Petromantis maculipennis20 Martynov 1932 scorpionfly
PIN 117/1332, 1334, 1340, 1367, 1376, 1379, 1393, 1403, 1411, 1433, 1440, 2687; var. major 117/1355
"Permochorista venosa" = Petromantis maculipennis20 Martynov 1932 scorpionfly
PIN 117/1111, 1448
"Petrochorista dubia" = Petromantis dubia20 Martynov 1932 scorpionfly
PIN 117/1320
Petrochorista ivae n. sp.20 Martynova 1948 scorpionfly
PIN 117/1341, 1407
Petrochorista sojanensis20 Martynova 1948 scorpionfly
PIN 117/1404, 2685
"Agetochorista letopalae" = Agetopanorpa letopalae20 Martynova 1948 scorpionfly
PIN 117/1339, 1414
"Agetochorista ornata" = Agetopanorpa ornata20 Martynov 1932 scorpionfly
PIN 117/1335, 1347, 1368, 1373, 1377, 1380, 1381, 1383, 1395, 1415, 1423, 1426, 1428, 1429, 1437, 1445
"Agetochorista splendida" = Agetopanorpa ornata20 Martynov 1932 scorpionfly
PIN 117/1333, 1342(1390), 1398; var. differens 117/1436
"Agetochorista dubia" = Agetopanorpa dubia20 Martynov 1932 scorpionfly
PIN 117/1401
"Agetochorista fasciata" = Agetopanorpa fasciata20 Martynov 1932 scorpionfly
PIN 117/1325, 1343, 2745
 Mecoptera - Permopanorpidae
"Permopanorpa angustata" = Martynopanorpa angustata20
"Permopanorpa angustata" = Martynopanorpa angustata20 Martynov 1932 scorpionfly
PIN 117/2695
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis20
Kamopanorpa rotundipennis20 Martynov 1932 winged insect
PIN 117/2695
 Raphidioptera - Parasialidae
Parasialis dissedens27, Sojanasialis longipennis n. gen. n. sp.27
Parasialis dissedens27 Ponomarenko 1977 snakefly
PIN 117/2146, 2335, 2600, 26220
Sojanasialis longipennis n. gen. n. sp.27 Ponomarenko 1977 snakefly
PIN 117/2459
 Dicondylia - Permithonidae
Permithonopsis cellulosa n. sp.21, "Permithonopsis ivensis" = Permithonopsis ivensis21, "Permithonopsis proxima" = Permithonopsis ivensis21, Permithonopsis longifurcata n. sp.21, "Bianchia spectabilis n. gen. n. sp." = Permegalomus spectabilis21, Palaemerobius avus n. sp.21
Permithonopsis cellulosa n. sp.21 Martynova 1952 winged insect
PIN 117/2533, 2560
"Permithonopsis ivensis" = Permithonopsis ivensis21 Martynov 1932 winged insect
PIN 117/2537(2598)
"Permithonopsis proxima" = Permithonopsis ivensis21 Martynov 1932 winged insect
PIN 117/2524
Permithonopsis longifurcata n. sp.21 Martynova 1952 winged insect
PIN 117/2515
"Bianchia spectabilis n. gen. n. sp." = Permegalomus spectabilis21 Martynova 1952 winged insect
PIN 117/2513
Palaemerobius avus n. sp.21 Martynova 1952 winged insect
PIN 117/2532(2592), 2540
 Palaeodictyoptera - Eukulojidae
Eukuloja atrosumma32, Eukuloja cubitalis32
Eukuloja atrosumma32 Sinitshenkova 1981 winged insect
PIN 117/24, 23(28), 47
Eukuloja cubitalis32 Martynov 1932 winged insect
PIN 117/18, 20-22, 25-27, 29-31, 34-36, 41(42), 46
 Palaeodictyoptera - Permothemistidae
Permothemis lepidus n. sp.31, Permothemis intermedius n. sp.31
Permothemis lepidus n. sp.31 Sinitshenkova 1980 winged insect
PIN 117/2554
Permothemis intermedius n. sp.31 Sinitshenkova 1980 winged insect
PIN 117/2529(16), 45
 Reculida - Geinitziidae
Permoshurabia mesenensis n. gen. n. sp.2, Shurabia inferior n. sp.7, Geinitzia ima n. sp.7, Stegopterum primus n. sp.7
Permoshurabia mesenensis n. gen. n. sp.2 Aristov 2009 winged insect
PIN 117/806, 102, 2774
Shurabia inferior n. sp.7 Aristov 2015 winged insect
PIN 117/282
Geinitzia ima n. sp.7 Aristov 2015 winged insect
PIN 117/805
