PBDB

Bore Kevelaer, depth 70m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.6° N, 6.3° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage or Reinbek-Tichelh. stage

• bed-level stratigraphic resolution

Environment/lithology: marine; unlithified, fine-grained, brown sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1994

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 62887: authorized by Martin Aberhan, entered by Sabine Nürnberg on 01.08.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Anthozoa
Scleractinia - Flabellidae	Flabellum sp., Flabellum pompeckji, Flabellum tuberculatum, Flabellum cristatum Flabellum sp. Lesson 1831 stony coral Flabellum pompeckji Krejci 1926 stony coral Flabellum tuberculatum Keferstein 1859 stony coral Flabellum cristatum Milne-Edwards and Haime 1851 stony coral
Scleractinia - Caryophylliidae	Deltocyathus reussi Deltocyathus reussi stony coral
Scleractinia - Micrabaciidae	Stephanophyllia imperialis Stephanophyllia imperialis Michelin 1841 stony coral
Polychaeta
Sabellida - Serpulidae	Serpulidae indet.¹ Serpulidae indet.¹ Rafinesque 1835 Serpulidae indet. D
Gymnolaemata
Cheilostomata - Membraniporidae	Reussirella haidingeri¹ Reussirella haidingeri¹
Cheilostomata - Lunulitidae	Lunulites voigti¹, Lunulites burdigalensis¹ Lunulites voigti¹ Lunulites burdigalensis¹
Bivalvia
Arcida - Limopsidae	Limopsis (Limopsis) aurita¹, "Limopsis (Pectunculina) anomala" = Limopsis¹, Limopsis (Pectunculina) lamellata¹ Limopsis (Limopsis) aurita¹ Brocchi 1814 clam "Limopsis (Pectunculina) anomala" = Limopsis¹ clam Limopsis (Pectunculina) lamellata¹ Lehmann 1885 clam
Arcida - Arcidae	Scapharca (Scapharca) diluvii¹ Scapharca (Scapharca) diluvii¹ Lamarck 1805 clam
Pectinida - Pectinoidae	Pseudamussium lilli¹, Korobkovia woodi¹, Aequipecten seniensis¹ Pseudamussium lilli¹ Pusch 1837 scallop Korobkovia woodi¹ scallop Aequipecten seniensis¹ scallop
Ostreida - Pinnidae	Atrina pectinata¹ Atrina pectinata¹ Linnaeus 1758 oyster
Carditida - Astartidae	Goodallia (Goodallia) angulata¹, Astarte (Astarte) radiata¹ Goodallia (Goodallia) angulata¹ Lehmann 1885 clam Astarte (Astarte) radiata¹ Nyst and Westendorp 1839 clam Astarte radiata fa. radiata 28 specimens, Astarte radiata fa. convexior 33 specimens, Astarte radiata fa. goldfussi 2 specimens
Carditida - Carditidae	Cyclocardia sp.¹ Cyclocardia sp.¹ Conrad 1867 clam
Lucinida - Lucinidae	"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti¹ "Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti¹ Nyst 1861 clam C. droueti droueti
Pandorida - Lyonsiidae	Lyonsia (Lyonsia) aff. norwegica¹ Lyonsia (Lyonsia) aff. norwegica¹ clam
Pholadida - Corbulidae	Corbula (Varicorbula) gibba¹ Corbula (Varicorbula) gibba¹ Oliva 1792 clam C. (V.) gibba gibba
Cardiida - Semelidae	sensu lato Abra lehmanni¹ sensu lato Abra lehmanni¹ abra
Cardiida - Tellinidae	"Macoma elliptica" = Laciolina magna¹ "Macoma elliptica" = Laciolina magna¹ Spengler 1798 tellin clam
