Soyana (PIN collection 117) (Permian of Russian Federation)

Where: Arkangelsk, Russian Federation (65.6° N, 43.7° E)

• Paleocoordinates: 33.4° N, 36.1° E (Wright 2013)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Iva-Gora Beds Formation, Roadian (274.4 - 266.9 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified siliciclastic sediments

• Probable brackish lagoon with some access to sea

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by Yu. D. Zekkel in 1935

Primary reference: A. V. Martynov. 1938. Permian fossil Insects from the Arkhangelsk district. Part V. The family Euthygrammidae and its relationships (with the description of a new genus and family from Chekarda). Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7(3):69-80 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 104716: authorized by Matthew Clapham, entered by Jered Karr on 08.02.2011, edited by Matthew Clapham, Lindy Edwards and Amy Delelli

Creative Commons license: CC0 (CC0)

Taxonomic list

Show authors, comments, and common names
Insecta
 Dicondylia - Kennedyidae
Kennedya ivensis n. sp.22
Kennedya ivensis n. sp.22 Nel et al. 2012 winged insect
PIN 117/15; 117/14(128) in Felker 2023
 Dicondylia - Permepallagidae
Permepallage angustissima n. gen. n. sp.19
Permepallage angustissima n. gen. n. sp.19 Martynov 1938 winged insect
PIN 117/65 (says Tshekarda in Nel et al. 2012)
 Dicondylia - Permagrionidae
Epilestes angustapterix n. sp.22, Solikamptilon pectinatus n. sp.22
Epilestes angustapterix n. sp.22 Nel et al. 2012 winged insect
PIN 117/63
Solikamptilon pectinatus n. sp.22 Nel et al. 2012 winged insect
PIN 117/13
 Permothemistida - Permothemistidae
Permothemis intermedius n. sp.31, Permothemis lepidus n. sp.31
Permothemis intermedius n. sp.31 Sinitshenkova 1980 winged insect
PIN 117/2529(16), 45
Permothemis lepidus n. sp.31 Sinitshenkova 1980 winged insect
PIN 117/2554
 Palaeodictyoptera - Eukulojidae
Eukuloja atrosumma32, Eukuloja cubitalis32
Eukuloja atrosumma32 Sinitshenkova 1981 winged insect
PIN 117/24, 23(28), 47
Eukuloja cubitalis32 Martynov 1932 winged insect
PIN 117/18, 20-22, 25-27, 29-31, 34-36, 41(42), 46
 Dicondylia - Permosialidae
"Permosialis bifasciata" = Permosialis paucinervis21, "Permosialis defurcata n. sp." = Permosialis paucinervis21, "Permosialis fasciata n. sp." = Permosialis paucinervis21, Permosialis perfecta n. sp.21, Permosialis latiformis n. sp.21, "Permosialis quadriramosa n. sp." = Palaeomantopsis furcatella21
"Permosialis bifasciata" = Permosialis paucinervis21 Martynov 1928 winged insect
PIN 117/1688(2656)
"Permosialis defurcata n. sp." = Permosialis paucinervis21 Martynov 1928 winged insect
PIN 117/1316
"Permosialis fasciata n. sp." = Permosialis paucinervis21 Martynov 1928 winged insect
PIN 117/2651
Permosialis perfecta n. sp.21 Martynova 1952 winged insect
PIN 117/1772
Permosialis latiformis n. sp.21 Martynova 1952 winged insect
PIN 117/1153, 2653
"Permosialis quadriramosa n. sp." = Palaeomantopsis furcatella21 Martynov 1928 winged insect
PIN 117/1444(2652)
 Hypoperlida - Hypoperlidae
Boreopsocus danksae30, Hypoperla elegans24
Boreopsocus danksae30 Shcherbakov 1994 winged insect
Hypoperla elegans24 Martynov 1928 winged insect
PIN 117/968
 Hemiptera - Archescytinidae
Archescytinidae indet.15, Archescytina fusca15, Archescytina gunderseni15
Archescytinidae indet.15 Tillyard 1926 true bug
PIN 117/1703
