Mammal Quarry No. 2 (UCMP V-5815) (Cretaceous of the United States)

Also known as Lance Quarry 2, Hatcher No. 2

Where: Niobrara County, Wyoming (43.1° N, 104.5° W: paleocoordinates 49.6° N, 79.4° W)

• coordinate estimated from map

• small collection-level geographic resolution

When: Lance Formation, Lancian (70.6 - 66.0 Ma)

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; sandstone

• "cross-stratified sandstone"

Size class: mesofossils

Collected by J. B. Hatcher, W. A. Clemens in 1889, 1958; reposited in the UCMP, YPM

Collection methods: bulk, sieve,

Primary reference: R. S. Lull. 1915. The mammals and horned dinosaurs of the Lance Formation of Niobrara County, Wyoming. The American Journal of Science, series 4 40(238):319-348 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14566: authorized by John Alroy, entered by John Alroy on 26.07.1994, edited by Matthew Carrano, Roger Benson and Patricia Holroyd

Creative Commons license: CC BY (attribution)

Taxonomic list

Gastropoda
 Sorbeoconcha - Hydrobiidae
Reesidella sp.8 Yen 1952 snail
Bivalvia
 Unionida - Unionidae
Unionidae indet.8 Rafinesque 1820 freshwater mussel
Chondrichthyes
 Hybodontiformes - Lonchidiidae
Lonchidion selachos8 Estes 1964 elasmobranch
Osteichthyes
 Acipenseriformes - Acipenseridae
Acipenser eruciferus8 Cope 1876 sturgeon
Amphibia
 Salientia - Pelobatidae
Eopelobates sp.9 Parker 1929 spadefoot toad
 Proteida - Batrachosauroididae
Opisthotriton kayi8, "Prodesmodon copei" = Prodesmodon copei8, "Cuttysarkus mcnallyi" = Prodesmodon copei8
Opisthotriton kayi8 Auffenberg 1961 salamander
"Prodesmodon copei" = Prodesmodon copei8 Estes 1964 salamander
"Cuttysarkus mcnallyi" = Prodesmodon copei8 Estes 1964 salamander
 Caudata - Urodela
Habrosaurus dilatus8 Gilmore 1928 siren
Scapherpeton tectum8 Cope 1876 salamander
Mammalia
 Multituberculata - Cimolomyidae
"Dipriodon robustus n. sp." = Meniscoessus robustus, Cimolomys gracilis4
"Dipriodon robustus n. sp." = Meniscoessus robustus Marsh 1889 multituberculate
Cimolomys gracilis4 Marsh 1889 multituberculate
present in both UCMP and Hatcher collections
 Multituberculata - Cimolodontidae
Cimolodon nitidus4 Marsh 1889 multituberculate
 Multituberculata - Neoplagiaulacidae
Mesodma sp.1 Jepsen 1940 multituberculate
Mesodma thompsoni4 Clemens 1964 multituberculate
 Theriamorpha - Gypsonictopidae
Gypsonictops hypoconus6 Simpson 1927 eutherian
 Cimolesta - Cimolestidae
Cimolestes incisus6 Marsh 1889 eutherian
 Theriamorpha - Herpetotheriidae
"Nortedelphys intermedius" = Nortedelphys jasoni3
"Nortedelphys intermedius" = Nortedelphys jasoni3 Storer 1991 marsupial
 Theriamorpha - Pediomyidae
Pediomys elegans5, "Pediomys cooki" = Leptalestes cooki5, "Pediomys krejcii" = Leptalestes krejcii5, "Pediomys hatcheri" = Protolambda hatcheri5, "Pediomys florencae" = Protolambda florencae5
Pediomys elegans5 Marsh 1889 metatherian
"Pediomys cooki" = Leptalestes cooki5 Clemens 1966 metatherian
"Pediomys krejcii" = Leptalestes krejcii5 Clemens 1966 metatherian
"Pediomys hatcheri" = Protolambda hatcheri5 Osborn 1898 metatherian
"Pediomys florencae" = Protolambda florencae5 Clemens 1966 metatherian
 Theriamorpha - Stagodontidae