Cardiida - Cardiidae	Habecardium dingdense¹, Parvicardium straeleni¹, Parvicardium scabrum¹ Habecardium dingdense¹ Lehmann 1885 cockle Parvicardium straeleni¹ cockle Parvicardium scabrum¹ Philippi 1844 cockle
Cardiida - Mactridae	Spisula elliptica¹ Spisula elliptica¹ Brown 1827 clam
Cardiida - Glossidae	Glossus (Glossus) lunulatus¹ Glossus (Glossus) lunulatus¹ clam
Cardiida - Veneridae	"Gouldia minima" = Gouldia (Gouldia) minima¹, Venus (Ventricoloidea) multilamella¹ "Gouldia minima" = Gouldia (Gouldia) minima¹ Montagu 1803 venus clam Venus (Ventricoloidea) multilamella¹ Lamarck 1818 venus clam V. (V.) multilamella multilamella
Nuculida - Nuculidae	Nuculoma haesendoncki¹ Nuculoma haesendoncki¹ nut clam N. haesendoncki hanseata
Protobranchia - Nuculanidae	"Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ "Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ Nyst and Westendorp 1839 pointed nut clam
Protobranchia - Yoldiidae	"Portlandia (Yoldiella) pygmaea" = Yoldiella¹, Yoldia (Yoldia) glaberrima¹ "Portlandia (Yoldiella) pygmaea" = Yoldiella¹ clam Yoldia (Yoldia) glaberrima¹ von Münster 1837 clam
Scaphopoda
Dentaliida - Dentaliidae	Lentigodentalium pseudomutabile¹, "Pseudantalis aturensis" = Dentalium (Pseudantilis)¹ Lentigodentalium pseudomutabile¹ tusk shell "Pseudantalis aturensis" = Dentalium (Pseudantilis)¹ tusk shell
Gadilida - Gadilidae	Gadila dingdenensis¹ Gadila dingdenensis¹ tusk shell
Gastropoda
Heterostropha - Ringiculidae	Ringicula (Ringiculina) buccinea³ Ringicula (Ringiculina) buccinea³ Sowerby 1823 snail
Heterostropha - Acteonidae	Crenilabium terebelloides³, Acteon (Acteon) semistriatus³ Crenilabium terebelloides³ snail Acteon (Acteon) semistriatus³ snail spelled as 'Actaeon'
Heterostropha - Pyramidellidae	Turbonilla (Turbonilla) gastaldi³, Turbonilla (Turbonilla) koeneniana³, Odostomia (Brachystomia) sp.³, Odostomia (Megastomia) cf. conoidea³, Eulimella (Eulimella) acicula³, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³, Pyramidella (Voluspa) plicosa³, Syrnola hoernesi³ Turbonilla (Turbonilla) gastaldi³ snail sensu JANSSEN Turbonilla (Turbonilla) koeneniana³ snail sensu JANSSEN Odostomia (Brachystomia) sp.³ Fleming 1817 snail Odostomia (Megastomia) cf. conoidea³ Brocchi 1814 snail Eulimella (Eulimella) acicula³ snail "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³ snail sensu JANSSEN Pyramidella (Voluspa) plicosa³ snail Syrnola hoernesi³ snail
Neotaenioglossa - Calyptraeidae	Calyptraea (Calyptraea) chinensis² Calyptraea (Calyptraea) chinensis² Linnaeus 1758 slipper shell
Sorbeoconcha - Turritellidae	Turritella subangulata² Turritella subangulata² turret shell
Sorbeoconcha - Cerithiidae	"Bittium (Bittium) spina" = Bittium spina² "Bittium (Bittium) spina" = Bittium spina² Partsch 1856 cerith snail
Sorbeoconcha - Aporrhaidae	Aporrhais (Aporrhais) alata² Aporrhais (Aporrhais) alata² snail
Neogastropoda - Olividae	Oliva (Strephona) dufresnei² Oliva (Strephona) dufresnei² Basterot 1825 olive snail
Neogastropoda - Ancillariidae	"Ancilla (Baryspira) obsoleta" = Amalda obsoleta² "Ancilla (Baryspira) obsoleta" = Amalda obsoleta² Brocchi 1814 snail