Archescytina fusca15 Becker-Migdisova 1960 true bug
PIN 117/1654(1685)
Archescytina gunderseni15 Becker-Migdisova 1960 true bug
PIN 117/1991, 1855, 1931, 1798, 1934
 Hemiptera - Prosbolidae
Sojanoneura "sp. 1 nov"12, Sojanoneura tincta n. sp.12, Sojanoneura triangularis12, Sojanoneura elytrata12, Sojanoneura edemskii12, Sojanoneura kazanensis12, Sojanoneura bimaculata12, Permocicada triramosa12, Permocicada integra12, Permocicada nigronervosa12, Permocicada simulatrix12, Permocicada elliptica12, Permocicada idelensis12, Prosbole dilatata12, Prosbole zekkeli n. sp.12, Prosbole curvimarginata n. sp.12, Permocicadopsis angustata12, Permocicadopsis aberrans12
Sojanoneura "sp. 1 nov"12 Martynov 1928 true bug
PIN 117/2065
Sojanoneura tincta n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/2129
Sojanoneura triangularis12 Martynov 1935 true bug
PIN 117/1660, 1697(2014)
Sojanoneura elytrata12 Martynov 1928 true bug
PIN 117/1654(1641)
Sojanoneura edemskii12 Martynov 1928 true bug
PIN 117/1675(2051), 2037(1691), 1646(2059), 2134(2062)
Sojanoneura kazanensis12 Zalessky 1929 true bug
PIN 117/2064(1652)
Sojanoneura bimaculata12 Martynov 1935 true bug
PIN 117/2009, 1695, 2059, 1713, 2058
Permocicada triramosa12 Becker-Migdisova 1940 true bug
PIN 117/1709
Permocicada integra12 Becker-Migdisova 1940 true bug
PIN 117/1705, 1662
Permocicada nigronervosa12 Martynov 1928 true bug
PIN 117/2016(2019), 1702, 1648, (2046, 2070(2057) (var. longifurcata))
Permocicada simulatrix12 Martynov 1935 true bug
PIN 117/2049
Permocicada elliptica12 Martynov 1935 true bug
PIN 117/2135
Permocicada idelensis12 Zalessky 1929 true bug
PIN 117/1655, 1701(2063), 2060, 1696
Prosbole dilatata12 Martynov 1935 true bug
PIN 117/2050
Prosbole zekkeli n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/2047(2139)
Prosbole curvimarginata n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/1687
Permocicadopsis angustata12 Martynov 1935 true bug
PIN 117/2131; 1665, 1679(2136) (var. typica); 1642(var. brevifurcillata); 1647(var. devia); 2132(var. remota)
Permocicadopsis aberrans12 Martynov 1935 true bug
PIN 117/1684, 1656, 1651
 Hemiptera - Prosbolopseidae
Prosbolopsis ovalis15, Anaprosbole ivensis n. gen. n. sp.15, Scytoneura elliptica15, Ivaia procercopoides n. sp.15
Prosbolopsis ovalis15 Martynov 1935 true bug
PIN 117/1819
Anaprosbole ivensis n. gen. n. sp.15 Becker-Migdisova 1960 true bug
PIN 117/2020
Scytoneura elliptica15 Martynov 1935 true bug
PIN 117/1659(83), 1711, 2040, 1817(2127),1644(2039), 2068, 1694, 1693, 1876, 2044
Ivaia procercopoides n. sp.15 Becker-Migdisova 1960 true bug
PIN 117/1914
 Hemiptera - Boreoscytidae
"Boreoscyta nefasta n. sp." = Boreoscyta nefasta14, "Boreoscyta imperfecta n. sp." = Boreoscyta nefasta14, "Boreoscyta mirabilis n. gen. n. sp." = Boreoscyta nefasta14
"Boreoscyta nefasta n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
PIN 117/1854
"Boreoscyta imperfecta n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
PIN 117/1673
"Boreoscyta mirabilis n. gen. n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
no number given
 Hemiptera - Scytinopteridae
Scytinoptera reducta13
Scytinoptera reducta13 Martynov 1928 true bug
PIN 117/1667, 1666, 2022, 1963, 577
 Lophioneurida - Lophioneuridae
Lophiocypha incompleta n. sp.35
Lophiocypha incompleta n. sp.35 Vishnyakova 1981 winged insect
PIN 117/2442
 Protorthoptera - Soyaniidae
Soyania logica8
Soyania logica8 Aristov and Rasnitsyn 2008 winged insect