Stegopterum primus n. sp.7 Aristov 2015 winged insect
PIN 117/765
 Reculida - Ivapteridae
Ivaptera sharovi n. gen. n. sp.3
Ivaptera sharovi n. gen. n. sp.3 Aristov 2009 winged insect
PIN 117/852, 140, 1085, 113, 123, 139, 224, 284, 322, 339, 379, 539, 565, 639, 783, 785, 825, 865, 873, 887, 897,945,950,964,979,990,992,994,995,999,1004,1005,1007,1009,1014,1019,1029,1038,1049,1050,1061,1071,1077,1081; PIN 117/477, 1134 in Aristov, 2015
 Reculida - Lemmatophoridae
Uraloprisca torta n. sp.7, Artinska sojanensis n. sp.4, Sylvaprisca colorata n. sp.4
Uraloprisca torta n. sp.7 Aristov 2015 winged insect
PIN 117/335
Artinska sojanensis n. sp.4 Aristov 2009 winged insect
PIN 117/1008, 1060
Sylvaprisca colorata n. sp.4 Aristov 2009 winged insect
PIN 117/855,444,1155
 Dicondylia - Permepallagidae
Permepallage angustissima n. gen. n. sp.19
Permepallage angustissima n. gen. n. sp.19 Martynov 1938 winged insect
PIN 117/65 (says Tshekarda in Nel et al. 2012)
 Dicondylia - Kennedyidae
Kennedya ivensis n. sp.22
Kennedya ivensis n. sp.22 Nel et al. 2012 winged insect
PIN 117/15
 Dicondylia - Permagrionidae
Epilestes angustapterix n. sp.22, Solikamptilon pectinatus n. sp.22
Epilestes angustapterix n. sp.22 Nel et al. 2012 winged insect
PIN 117/63
Solikamptilon pectinatus n. sp.22 Nel et al. 2012 winged insect
PIN 117/13
 Eoblattida - Permotermopsidae
Permotermopsis pectinata34, Permotermopsis roseni34
Permotermopsis pectinata34 Martynov 1937 insect
PIN 117/815(831), 830(7)
Permotermopsis roseni34 Martynov 1937 insect
PIN 117/265(6)
 Eoblattida - Mesorthopteridae
"Megakhosarina intricata n. sp." = Parastenaropodites intricatus1, Parastenaropodites aquilonius n. sp.6
"Megakhosarina intricata n. sp." = Parastenaropodites intricatus1 Aristov 2008 insect
PIN 117/144, 176, 740, 975, 982, 1082
Parastenaropodites aquilonius n. sp.6 Aristov 2014 insect
PIN 117/223
 Eoblattida - Megakhosaridae
"Megakhosarina minuscula n. sp." = Blattokhosara minuscula1, Megakhosarella paula n. sp.7, Ivakhosara pectinimedia n. gen. n. sp.1
"Megakhosarina minuscula n. sp." = Blattokhosara minuscula1 Aristov 2008 insect
PIN 117/86; 117/820 in Storozhenko & Aristov, 2014
Megakhosarella paula n. sp.7 Aristov 2015 insect
PIN 117/996
Ivakhosara pectinimedia n. gen. n. sp.1 Aristov 2008 insect
PIN 117/2488
 Eoblattida - Bardapteridae
Letopala costalis n. gen. n. sp.9, Olgaephilus emendatus33
Letopala costalis n. gen. n. sp.9 Aristov and Rasnitsyn 2011 insect
PIN 117/617
Olgaephilus emendatus33 Storozhenko 1990 insect
PIN 117/844, 383, 842
 Eoblattida - Blattogryllidae
Permoblattogryllus praecox n. gen. n. sp.5
Permoblattogryllus praecox n. gen. n. sp.5 Aristov 2011 insect
PIN 117/83
1Aristov 2008; 2Aristov 2009; 3Aristov 2009; 4Aristov 2009; 5Aristov 2011; 6Aristov 2014; 7Aristov 2015; 8Aristov and Rasnitsyn 2008; 9Aristov and Rasnitsyn 2011; 10Aristov and Storozhenko 2011; 11Aristov et al. 2010; 12Becker-Migdisova 1940; 13Becker-Migdisova 1948; 14Becker-Migdisova 1949; 15Becker-Migdisova 1960; 16Béthoux and Nel 2002; 17Gorochov 1987; 18Martynov 1938; 19Martynov 1938; 20Martynova 1948; 21Martynova 1952; 22Nel et al. 2012; 23Novokshonov 1994; 24Novokshonov 2001; 25Novokshonov and Aristov 2004; 26Ponomarenko 1963; 27Ponomarenko 1977; 28Sharov 1962; 29Sharov 1968; 30Shcherbakov 1994; 31Sinitshenkova 1980; 32Sinitshenkova 1981; 33Storozhenko 1990; 34Storozhenko 1992; 35Vishnyakova 1981