Neogastropoda - Nassariidae	Nassarius bocholtensis², Nassarius tenuistriatus², Nassarius voorthuyseni², Nassarius karinae² Nassarius bocholtensis² snail Nassarius tenuistriatus² snail Nassarius voorthuyseni² snail Nassarius karinae² Wienrich 2001 snail
Neogastropoda - Fasciolariidae	Streptodictyon sexcostatus² Streptodictyon sexcostatus² snail
Neogastropoda - Buccinidae	Phos (Phos) decussatus² Phos (Phos) decussatus² true whelk
Neogastropoda - Columbellidae	Mitrella (Macrurella) nassoides², Anachis (Costoanachis) hosiusi², Anachis (Costoanachis) terebralis² Mitrella (Macrurella) nassoides² snail Anachis (Costoanachis) hosiusi² snail Anachis (Costoanachis) terebralis² snail
Neogastropoda - Muricidae	"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta², "Murex (Haustellum) inornatus" = Haustellum², Typhis (Typhis) ? pungens², Lyrotyphis (Eotyphis) fistulosus² "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta² Bronn 1831 murex snail "Murex (Haustellum) inornatus" = Haustellum² murex snail Typhis (Typhis) ? pungens² Solander in Brander and Brander 1766 murex snail Lyrotyphis (Eotyphis) fistulosus² murex snail
Neogastropoda - Drilliidae	Splendrillia selenkae³ Splendrillia selenkae³ snail
Neogastropoda - Clathurellidae	Pleurotomoides luisae³, Pleurotomoides simplex³, ? Ceritoturris hannoverana³ Pleurotomoides luisae³ snail Pleurotomoides simplex³ snail ? Ceritoturris hannoverana³ snail
Neogastropoda - Terebridae	Strioterebrum hoernesi³, Strioterebrum ronaldi³ Strioterebrum hoernesi³ auger snail Strioterebrum ronaldi³ auger snail
Neogastropoda - Clavatulidae	Clavatula boreointerrupta³ Clavatula boreointerrupta³ snail
Neogastropoda - Borsoniidae	Asthenotoma pannoides³, Asthenotoma pannus³, Bathytoma jugleri³ Asthenotoma pannoides³ snail Asthenotoma pannus³ snail Bathytoma jugleri³ snail
Neogastropoda - Mangeliidae	sensu lato Mangelia andersoni n. sp.³, Sorgenfreispira "sp. 2"³, Sorgenfreispira tenella³, Benthomangelia aff. obtusangula³ sensu lato Mangelia andersoni n. sp.³ Wienrich and Janssen 2007 snail Sorgenfreispira "sp. 2"³ Moroni 1979 snail Sorgenfreispira tenella³ Conrad 1848 snail Benthomangelia aff. obtusangula³ snail
Neogastropoda - Turridae	Fusiturris aquensis³, Fusiturris duchastelii³, Nitidiclavus maitrejus³, Boreodrillia hosiusi³, sensu lato Unedogemmula boreoturricula³, sensu lato Unedogemmula hanseata³, sensu lato Unedogemmula stoffelsii³, sensu lato Inquisitor borealis³, Gemmula (Gemmula) denticula³, Gemmula (Gemmula) zimmermanni³ Fusiturris aquensis³ turrid Fusiturris duchastelii³ turrid Nitidiclavus maitrejus³ turrid Boreodrillia hosiusi³ turrid sensu lato Unedogemmula boreoturricula³ turrid sensu lato Unedogemmula hanseata³ turrid sensu lato Unedogemmula stoffelsii³ turrid sensu lato Inquisitor borealis³ turrid Gemmula (Gemmula) denticula³ turrid G. denticula borealis Gemmula (Gemmula) zimmermanni³ turrid
Neogastropoda - Conidae	"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ "Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ cone shell
