PBDB Collection

Mammal Quarry No. 2 (UCMP V-5815) (Cretaceous of the United States)

Also known as Lance Quarry 2, Hatcher No. 2

Where: Niobrara County, Wyoming (43.1° N, 104.5° W)

• Paleocoordinates: 53.6° N, 65.8° W (Wright 2013)

• coordinate estimated from map

• small collection-level geographic resolution

When: Lance Formation, Lancian (72.2 - 66.0 Ma)

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; sandstone

• "cross-stratified sandstone"

Size class: mesofossils

Collected by J. B. Hatcher, W. A. Clemens in 1889, 1958; reposited in the UCMP, YPM

Collection methods: bulk, sieve,

Primary reference: R. S. Lull. 1915. The mammals and horned dinosaurs of the Lance Formation of Niobrara County, Wyoming. The American Journal of Science, series 4 40(238):319-348 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14566: authorized by John Alroy, entered by John Alroy on 26.07.1994, edited by Matthew Carrano, Roger Benson and Patricia Holroyd

Creative Commons license: CC0 (CC0)

Taxonomic list

Show authors, comments, and common names

Gastropoda
Sorbeoconcha - Hydrobiidae	Reesidella sp.⁹ Reesidella sp.⁹ Yen 1952 snail
Bivalvia
Unionida - Unionidae	Unionidae indet.⁹ Unionidae indet.⁹ Rafinesque 1820 freshwater mussel
Chondrichthyes
Hybodontiformes - Lonchidiidae	Lonchidion selachos⁹ Lonchidion selachos⁹ Estes 1964 elasmobranch
Amphibia
Salientia - Pelobatidae	Eopelobates sp.¹⁰ Eopelobates sp.¹⁰ Parker 1929 spadefoot toad
Caudata - Urodela	Habrosaurus dilatus⁹, Scapherpeton tectum⁹ Habrosaurus dilatus⁹ Gilmore 1928 siren Scapherpeton tectum⁹ Cope 1876 salamander
Proteida - Batrachosauroididae	"Prodesmodon copei" = Prodesmodon copei⁹, "Cuttysarkus mcnallyi" = Prodesmodon copei⁹, Opisthotriton kayi⁹ "Prodesmodon copei" = Prodesmodon copei⁹ Estes 1964 salamander "Cuttysarkus mcnallyi" = Prodesmodon copei⁹ Estes 1964 salamander Opisthotriton kayi⁹ Auffenberg 1961 salamander
Mammalia
Cimolesta - Cimolestidae	Cimolestes incisus⁸ Cimolestes incisus⁸ Marsh 1889 eutherian
Leptictida - Gypsonictopidae	Gypsonictops hypoconus⁸ Gypsonictops hypoconus⁸ Simpson 1927 eutherian
Tribosphenida - Stagodontidae	Didelphodon vorax⁷ Didelphodon vorax⁷ Marsh 1889 metatherian
Tribosphenida - Alphadontidae	"Alphadon rhaister n. sp." = Turgidodon rhaister⁶, Alphadon marshi⁷ "Alphadon rhaister n. sp." = Turgidodon rhaister⁶ Clemens 1966 metatherian Alphadon marshi⁷ Simpson 1927 metatherian
Tribosphenida - Herpetotheriidae	"Nortedelphys intermedius" = Nortedelphys jasoni⁵ "Nortedelphys intermedius" = Nortedelphys jasoni⁵ Storer 1991 marsupial
Tribosphenida -	"Alphadon lulli" = Protalphadon lulli⁶ "Alphadon lulli" = Protalphadon lulli⁶ Clemens 1966 metatherian
Tribosphenida - Pediomyidae	"Pediomys krejcii" = Leptalestes krejcii⁷, "Pediomys cooki" = Leptalestes cooki⁷, Pediomys elegans⁷, "Pediomys hatcheri" = Protolambda hatcheri⁷, "Pediomys florencae" = Protolambda florencae⁷ "Pediomys krejcii" = Leptalestes krejcii⁷ Clemens 1966 metatherian "Pediomys cooki" = Leptalestes cooki⁷ Clemens 1966 metatherian Pediomys elegans⁷ Marsh 1889 metatherian "Pediomys hatcheri" = Protolambda hatcheri⁷ Osborn 1898 metatherian "Pediomys florencae" = Protolambda florencae⁷ Clemens 1966 metatherian
Multituberculata - Cimolomyidae	Cimolomys gracilis⁶, "Dipriodon robustus n. sp." = Meniscoessus robustus Cimolomys gracilis⁶ Marsh 1889 multituberculate present in both UCMP and Hatcher collections "Dipriodon robustus n. sp." = Meniscoessus robustus Marsh 1889 multituberculate
Multituberculata - Cimolodontidae	Cimolodon nitidus⁶ Cimolodon nitidus⁶ Marsh 1889 multituberculate
Multituberculata - Neoplagiaulacidae	Mesodma sp.¹, Mesodma thompsoni⁶ Mesodma sp.¹ Jepsen 1940 multituberculate Mesodma thompsoni⁶ Clemens 1964 multituberculate
Reptilia
Testudinata -	Compsemys victa⁹ Compsemys victa⁹ Leidy 1856 turtle
Testudinata - Baenidae	Baenidae indet.⁴ Baenidae indet.⁴ Cope 1882 turtle Including UCMP 173424, listed by Holroyd & Hutchison (2002)