PIN 177/1314,2759,2803
 Orthoptera - Oedischiidae
Elcanoedischia minor n. sp.17, Elcanoedischia major n. gen. n. sp.17, "Pinegia martynovi n. sp." = Tcholmanvissia noinskii16
Elcanoedischia minor n. sp.17 Gorochov 1987 grasshopper
PIN 117/645+150, 702
Elcanoedischia major n. gen. n. sp.17 Gorochov 1987 grasshopper
PIN 117/463
"Pinegia martynovi n. sp." = Tcholmanvissia noinskii16 Zalessky 1929 grasshopper
PIN 117/258+259
 Orthoptera - Pruvostitidae
Sylvoedischia curvovenosa n. sp.29
Sylvoedischia curvovenosa n. sp.29 Sharov 1968 grasshopper
PIN 117/426
 Orthoptera - Permelcanidae
Permelcana sojanense n. gen. n. sp.28
Permelcana sojanense n. gen. n. sp.28 Sharov 1962 grasshopper
PIN 117/346, 708
 Grylloblattodea - Blattogryllidae
Permoblattogryllus praecox n. gen. n. sp.5
Permoblattogryllus praecox n. gen. n. sp.5 Aristov 2011 ice crawler
PIN 117/83
 Grylloblattodea - Euremiscidae
Euremisca kazanica n. sp.4
Euremisca kazanica n. sp.4 Aristov 2009 ice crawler
PIN 117/1140
 Grylloblattodea - Lemmatophoridae
Artinska sojanensis n. sp.3, Sylvaprisca colorata n. sp.3, Uraloprisca torta n. sp.7
Artinska sojanensis n. sp.3 Aristov 2009 ice crawler
PIN 117/1008, 1060
Sylvaprisca colorata n. sp.3 Aristov 2009 ice crawler
PIN 117/855,444,1155
Uraloprisca torta n. sp.7 Aristov 2015 ice crawler
PIN 117/335
 Grylloblattodea - Camptoneuritidae
Camptoneurites soyanensis n. sp.11
Camptoneurites soyanensis n. sp.11 Aristov et al. 2010 ice crawler
PIN 117/244,807,871,2660
 Paoliida - Paoliidae
Permotermopsis roseni34, Permotermopsis pectinata34, Sojanidelia longula n. sp.34, Sojanidelia parvula34, Sojanidelia lineata34, Sojanidelia fluctuosa34, Sojanidelia makarkini34, Stenaropodites sojanensis34, Stenaropodites mistshenkoi n. sp.34
Permotermopsis roseni34 Martynov 1937 winged insect
PIN 117/265(6)
Permotermopsis pectinata34 Martynov 1937 winged insect
PIN 117/815(831), 830(7)
Sojanidelia longula n. sp.34 Storozhenko 1992 winged insect
PIN 117/814(869)
Sojanidelia parvula34 Storozhenko 1992 winged insect
PIN 117/2642
Sojanidelia lineata34 Storozhenko 1992 winged insect
PIN 117/816, 817, 861, 872
Sojanidelia fluctuosa34 Storozhenko 1992 winged insect
PIN 117/874
Sojanidelia makarkini34 Storozhenko 1992 winged insect
PIN 117/368, 1133
Stenaropodites sojanensis34 Storozhenko 1992 winged insect
PIN 117/362(372), 374, 722, 794, 979(798), 814, 817, 835(840), 846, 860, 866, 2645, 103,104, 111, 356, 384(486), 508(9), 786, 888
Stenaropodites mistshenkoi n. sp.34 Storozhenko 1992 winged insect
PIN 117/801(4), 793, 353(836), 803
 Paoliida - Anthracoptilidae
Strephoneura vasta n. sp.25, Strephoneura pallida n. sp.25
Strephoneura vasta n. sp.25 Novokshonov and Aristov 2004 winged insect
PIN 117/90
Strephoneura pallida n. sp.25 Novokshonov and Aristov 2004 winged insect
PIN 117/376
 Caloneurodea - Caloneuridae
Euthygramma parallellum
Euthygramma parallellum Martynov 1928 winged insect
PIN 117/315, 316, 336, 357, 283, 160, 191, 101, 603, 182, 625
 Cnemidolestodea - Sylvabestiidae
Kazanalicula reducta n. gen. n. sp.10, Sojanopermula rasnitsyni n. sp.10
Kazanalicula reducta n. gen. n. sp.10 Aristov and Storozhenko 2011 winged insect
PIN 117/467
Sojanopermula rasnitsyni n. sp.10 Aristov and Storozhenko 2011 winged insect
PIN 117/301,195
 Cnemidolestodea - Pinideliidae
Idelopterum samoedum n. sp.7
Idelopterum samoedum n. sp.7 Aristov 2015 winged insect
PIN 117/43
