Mammal Quarry No. 2 (UCMP V-5815) (Cretaceous of the United States)

Also known as Lance Quarry 2, Hatcher No. 2

Where: Niobrara County, Wyoming (43.1° N, 104.5° W)

• Paleocoordinates: 53.6° N, 65.8° W (Wright 2013)

• coordinate estimated from map

• small collection-level geographic resolution

When: Lance Formation, Lancian (72.2 - 66.0 Ma)

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; sandstone

• "cross-stratified sandstone"

Size class: mesofossils

Collected by J. B. Hatcher, W. A. Clemens in 1889, 1958; reposited in the UCMP, YPM

Collection methods: bulk, sieve,

Primary reference: R. S. Lull. 1915. The mammals and horned dinosaurs of the Lance Formation of Niobrara County, Wyoming. The American Journal of Science, series 4 40(238):319-348 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14566: authorized by John Alroy, entered by John Alroy on 26.07.1994, edited by Matthew Carrano, Roger Benson and Patricia Holroyd

Creative Commons license: CC0 (CC0)

Taxonomic list

Chondrichthyes
 Hybodontiformes - Lonchidiidae
Lonchidion selachos9 Estes 1964 elasmobranch
Actinopteri
 Lepisosteiformes - Lepisosteidae
Lepisosteus occidentalis9 Leidy 1856 gar
 Amiiformes - Amiidae
Melvius thomasi2 Bryant 1988 bowfin
cf. Protamia sp.9 Leidy 1873 bowfin
"Kindleia fragosa" = Cyclurus fragosus9 Jordan 1927 bowfin
 Esociformes - Esocidae
Estesesox foxi15 Wilson et al. 1992
 Albuliformes - Albulidae
Albulidae indet.9 Bleeker 1859 bonefish
 Elopiformes - Elopidae
Elopidae indet.9 Bonaparte 1846
 Aspidorhynchiformes - Aspidorhynchidae
Amphibia
 Salientia - Pelobatidae
Eopelobates sp.10 Parker 1929 spadefoot toad
 Proteida - Batrachosauroididae
"Prodesmodon copei" = Prodesmodon copei9, "Cuttysarkus mcnallyi" = Prodesmodon copei9, Opisthotriton kayi9
"Prodesmodon copei" = Prodesmodon copei9 Estes 1964 salamander
"Cuttysarkus mcnallyi" = Prodesmodon copei9 Estes 1964 salamander
Opisthotriton kayi9 Auffenberg 1961 salamander
 Caudata - Urodela
Habrosaurus dilatus9 Gilmore 1928 siren
Scapherpeton tectum9 Cope 1876 salamander
Reptilia
 Archosauria -
 Crocodylia -
"Leidyosuchus sternbergi" = Borealosuchus sternbergii9
"Leidyosuchus sternbergi" = Borealosuchus sternbergii9 Gilmore 1910 crocodilian
 Crocodylia - Alligatoridae
Brachychampsa montana9 Gilmore 1911 crocodilian
 Saurischia -
Paronychodon lacustris9 Cope 1876 maniraptoran
cf. Paronychodon lacustris14 Cope 1876 maniraptoran
UCMP 73076
 Saurischia - Troodontidae
Troodontidae indet.4 Gilmore 1924 maniraptoran
Troodon sp.4 Leidy 1856 maniraptoran
Pectinodon bakkeri3 Carpenter 1982 maniraptoran
 Saurischia - Dromaeosauridae
Saurornitholestinae indet.12 Longrich and Currie 2009 maniraptoran
UCMP 187054
Richardoestesia sp.4 Currie et al. 1990 maniraptoran
Aves
  -
Aves indet.4 Linnaeus 1758 bird
Reptilia
 Saurischia -
cf. Dryptosaurus sp.9 Marsh 1877 coelurosaur
 Ornithischia -
Thescelosaurus sp.3 Gilmore 1913 ornithopod
 Choristodera -
Champsosaurus sp.9 Cope 1876 choristodere
 Squamata - Xenosauridae
Exostinus lancensis9 Gilmore 1928 squamates
 Squamata - Anguidae
"Peltosaurus piger" = Odaxosaurus piger9
"Peltosaurus piger" = Odaxosaurus piger9 Gilmore 1928 squamates
 Squamata -