Testudines -	Cryptodira indet.⁴ Cryptodira indet.⁴ Cope 1868 turtle
Testudines - Trionychidae	Trionychidae indet.⁴ Trionychidae indet.⁴ Gray 1825 softshell turtle Including UCMP 173026, listed by Holroyd & Hutchison (2002)
Testudines - Emydidae	Emydinae indet.⁹ Emydinae indet.⁹ Rafinesque 1815 turtle "near Pseudemys?"
Testudines - Kinosternidae	"Kinosternidae indet." = Kinosternoidea⁴ "Kinosternidae indet." = Kinosternoidea⁴ Hay 1892 mud turtle Including UCMP 173422, listed by Holroyd & Hutchison (2002)
Testudines - Chelydridae	? Chelydridae indet.⁴, Chelydridae indet.⁴ ? Chelydridae indet.⁴ Swainson 1839 turtle Chelydridae indet.⁴ Swainson 1839 turtle Including UCMP 173420, listed by Holroyd & Hutchison (2002)
Archosauria -	Archosauria indet.⁴ Archosauria indet.⁴ archosaur
Crocodylia - Alligatoridae	Brachychampsa montana⁹ Brachychampsa montana⁹ Gilmore 1911 crocodilian
Crocodylia -	"Leidyosuchus sternbergi" = Borealosuchus sternbergii⁹ "Leidyosuchus sternbergi" = Borealosuchus sternbergii⁹ Gilmore 1910 crocodilian
Ornithischia -	Thescelosaurus sp.³ Thescelosaurus sp.³ Gilmore 1913 ornithopod
Saurischia -	cf. Dryptosaurus sp.⁹ cf. Dryptosaurus sp.⁹ Marsh 1877 coelurosaur
Saurischia - Troodontidae	Troodontidae indet.⁴, Troodon sp.⁴, Pectinodon bakkeri³ Troodontidae indet.⁴ Gilmore 1924 maniraptoran Troodon sp.⁴ Leidy 1856 maniraptoran Pectinodon bakkeri³ Carpenter 1982 maniraptoran
Saurischia - Dromaeosauridae	Saurornitholestinae indet.¹², Richardoestesia sp.⁴ Saurornitholestinae indet.¹² Longrich and Currie 2009 maniraptoran UCMP 187054 Richardoestesia sp.⁴ Currie et al. 1990 maniraptoran
Aves
-	Aves indet.⁴ Aves indet.⁴ Linnaeus 1758 bird
Reptilia
Saurischia -	Paronychodon lacustris⁹, cf. Paronychodon lacustris¹⁴ Paronychodon lacustris⁹ Cope 1876 maniraptoran cf. Paronychodon lacustris¹⁴ Cope 1876 maniraptoran UCMP 73076
Choristodera -	Champsosaurus sp.⁹ Champsosaurus sp.⁹ Cope 1876 choristodere
Squamata -	Paraderma bogerti⁹ Paraderma bogerti⁹ Estes 1964 squamates
Squamata - Anguidae	"Peltosaurus piger" = Odaxosaurus piger⁹ "Peltosaurus piger" = Odaxosaurus piger⁹ Gilmore 1928 squamates
Squamata -	Parasaniwa wyomingensis⁹, Colpodontosaurus cracens⁹, Varanoidea indet.¹¹ Parasaniwa wyomingensis⁹ Gilmore 1928 squamates Colpodontosaurus cracens⁹ Estes 1964 squamates Varanoidea indet.¹¹ Gray 1827 squamates
Squamata - Xenosauridae	Exostinus lancensis⁹ Exostinus lancensis⁹ Gilmore 1928 squamates
Squamata - Chamopsiidae	Leptochamops denticulatus⁹, Haptosphenus placodon⁹, Chamops segnis⁹, Meniscognathus altmani⁹ Leptochamops denticulatus⁹ Gilmore 1928 squamates Haptosphenus placodon⁹ Estes 1964 squamates Chamops segnis⁹ Marsh 1892 squamates Meniscognathus altmani⁹ Estes 1964 squamates
Squamata -	Coniophis precedens¹³ Coniophis precedens¹³ Marsh 1892 snake UCMP 53556‐53561 (vertebrae)
Osteichthyes
Acipenseriformes - Acipenseridae	Acipenser eruciferus⁹ Acipenser eruciferus⁹ Cope 1876 sturgeon
Lepisosteiformes - Lepisosteidae	Lepisosteus occidentalis⁹ Lepisosteus occidentalis⁹ Leidy 1856 gar
Amiiformes - Amiidae	"Kindleia fragosa" = Cyclurus fragosus⁹, cf. Protamia sp.⁹, Melvius thomasi² "Kindleia fragosa" = Cyclurus fragosus⁹ Jordan 1927 bowfin cf. Protamia sp.⁹ Leidy 1873 bowfin Melvius thomasi² Bryant 1988 bowfin
Esociformes - Esocidae	Estesesox foxi¹⁵ Estesesox foxi¹⁵ Wilson et al. 1992 ray-finned fish
Elopiformes - Elopidae	Elopidae indet.⁹ Elopidae indet.⁹ Bonaparte 1846 ray-finned fish
Albuliformes - Albulidae	Albulidae indet.⁹ Albulidae indet.⁹ Bleeker 1859 bonefish
Aspidorhynchiformes - Aspidorhynchidae	Belonostomus longirostris⁹ Belonostomus longirostris⁹ Lambe 1902 ray-finned fish

¹Alroy 2002; ²Bryant 1988; ³Carpenter 1982; ⁴Carrano 2005; ⁵Case et al. 2004; ⁶Clemens 1964; ⁷Clemens 1966; ⁸Clemens 1973; ⁹Estes 1964; ¹⁰Estes and Sanchíz 1982; ¹¹Gao and Fox 1996; ¹²Larson and Currie 2013; ¹³Longrich et al. 2012; ¹⁴Sankey 2008; ¹⁵Wilson et al. 1992