 Cnemidolestodea - Protembiidae
Tshekardomina novokshonovi n. sp.7, Sojanoraphidia rossica n. gen. n. sp.21
Tshekardomina novokshonovi n. sp.7 Aristov 2015 winged insect
PIN 117/1528
Sojanoraphidia rossica n. gen. n. sp.21 Martynova 1952 winged insect
PIN 117/2549
 Glosselytrodea - Jurinidae
Jurina marginata18
Jurina marginata18 Martynov 1938 winged insect
PIN 117/584
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis20
Kamopanorpa rotundipennis20 Martynov 1932 winged insect
PIN 117/2695
 Mecoptera - Permochoristidae
Petrochorista sojanensis20, Petrochorista ivae n. sp.20, Petromantis sp.23, ? Petromantis sp.23, "Permochorista maculipennis" = Petromantis maculipennis20, "Permochorista venosa" = Petromantis maculipennis20, "Petromantis rossica" = Petromantis rossica20, "Petromantis borealis" = Petromantis rossica20, "Petrochorista dubia" = Petromantis dubia20, "Petromantis robusta" = Petromantis robusta20, "Petromantis variolosa" = Petromantis robusta20, "Agetochorista letopalae" = Agetopanorpa letopalae20, "Agetochorista fasciata" = Agetopanorpa fasciata20, "Agetochorista ornata" = Agetopanorpa ornata20, "Agetochorista splendida" = Agetopanorpa ornata20, "Agetochorista dubia" = Agetopanorpa dubia20
Petrochorista sojanensis20 Martynova 1948 scorpionfly
PIN 117/1404, 2685
Petrochorista ivae n. sp.20 Martynova 1948 scorpionfly
PIN 117/1341, 1407
Petromantis sp.23 Handlirsch 1904 scorpionfly
PIN 117/2176, 1629-1851
? Petromantis sp.23 Handlirsch 1904 scorpionfly
PIN 117/276
"Permochorista maculipennis" = Petromantis maculipennis20 Martynov 1932 scorpionfly
PIN 117/1332, 1334, 1340, 1367, 1376, 1379, 1393, 1403, 1411, 1433, 1440, 2687; var. major 117/1355
"Permochorista venosa" = Petromantis maculipennis20 Martynov 1932 scorpionfly
PIN 117/1111, 1448
"Petromantis rossica" = Petromantis rossica20 Handlirsch 1904 scorpionfly
PIN 117/1321, 1336, 1358, 1361, 1392, 1397, 1406, 1439, 2133
"Petromantis borealis" = Petromantis rossica20 Handlirsch 1904 scorpionfly
PIN 117/1328, 1344, 1351, 1364, 1389, 1394, 1405, 1412, 1416, 1418, 1441, 2724, 2731
"Petrochorista dubia" = Petromantis dubia20 Martynov 1932 scorpionfly
PIN 117/1320
"Petromantis robusta" = Petromantis robusta20 Martynov 1932 scorpionfly
PIN 117/1446
"Petromantis variolosa" = Petromantis robusta20 Martynov 1932 scorpionfly
PIN 117/1420
"Agetochorista letopalae" = Agetopanorpa letopalae20 Martynova 1948 scorpionfly
PIN 117/1339, 1414
"Agetochorista fasciata" = Agetopanorpa fasciata20 Martynov 1932 scorpionfly
PIN 117/1325, 1343, 2745
"Agetochorista ornata" = Agetopanorpa ornata20 Martynov 1932 scorpionfly
PIN 117/1335, 1347, 1368, 1373, 1377, 1380, 1381, 1383, 1395, 1415, 1423, 1426, 1428, 1429, 1437, 1445
"Agetochorista splendida" = Agetopanorpa ornata20 Martynov 1932 scorpionfly
PIN 117/1333, 1342(1390), 1398; var. differens 117/1436
"Agetochorista dubia" = Agetopanorpa dubia20 Martynov 1932 scorpionfly
PIN 117/1401
 Mecoptera - Permopanorpidae
"Permopanorpa angustata" = Martynopanorpa angustata20
"Permopanorpa angustata" = Martynopanorpa angustata20 Martynov 1932 scorpionfly
PIN 117/2695
 Dicondylia - Permithonidae
Palaemerobius avus n. sp.21, "Bianchia spectabilis n. gen. n. sp." = Permegalomus spectabilis21, "Permithonopsis ivensis" = Permithonopsis ivensis21, "Permithonopsis proxima" = Permithonopsis ivensis21, Permithonopsis cellulosa n. sp.21, Permithonopsis longifurcata n. sp.21