Varanoidea indet.11 Gray 1827 squamates
Parasaniwa wyomingensis9 Gilmore 1928 squamates
Colpodontosaurus cracens9 Estes 1964 squamates
Paraderma bogerti9 Estes 1964 squamates
Coniophis precedens13 Marsh 1892 snake
UCMP 53556‐53561 (vertebrae)
 Squamata - Chamopsiidae
Haptosphenus placodon9 Estes 1964 squamates
Leptochamops denticulatus9 Gilmore 1928 squamates
Meniscognathus altmani9 Estes 1964 squamates
Chamops segnis9 Marsh 1892 squamates
 Testudines -
Cryptodira indet.4 Cope 1868 turtle
 Testudines - Chelydridae
? Chelydridae indet.4 Swainson 1839 turtle
Chelydridae indet.4 Swainson 1839 turtle
Including UCMP 173420, listed by Holroyd & Hutchison (2002)
 Testudines - Emydidae
Emydinae indet.9 Rafinesque 1815 turtle
"near Pseudemys?"
 Testudines - Kinosternidae
"Kinosternidae indet." = Kinosternoidea4
"Kinosternidae indet." = Kinosternoidea4 Hay 1892 mud turtle
Including UCMP 173422, listed by Holroyd & Hutchison (2002)
 Testudines - Trionychidae
Trionychidae indet.4 Gray 1825 softshell turtle
Including UCMP 173026, listed by Holroyd & Hutchison (2002)
 Testudinata - Baenidae
Baenidae indet.4 Cope 1882 turtle
Including UCMP 173424, listed by Holroyd & Hutchison (2002)
 Testudinata -
Compsemys victa9 Leidy 1856 turtle
Mammalia
 Multituberculata - Cimolomyidae
"Dipriodon robustus n. sp." = Meniscoessus robustus, Cimolomys gracilis6
"Dipriodon robustus n. sp." = Meniscoessus robustus Marsh 1889 multituberculate
Cimolomys gracilis6 Marsh 1889 multituberculate
present in both UCMP and Hatcher collections
 Multituberculata - Neoplagiaulacidae
Mesodma sp.1 Jepsen 1940 multituberculate
Mesodma thompsoni6 Clemens 1964 multituberculate
 Multituberculata - Cimolodontidae
Cimolodon nitidus6 Marsh 1889 multituberculate
 Leptictida - Gypsonictopidae
Gypsonictops hypoconus8 Simpson 1927 eutherian
 Cimolesta - Cimolestidae
Cimolestes incisus8 Marsh 1889 eutherian
 Tribosphenida - Stagodontidae
Didelphodon vorax7 Marsh 1889 metatherian
 Tribosphenida - Herpetotheriidae
"Nortedelphys intermedius" = Nortedelphys jasoni5
"Nortedelphys intermedius" = Nortedelphys jasoni5 Storer 1991 marsupial
 Tribosphenida - Alphadontidae
"Alphadon rhaister n. sp." = Turgidodon rhaister6, Alphadon marshi7
"Alphadon rhaister n. sp." = Turgidodon rhaister6 Clemens 1966 metatherian
Alphadon marshi7 Simpson 1927 metatherian
 Tribosphenida -
"Alphadon lulli" = Protalphadon lulli6
"Alphadon lulli" = Protalphadon lulli6 Clemens 1966 metatherian
 Tribosphenida - Pediomyidae
Pediomys elegans7, "Pediomys florencae" = Protolambda florencae7, "Pediomys hatcheri" = Protolambda hatcheri7, "Pediomys cooki" = Leptalestes cooki7, "Pediomys krejcii" = Leptalestes krejcii7
Pediomys elegans7 Marsh 1889 metatherian
"Pediomys florencae" = Protolambda florencae7 Clemens 1966 metatherian
"Pediomys hatcheri" = Protolambda hatcheri7 Osborn 1898 metatherian
"Pediomys cooki" = Leptalestes cooki7 Clemens 1966 metatherian
"Pediomys krejcii" = Leptalestes krejcii7 Clemens 1966 metatherian
Osteichthyes
 Acipenseriformes - Acipenseridae
Acipenser eruciferus9 Cope 1876 sturgeon
Gastropoda
 Sorbeoconcha - Hydrobiidae
Reesidella sp.9 Yen 1952 snail
Bivalvia
 Unionida - Unionidae
Unionidae indet.9 Rafinesque 1820 freshwater mussel