PBDB

Hatcher No. 2 (UCMP V-5815) (Cretaceous of the United States)

Also known as Lance Quarry 2

Where: Niobrara County, Wyoming (43.1° N, 104.5° W: paleocoordinates 49.6° N, 79.4° W)

• coordinate estimated from map

• small collection-level geographic resolution

When: Lance Formation, Lancian (70.6 - 66.0 Ma)

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; sandstone

• "cross-stratified sandstone"

Size class: mesofossils

Collected by J. B. Hatcher, W. A. Clemens in 1889; reposited in the UCMP, YPM

Collection methods: bulk, sieve,

Primary reference: R. S. Lull. 1915. The mammals and horned dinosaurs of the Lance Formation of Niobrara County, Wyoming. The American Journal of Science, series 4 40(238):319-348 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14566: authorized by John Alroy, entered by John Alroy on 26.07.1994, edited by Matthew Carrano and Roger Benson

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Bivalvia
Unionida - Unionidae	Unionidae indet.⁸ Unionidae indet.⁸ Rafinesque 1820 freshwater mussel
Gastropoda
Truncatelloidea - Hydrobiidae	Reesidella sp.⁸ Reesidella sp.⁸ Yen 1952 snail
Chondrichthyes
Hybodontiformes - Lonchidiidae	Lonchidion selachos⁸ Lonchidion selachos⁸ Estes 1964 elasmobranch
Actinopteri
Acipenseriformes - Acipenseridae	Acipenser eruciferus⁸ Acipenser eruciferus⁸ Cope 1876 sturgeon
Teleostei - Aspidorhynchidae	Belonostomus longirostris⁸ Belonostomus longirostris⁸ Lambe 1902
Elopiformes - Elopidae	Elopidae indet.⁸ Elopidae indet.⁸ Bonaparte 1846
Albuliformes - Albulidae	Albulidae indet.⁸ Albulidae indet.⁸ Bleeker 1859 bonefish
Acanthomorphata - Sciaenidae	Platacodon nanus⁸ Platacodon nanus⁸ Marsh 1889
Amiiformes - Amiidae	Melvius thomasi⁷, "Kindleia fragosa" = Cyclurus fragosus⁸, cf. Protamia sp.⁸ Melvius thomasi⁷ Bryant 1987 bowfin "Kindleia fragosa" = Cyclurus fragosus⁸ Jordan 1927 bowfin cf. Protamia sp.⁸ Leidy 1873 bowfin
Lepisosteiformes - Lepisosteidae	Lepisosteus occidentalis⁸ Lepisosteus occidentalis⁸ Leidy 1856 gar
Osteichthyes
Salientia - Pelobatidae	Eopelobates sp.⁹ Eopelobates sp.⁹ Parker 1929 spadefoot toad
Proteida - Batrachosauroididae	"Prodesmodon copei" = Prodesmodon copei⁸, "Cuttysarkus mcnallyi" = Prodesmodon copei⁸, Opisthotriton kayi⁸ "Prodesmodon copei" = Prodesmodon copei⁸ Estes 1964 salamander "Cuttysarkus mcnallyi" = Prodesmodon copei⁸ Estes 1964 salamander Opisthotriton kayi⁸ Auffenberg 1961 salamander
Caudata - Urodela	Habrosaurus dilatus⁸, Scapherpeton tectum⁸ Habrosaurus dilatus⁸ Gilmore 1928 siren Scapherpeton tectum⁸ Cope 1876 salamander
Reptilia
Testudines - Trionychidae	Trionychidae indet.⁷ Trionychidae indet.⁷ Gray 1825 softshell turtle Including UCMP 173026, listed by Holroyd & Hutchison (2002)
Testudines -	Compsemys victa⁸ Compsemys victa⁸ Leidy 1856 turtle
Testudines - Baenidae	Baenidae indet.⁷ Baenidae indet.⁷ Cope 1882 turtle Including UCMP 173424, listed by Holroyd & Hutchison (2002)
Testudines -	Cryptodira indet.⁷ Cryptodira indet.⁷ Cope 1868 turtle
Testudines - Kinosternidae	"Kinosternidae indet." = Kinosternoidea⁷ "Kinosternidae indet." = Kinosternoidea⁷ Hay 1892 mud turtle Including UCMP 173422, listed by Holroyd & Hutchison (2002)
Testudines - Panchelydridae	? Chelydridae indet.⁷, Chelydridae indet.⁷ ? Chelydridae indet.⁷ turtle Chelydridae indet.⁷ turtle Including UCMP 173420, listed by Holroyd & Hutchison (2002)
Testudines - Panemydidae	Emydinae indet.⁸ Emydinae indet.⁸ Rafinesque 1815 turtle "near Pseudemys?"
Archosauria -	Archosauria indet.⁷ Archosauria indet.⁷ archosaur
Avetheropoda - Dromaeosauridae	Richardoestesia sp.⁷, Saurornitholestinae indet.¹¹ Richardoestesia sp.⁷ Currie et al. 1990 coelurosaur Saurornitholestinae indet.¹¹ Longrich and Currie 2009 coelurosaur UCMP 187054