Palaemerobius avus n. sp.21 Martynova 1952 winged insect
PIN 117/2532(2592), 2540
"Bianchia spectabilis n. gen. n. sp." = Permegalomus spectabilis21 Martynova 1952 winged insect
PIN 117/2513
"Permithonopsis ivensis" = Permithonopsis ivensis21 Martynov 1932 winged insect
PIN 117/2537(2598)
"Permithonopsis proxima" = Permithonopsis ivensis21 Martynov 1932 winged insect
PIN 117/2524
Permithonopsis cellulosa n. sp.21 Martynova 1952 winged insect
PIN 117/2533, 2560
Permithonopsis longifurcata n. sp.21 Martynova 1952 winged insect
PIN 117/2515
 Raphidioptera - Parasialidae
Parasialis dissedens27, Sojanasialis longipennis n. gen. n. sp.27
Parasialis dissedens27 Ponomarenko 1977 snakefly
PIN 117/2146, 2335, 2600, 26220
Sojanasialis longipennis n. gen. n. sp.27 Ponomarenko 1977 snakefly
PIN 117/2459
 Coleoptera - Permocupedidae
Permocupes semenovi26
Permocupes semenovi26 Martynov 1932 beetle
PIN 117/399(408)
 Reculida - Geinitziidae
Geinitzia ima n. sp.7, Stegopterum primus n. sp.7, Permoshurabia mesenensis n. gen. n. sp.4, Shurabia inferior n. sp.7
Geinitzia ima n. sp.7 Aristov 2015 winged insect
PIN 117/805
Stegopterum primus n. sp.7 Aristov 2015 winged insect
PIN 117/765
Permoshurabia mesenensis n. gen. n. sp.4 Aristov 2009 winged insect
PIN 117/806, 102, 2774
Shurabia inferior n. sp.7 Aristov 2015 winged insect
PIN 117/282
 Reculida - Ivapteridae
Ivaptera sharovi n. gen. n. sp.2
Ivaptera sharovi n. gen. n. sp.2 Aristov 2009 winged insect
PIN 117/852, 140, 1085, 113, 123, 139, 224, 284, 322, 339, 379, 539, 565, 639, 783, 785, 825, 865, 873, 887, 897,945,950,964,979,990,992,994,995,999,1004,1005,1007,1009,1014,1019,1029,1038,1049,1050,1061,1071,1077,1081; PIN 117/477, 1134 in Aristov, 2015
 Eoblattida - Megakhosaridae
Ivakhosara pectinimedia n. gen. n. sp.1, Megakhosarella paula n. sp.7, "Megakhosarina minuscula n. sp." = Blattokhosara minuscula1
Ivakhosara pectinimedia n. gen. n. sp.1 Aristov 2008 insect
PIN 117/2488
Megakhosarella paula n. sp.7 Aristov 2015 insect
PIN 117/996
"Megakhosarina minuscula n. sp." = Blattokhosara minuscula1 Aristov 2008 insect
PIN 117/86; 117/820 in Storozhenko & Aristov, 2014
 Eoblattida - Bardapteridae
Olgaephilus emendatus33, Letopala costalis n. gen. n. sp.9
Olgaephilus emendatus33 Storozhenko 1990 insect
PIN 117/844, 383, 842
Letopala costalis n. gen. n. sp.9 Aristov and Rasnitsyn 2011 insect
PIN 117/617
 Eoblattida - Mesorthopteridae
Parastenaropodites aquilonius n. sp.6, "Megakhosarina intricata n. sp." = Parastenaropodites intricatus1
Parastenaropodites aquilonius n. sp.6 Aristov 2014 insect
PIN 117/223
"Megakhosarina intricata n. sp." = Parastenaropodites intricatus1 Aristov 2008 insect
PIN 117/144, 176, 740, 975, 982, 1082
1Aristov 2008; 2Aristov 2009; 3Aristov 2009; 4Aristov 2009; 5Aristov 2011; 6Aristov 2014; 7Aristov 2015; 8Aristov and Rasnitsyn 2008; 9Aristov and Rasnitsyn 2011; 10Aristov and Storozhenko 2011; 11Aristov et al. 2010; 12Becker-Migdisova 1940; 13Becker-Migdisova 1948; 14Becker-Migdisova 1949; 15Becker-Migdisova 1960; 16Béthoux and Nel 2002; 17Gorochov 1987; 18Martynov 1938; 19Martynov 1938; 20Martynova 1948; 21Martynova 1952; 22Nel et al. 2012; 23Novokshonov 1994; 24Novokshonov 2001; 25Novokshonov and Aristov 2004; 26Ponomarenko 1963; 27Ponomarenko 1977; 28Sharov 1962; 29Sharov 1968; 30Shcherbakov 1994; 31Sinitshenkova 1980; 32Sinitshenkova 1981; 33Storozhenko 1990; 34Storozhenko 1992; 35Vishnyakova 1981