Neogastropoda - Raphitomidae	Teretia anceps³, sensu lato Teretia "sp. 2"³, Mioawateria andersoni³, Metuonella grippi³ Teretia anceps³ snail sensu lato Teretia "sp. 2"³ Norman 1888 snail Mioawateria andersoni³ snail Metuonella grippi³ snail
Neogastropoda - Costellariidae	Orthosurcula steinvorthi³ Orthosurcula steinvorthi³ ribbed mitre
Neogastropoda - Cancellariidae	"Trigonostoma (Ventrilia) pouwi" = Ventrilia², Cancellaria (Merica) contorta², Babylonella fusiformis², Trigonostoma (Trigonostoma) aperta², Trigonostoma (Trigonostoma) extractrix², Trigonostoma (Trigonostoma) cf. spinifera², Sveltia varicosa² "Trigonostoma (Ventrilia) pouwi" = Ventrilia² snail Cancellaria (Merica) contorta² snail C. contorta gelriana Babylonella fusiformis² snail Trigonostoma (Trigonostoma) aperta² snail Trigonostoma (Trigonostoma) extractrix² snail Trigonostoma (Trigonostoma) cf. spinifera² snail Sveltia varicosa² snail S. varicosa paucicostata
Sorbeoconcha - Cassidae	"Semicassis (Echinophoria) bicoronata" = Echinophoria², "Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata² "Semicassis (Echinophoria) bicoronata" = Echinophoria² snail "Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata² Sacco 1890 snail
Sorbeoconcha - Triviidae	Erato (Erato) germanica² Erato (Erato) germanica² Schilder 1929 snail
Sorbeoconcha - Xenophoridae	Xenophora deshayesi² Xenophora deshayesi² Michelotti 1847 carrier shell
Sorbeoconcha - Vitrinellidae	Circulus praecedens², Circulus hennei² Circulus praecedens² snail C. praecedens for. praecedens Circulus hennei² Glibert 1952 snail
Sorbeoconcha - Rissoidae	sensu lato Alvania holsatica², Alvania (Alvania) basisulcata² sensu lato Alvania holsatica² snail Alvania (Alvania) basisulcata² snail
Sorbeoconcha - Eulimidae	Balcis (Balcis) alba², Eulima (Eulima) bilineata², Eulima (Eulima) glabra² Balcis (Balcis) alba² snail Eulima (Eulima) bilineata² snail Eulima (Eulima) glabra² snail
Sorbeoconcha - Naticidae	Naticidae indet.², "Natica sensu lato tigrina" = Cochlis tigrina², Tectonatica miopusilla², Euspira edithae², Euspira staringi², Euspira helicina² Naticidae indet.² Guilding 1834 moon snail Naticidae juv. et indet. "Natica sensu lato tigrina" = Cochlis tigrina² Röding 1798 moon snail Tectonatica miopusilla² moon snail Euspira edithae² moon snail Euspira staringi² moon snail Euspira helicina² Brocchi 1814 moon snail E. aggr. helicina
Sorbeoconcha - Triphoridae	Norephora (Norephora) fritschi² Norephora (Norephora) fritschi² snail
Sorbeoconcha - Triforidae	Cerithiella (Cerithiella) genei² Cerithiella (Cerithiella) genei² snail
Sorbeoconcha - Epitoniidae	Opalia (Pliciscala) pertusa², Cirsotrema (Cirsotrema) crassicostatum², "Amaea (Scalina) subreticula" = Scalina² Opalia (Pliciscala) pertusa² wentletrap Cirsotrema (Cirsotrema) crassicostatum² Deshayes 1839 wentletrap "Amaea (Scalina) subreticula" = Scalina² wentletrap
Opisthobranchia -	Roxania utriculus³ Roxania utriculus³ snail