Didelphodon vorax5 Marsh 1889 metatherian
 Theriamorpha - Alphadontidae
Alphadon marshi5, "Alphadon rhaister n. sp." = Turgidodon rhaister4
Alphadon marshi5 Simpson 1927 metatherian
"Alphadon rhaister n. sp." = Turgidodon rhaister4 Clemens 1966 metatherian
 Theriamorpha -
"Alphadon lulli" = Protalphadon lulli4
"Alphadon lulli" = Protalphadon lulli4 Clemens 1966 metatherian
Reptilia
 Testudinata -
Compsemys victa8 Leidy 1856 turtle
 Testudinata - Baenidae
Baenidae indet.7 Cope 1882 turtle
Including UCMP 173424, listed by Holroyd & Hutchison (2002)
 Testudines -
Cryptodira indet.7 Cope 1868 turtle
 Testudines - Chelydridae
? Chelydridae indet.7 Swainson 1839 turtle
Chelydridae indet.7 Swainson 1839 turtle
Including UCMP 173420, listed by Holroyd & Hutchison (2002)
 Testudines - Kinosternidae
"Kinosternidae indet." = Kinosternoidea7
"Kinosternidae indet." = Kinosternoidea7 Hay 1892 mud turtle
Including UCMP 173422, listed by Holroyd & Hutchison (2002)
 Testudines - Emydidae
Emydinae indet.8 Rafinesque 1815 turtle
"near Pseudemys?"
 Testudines - Trionychidae
Trionychidae indet.7 Gray 1825 softshell turtle
Including UCMP 173026, listed by Holroyd & Hutchison (2002)
 Squamata -
Coniophis precedens12 Marsh 1892 snake
UCMP 53556‐53561 (vertebrae)
Paraderma bogerti8 Estes 1964 squamates
Parasaniwa wyomingensis8 Gilmore 1928 squamates
Varanoidea indet.10 Gray 1827 squamates
Colpodontosaurus cracens8 Estes 1964 squamates
 Squamata - Anguidae
"Peltosaurus piger" = Odaxosaurus piger8
"Peltosaurus piger" = Odaxosaurus piger8 Gilmore 1928 squamates
 Squamata - Xenosauridae
Exostinus lancensis8 Gilmore 1928 squamates
 Squamata - Chamopsiidae
Meniscognathus altmani8 Estes 1964 squamates
Haptosphenus placodon8 Estes 1964 squamates
Leptochamops denticulatus8 Gilmore 1928 squamates
Chamops segnis8 Marsh 1892 squamates
 Archosauria -
 Crocodylia - Alligatoridae
Brachychampsa montana8 Gilmore 1911 crocodilian
 Loricata -
"Leidyosuchus sternbergi" = Borealosuchus sternbergii8
"Leidyosuchus sternbergi" = Borealosuchus sternbergii8 Gilmore 1910 crocodilian
 Ornithischia -
Thescelosaurus sp.2 Gilmore 1913 ornithopod
 Theropoda - Troodontidae
Troodontidae indet.7 Gilmore 1924 maniraptoran
Troodon sp.7 Leidy 1856 maniraptoran
Pectinodon bakkeri2 Carpenter 1982 maniraptoran
 Theropoda - Dromaeosauridae
Richardoestesia sp.7 Currie et al. 1990 maniraptoran
Saurornitholestinae indet.11 Longrich and Currie 2009 maniraptoran
UCMP 187054
 Theropoda -
Paronychodon lacustris8 Cope 1876 maniraptoran
cf. Paronychodon lacustris13 Cope 1876 maniraptoran
UCMP 73076
Aves
  -
Aves indet.7 Linnaeus 1758 bird
Reptilia
 Theropoda -
cf. Dryptosaurus sp.8 Marsh 1877 coelurosaur
 Choristodera -
Champsosaurus sp.8 Cope 1876 choristodere
Actinopteri
 Lepisosteiformes - Lepisosteidae
Lepisosteus occidentalis8 Leidy 1856 gar
 Amiiformes - Amiidae
"Kindleia fragosa" = Cyclurus fragosus8 Jordan 1927 bowfin
cf. Protamia sp.8 Leidy 1873 bowfin
Melvius thomasi7 Bryant 1987 bowfin
 Esociformes - Esocidae
Estesesox foxi14 Wilson et al. 1992
 Elopiformes - Elopidae
Elopidae indet.8 Bonaparte 1846
 Albuliformes - Albulidae
Albulidae indet.8 Bleeker 1859 bonefish
 Aspidorhynchiformes - Aspidorhynchidae