Milia 5 (=MIL 5) (Pliocene of Greece)

Also known as Arap Tatsera

Where: Grevena, Greece (40.2° N, 21.5° E: paleocoordinates 40.1° N, 21.4° E)

• coordinate stated in text

When: MN 16a mammal zone, Late/Upper Pliocene (3.6 - 2.6 Ma)

Environment/lithology: fluvial; unlithified, medium-grained, brown, yellow sandstone

Size classes: macrofossils, mesofossils

Collection methods: quarrying, surface (in situ), sieve,

Primary reference: E. Vlachos, E. Tsoukala, E. Cregut-Bonnoure, C. Guerin, and D. Mol. 2018. The Paradise Lost of Milia (Grevena, Greece; Late Pliocene, Early Villafranchian, MN15/MN16a): faunal composition and diversity. Quaternary MDPI 1(13):1-18 [E. Vlachos/E. Vlachos/E. Vlachos]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 185859: authorized by Evangelos Vlachos, entered by Evangelos Vlachos on 22.05.2017

Creative Commons license: CC BY (attribution)

Taxonomic list

 Testudines - Pantestudinidae
Titanochelon sp.4 Pérez-García and Vlachos 2014 turtle
possible coracoid
Testudo brevitesta4 Vlachos and Tsoukala 2016 turtle
shell fragments
 Testudines - Pangeoemydidae
Mauremys sp.4 Gray 1870 pond turtle
shell fragments
 Rodentia - Hystricidae
Hystrix refossa2 porcupine
 Perissodactyla - Rhinocerotidae
 Perissodactyla - Tapiridae
Tapirus arvernensis Croizet and Jobert 1828 tapir
 Perissodactyla - Equidae
Hipparion sp. de Christol 1832 hipparionine horse
 Carnivora - Ursidae
Ursus etruscus Cuvier 1823 bear
Agriotherium sp. Wagner 1837 bear
 Carnivora - Felidae
 Proboscidea - Gomphotheriidae
Anancus arvernensis3 Croizet and Jobert 1828 gomphothere
 Proboscidea - Mammutidae
Mammut borsoni3 Hays 1834 mastodon
partial skeleton, including two complete tusks, the longest in the world
 Ungulata - Suidae
 Ungulata - Bovidae
Grevenobos antiquus n. gen. n. sp.1 Cregut-Bonnoure and Tsoukala 2017 antelope
 Ungulata - Cervidae
Croizetoceros ramosus Croizet and Jobert 1828 deer