Opisthobranchia - Retusidae	Volvulella (Volvulella) acuminata³, Retusa (Cylichnina) elongata³ Volvulella (Volvulella) acuminata³ snail Retusa (Cylichnina) elongata³ snail
Opisthobranchia - Cylichnidae	Scaphander grateloupi³, Cylichna (Cylichna) pseudoconvoluta³ Scaphander grateloupi³ snail Cylichna (Cylichna) pseudoconvoluta³ snail
Foraminifera
Textulariacea - Textulariidae	Bigenerina nodosaria, Textularia gramen Bigenerina nodosaria Textularia gramen dOrbigny 1846
Textulariacea - Valvulinidae	Martinottiella communis Martinottiella communis Huang et al. 2019
- Spiroplectamminidae	Spiroplectammina carinata Spiroplectammina carinata d'Orbigny 1846
- Buliminidae	Virgulina schreibersiana, Virgulina (Virgulinella) pertusa Virgulina schreibersiana Virgulina (Virgulinella) pertusa
Tubothalamea
Miliolida - Miliolidae	Sigmoilina tenuis Sigmoilina tenuis
Foraminifera
Discorbacea - Bagginidae	Cancris auriculus Cancris auriculus Fichtel and Moll 1798
Rotaliacea - Elphidiidae	Elphidium inflatum, Elphidium hauerinum Elphidium inflatum Elphidium hauerinum
Nonionacea - Nonionidae	Cribrononion subnodosum, Nonion boueanum, Nonion affine, Nonion perfossum Cribrononion subnodosum Nonion boueanum Nonion affine Nonion perfossum
Rotaliina - Cassidulinidae	Ceratobulimina haueri Ceratobulimina haueri
Asterigerinacea - Asterigerinidae	Asterigerina guerichi Asterigerina guerichi A. gürichi staeschi
Planorbulinacea - Cibicididae	Cibicides lobatulus, Cibicides pseudoungerianus, Cibicides dutemplei Cibicides lobatulus incl. 2 specimens C. lobatulus var. ornatus Cibicides pseudoungerianus Cushman 1922 Cibicides dutemplei dOrbigny 1846
Planorbulinacea - Planulinidae	Planulina austriaca Planulina austriaca
Buliminacea - Uvigerinidae	Uvigerina semiornata Uvigerina semiornata U. semiornata for. semiornata
- Nodosariidae	Lenticulina (Lenticulina) calcar, Lenticulina (Robulus) cultrata, Lenticulina (Lenticulina) inornata, Lenticulina (Robulus) similis, Lenticulina (Robulus) ornata Lenticulina (Lenticulina) calcar Lenticulina (Robulus) cultrata Lenticulina (Lenticulina) inornata Lenticulina (Robulus) similis Lenticulina (Robulus) ornata
Nodosariacea - Polymorphinidae	Guttulina problema, Globulina gibba, Pseudopolymorphina subnodosa, Sigmomorphina schwageri, Polymorphina charlottensis Guttulina problema d'Orbigny 1826 Globulina gibba incl. 1 specimen G. gibba for. punctata and 1 specimen G. gibba rugosa Pseudopolymorphina subnodosa Sigmomorphina schwageri Polymorphina charlottensis
Nodosariacea - Lagenidae	Dentalina intermittens, Dentalina bifurcata, Dentalina grossicosta, Dentalina acuticauda, Dentalina konincki, Dentalina pauperata, Lagena globosa, Lagena cf. emaciata, Lagena sulcata, Nodosaria bacillum, Nodosaria longiscata Dentalina intermittens Dentalina bifurcata Dentalina grossicosta Dentalina acuticauda Dentalina konincki Reuss 1861 Dentalina pauperata d'Orbigny 1846 Lagena globosa Lagena cf. emaciata Lagena sulcata Williamson 1848 Nodosaria bacillum Nodosaria longiscata d'Orbigny 1846

¹Wienrich 1999; ²Wienrich 2001; ³Wienrich and Janssen 2007