PBDB Collection

Bore Kevelaer, depth 70m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.6° N, 6.3° E)

• Paleocoordinates: 49.0° N, 8.5° E (Wright 2013)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage or Reinbek-Tichelh. stage

• bed-level stratigraphic resolution

Environment/lithology: marine; unlithified, fine-grained, brown sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1994

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 62887: authorized by Martin Aberhan, entered by Sabine Nürnberg on 01.08.2006

Creative Commons license: CC0 (CC0)

Taxonomic list

Show authors, comments, and common names

Globothalamea
Rotaliida - Nonionidae	Cribrononion subnodosum, Nonion boueanum, Nonion affine, Nonion perfossum Cribrononion subnodosum Nonion boueanum Nonion affine Nonion perfossum
Rotaliida - Cibicididae	Planulina austriaca, Cibicides pseudoungerianus, Cibicides dutemplei, Cibicides lobatulus Planulina austriaca Cibicides pseudoungerianus Cushman 1922 Cibicides dutemplei dOrbigny 1846 Cibicides lobatulus Walker and Jacob 1798 incl. 2 specimens C. lobatulus var. ornatus
Foraminifera
Rotalioidea - Elphidiidae	Elphidium inflatum, Elphidium hauerinum Elphidium inflatum Elphidium hauerinum
Nodosariata
Robertinida - Ceratobuliminidae	Ceratobulimina haueri Ceratobulimina haueri
Polymorphinida - Polymorphinidae	Sigmomorphina schwageri, Polymorphina charlottensis, Pseudopolymorphina subnodosa, Globulina gibba, Guttulina problema Sigmomorphina schwageri Polymorphina charlottensis Pseudopolymorphina subnodosa Globulina gibba d'Orbigny 1832 incl. 1 specimen G. gibba for. punctata and 1 specimen G. gibba rugosa Guttulina problema d'Orbigny 1826
Nodosariida - Nodosariidae	Lagena cf. emaciata, Lagena sulcata, Lagena globosa, Nodosaria bacillum, Nodosaria longiscata, Dentalina intermittens, Dentalina bifurcata, Dentalina grossicosta, Dentalina acuticauda, Dentalina konincki, Dentalina pauperata Lagena cf. emaciata Lagena sulcata Williamson 1848 Lagena globosa Montagu 1803 Nodosaria bacillum Nodosaria longiscata d'Orbigny 1846 Dentalina intermittens Dentalina bifurcata Dentalina grossicosta Dentalina acuticauda Dentalina konincki Reuss 1861 Dentalina pauperata d'Orbigny 1846
Vaginulinida - Vaginulinidae	Lenticulina (Lenticulina) inornata, Lenticulina (Lenticulina) calcar, Lenticulina (Robulus) cultrata, Lenticulina (Robulus) similis, Lenticulina (Robulus) ornata Lenticulina (Lenticulina) inornata d'Orbigny 1846 Lenticulina (Lenticulina) calcar Lenticulina (Robulus) cultrata Lenticulina (Robulus) similis Lenticulina (Robulus) ornata
Foraminifera
Lituolacea - Ataxophragmiidae	Martinottiella communis Martinottiella communis Huang et al. 2019
Lituolida - Lituolidae	Bigenerina nodosaria, Textularia gramen Bigenerina nodosaria Textularia gramen dOrbigny 1846
Buliminoidea - Buliminidae	Virgulina schreibersiana, Virgulina (Virgulinella) pertusa Virgulina schreibersiana Virgulina (Virgulinella) pertusa
Spiroplectamminacea - Spiroplectamminidae	Spiroplectammina carinata Spiroplectammina carinata d'Orbigny 1846
Tubothalamea
Miliolida - Miliolidae	Sigmoilina tenuis Sigmoilina tenuis
Foraminifera
Buliminacea - Uvigerinidae	Uvigerina semiornata Uvigerina semiornata U. semiornata for. semiornata
Asterigerinacea - Asterigerinidae	Asterigerina guerichi Asterigerina guerichi A. gürichi staeschi
Discorbacea - Bagginidae	Cancris auriculus Cancris auriculus Fichtel and Moll 1798
Gymnolaemata