Avetheropoda - Troodontidae	Troodontidae indet.⁷, Troodon sp.⁷, Pectinodon bakkeri² Troodontidae indet.⁷ Gilmore 1924 coelurosaur Troodon sp.⁷ Leidy 1856 coelurosaur Pectinodon bakkeri² Carpenter 1982 coelurosaur
Avetheropoda -	Paronychodon lacustris⁸, cf. Paronychodon lacustris¹³ Paronychodon lacustris⁸ Cope 1876 coelurosaur cf. Paronychodon lacustris¹³ Cope 1876 coelurosaur UCMP 73076
Aves
-	Aves indet.⁷ Aves indet.⁷ Linnaeus 1758 bird
Reptilia
Avetheropoda -	cf. Dryptosaurus sp.⁸ cf. Dryptosaurus sp.⁸ Marsh 1877 coelurosaur
Neornithischia - Thescelosauridae	Thescelosaurus sp.² Thescelosaurus sp.² Gilmore 1913 ornithischian
Crocodylia -	"Leidyosuchus sternbergi" = Borealosuchus sternbergii⁸, Brachychampsa montana⁸ "Leidyosuchus sternbergi" = Borealosuchus sternbergii⁸ Gilmore 1910 crocodilian Brachychampsa montana⁸ Gilmore 1911 crocodilian
Squamata - Xenosauridae	Exostinus lancensis⁸ Exostinus lancensis⁸ Gilmore 1928 squamates
Squamata -	Paraderma bogerti⁸, Parasaniwa wyomingensis⁸, Varanoidea indet.¹⁰, Colpodontosaurus cracens⁸ Paraderma bogerti⁸ Estes 1964 squamates Parasaniwa wyomingensis⁸ Gilmore 1928 squamates Varanoidea indet.¹⁰ Gray 1827 squamates Colpodontosaurus cracens⁸ Estes 1964 squamates
Squamata - Anguidae	"Peltosaurus piger" = Odaxosaurus piger⁸ "Peltosaurus piger" = Odaxosaurus piger⁸ Gilmore 1928 squamates
Squamata -	Coniophis precedens¹² Coniophis precedens¹² Marsh 1892 snake UCMP 53556‐53561 (vertebrae)
Squamata - Chamopsiidae	Haptosphenus placodon⁸, Leptochamops denticulatus⁸, Chamops segnis⁸, Meniscognathus altmani⁸ Haptosphenus placodon⁸ Estes 1964 lizard Leptochamops denticulatus⁸ Gilmore 1928 lizard Chamops segnis⁸ Marsh 1892 lizard Meniscognathus altmani⁸ Estes 1964 lizard
Choristodera -	Champsosaurus sp.⁸ Champsosaurus sp.⁸ Cope 1876 choristodere
Mammalia
Cimolesta - Cimolestidae	Cimolestes incisus⁶ Cimolestes incisus⁶ Marsh 1889 eutherian
Leptictida - Gypsonictopidae	Gypsonictops hypoconus⁶ Gypsonictops hypoconus⁶ Simpson 1927 eutherian
Theriiformes -	"Alphadon lulli" = Protalphadon lulli⁴ "Alphadon lulli" = Protalphadon lulli⁴ Clemens 1966 metatherian
Theriiformes - Peradectidae	Pediomys elegans⁵, "Pediomys hatcheri" = Aletridelphys hatcheri⁵, "Pediomys florencae" = Aletridelphys florencae⁵, "Pediomys krejcii" = Leptalestes krejcii⁵, "Pediomys cooki" = Leptalestes cooki⁵ Pediomys elegans⁵ Marsh 1889 metatherian "Pediomys hatcheri" = Aletridelphys hatcheri⁵ Osborn 1898 metatherian "Pediomys florencae" = Aletridelphys florencae⁵ Clemens 1966 metatherian "Pediomys krejcii" = Leptalestes krejcii⁵ Clemens 1966 metatherian "Pediomys cooki" = Leptalestes cooki⁵ Clemens 1966 metatherian
Theriiformes - Herpetotheriidae	"Nortedelphys intermedius" = Nortedelphys jasoni³ "Nortedelphys intermedius" = Nortedelphys jasoni³ Storer 1991 marsupial
Didelphimorphia - Alphadontidae	Alphadon marshi⁵, "Alphadon rhaister n. sp." = Turgidodon rhaister⁴ Alphadon marshi⁵ Simpson 1927 marsupial "Alphadon rhaister n. sp." = Turgidodon rhaister⁴ Clemens 1966 marsupial
Didelphimorphia - Stagodontidae	Didelphodon vorax⁵ Didelphodon vorax⁵ Marsh 1889 marsupial
Multituberculata - Cimolomyidae	"Dipriodon robustus n. sp." = Meniscoessus robustus, Cimolomys gracilis⁴ "Dipriodon robustus n. sp." = Meniscoessus robustus Marsh 1889 multituberculate Cimolomys gracilis⁴ Marsh 1889 multituberculate present in both UCMP and Hatcher collections
Multituberculata - Neoplagiaulacidae	Mesodma sp.¹, Mesodma thompsoni⁴ Mesodma sp.¹ Jepsen 1940 multituberculate Mesodma thompsoni⁴ Clemens 1964 multituberculate
Multituberculata - Cimolodontidae	Cimolodon nitidus⁴ Cimolodon nitidus⁴ Marsh 1889 multituberculate

¹Alroy 2002; ²Carpenter 1982; ³Case et al. 2004; ⁴Clemens 1964; ⁵Clemens 1966; ⁶Clemens 1973; ⁷Database 2005; ⁸Estes 1964; ⁹Estes and Sanchíz 1982; ¹⁰Gao and Fox 1996; ¹¹Larson and Currie 2013; ¹²Longrich et al. 2012; ¹³Sankey 2008