Cheilostomata - Lunulitidae	Lunulites voigti¹, Lunulites burdigalensis¹ Lunulites voigti¹ Lunulites burdigalensis¹
Cheilostomata - Cupuladriidae	Reussirella haidingeri¹ Reussirella haidingeri¹
Scaphopoda
Gadilida - Gadilidae	Gadila dingdenensis¹ Gadila dingdenensis¹ tusk shell
Dentaliida - Dentaliidae	Lentigodentalium pseudomutabile¹, "Pseudantalis aturensis" = Dentalium (Pseudantilis)¹ Lentigodentalium pseudomutabile¹ tusk shell "Pseudantalis aturensis" = Dentalium (Pseudantilis)¹ tusk shell
Gastropoda
Opisthobranchiata -	Roxania utriculus³ Roxania utriculus³ snail
Opisthobranchiata - Cylichnidae	Scaphander grateloupi³, Cylichna (Cylichna) pseudoconvoluta³ Scaphander grateloupi³ snail Cylichna (Cylichna) pseudoconvoluta³ snail
Opisthobranchiata - Retusidae	Retusa (Cylichnina) elongata³, Volvulella (Volvulella) acuminata³ Retusa (Cylichnina) elongata³ snail Volvulella (Volvulella) acuminata³ snail
Heterostropha - Pyramidellidae	Pyramidella (Voluspa) plicosa³, Eulimella (Eulimella) acicula³, Turbonilla (Turbonilla) gastaldi³, Turbonilla (Turbonilla) koeneniana³, Syrnola hoernesi³, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³, Odostomia (Brachystomia) sp.³, Odostomia (Megastomia) cf. conoidea³ Pyramidella (Voluspa) plicosa³ snail Eulimella (Eulimella) acicula³ snail Turbonilla (Turbonilla) gastaldi³ snail sensu JANSSEN Turbonilla (Turbonilla) koeneniana³ snail sensu JANSSEN Syrnola hoernesi³ snail "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³ snail sensu JANSSEN Odostomia (Brachystomia) sp.³ Fleming 1817 snail Odostomia (Megastomia) cf. conoidea³ Brocchi 1814 snail
Heterostropha - Ringiculidae	Ringicula (Ringiculina) buccinea³ Ringicula (Ringiculina) buccinea³ Sowerby 1823 snail
Heterostropha - Acteonidae	Crenilabium terebelloides³, Acteon (Acteon) semistriatus³ Crenilabium terebelloides³ snail Acteon (Acteon) semistriatus³ snail spelled as 'Actaeon'
Sorbeoconcha - Naticidae	Naticidae indet.², Euspira edithae², Euspira staringi², Euspira helicina², "Natica sensu lato tigrina" = Cochlis tigrina², Tectonatica miopusilla² Naticidae indet.² Guilding 1834 moon snail Naticidae juv. et indet. Euspira edithae² moon snail Euspira staringi² moon snail Euspira helicina² Brocchi 1814 moon snail E. aggr. helicina "Natica sensu lato tigrina" = Cochlis tigrina² Röding 1798 moon snail Tectonatica miopusilla² moon snail
Sorbeoconcha - Rissoidae	sensu lato Alvania holsatica², Alvania (Alvania) basisulcata² sensu lato Alvania holsatica² snail Alvania (Alvania) basisulcata² snail
Sorbeoconcha - Triviidae	Erato (Erato) germanica² Erato (Erato) germanica² Schilder 1929 snail
Sorbeoconcha - Aporrhaidae	Aporrhais (Aporrhais) alata² Aporrhais (Aporrhais) alata² snail
Sorbeoconcha - Xenophoridae	Xenophora deshayesi² Xenophora deshayesi² Michelotti 1847 carrier shell
Neogastropoda - Cancellariidae	Trigonostoma (Trigonostoma) aperta², Trigonostoma (Trigonostoma) extractrix², Trigonostoma (Trigonostoma) cf. spinifera², Sveltia varicosa², "Trigonostoma (Ventrilia) pouwi" = Ventrilia², Cancellaria (Merica) contorta², Babylonella fusiformis² Trigonostoma (Trigonostoma) aperta² snail Trigonostoma (Trigonostoma) extractrix² snail Trigonostoma (Trigonostoma) cf. spinifera² snail Sveltia varicosa² snail S. varicosa paucicostata "Trigonostoma (Ventrilia) pouwi" = Ventrilia² snail Cancellaria (Merica) contorta² snail C. contorta gelriana Babylonella fusiformis² snail
Neogastropoda - Costellariidae	Orthosurcula steinvorthi³ Orthosurcula steinvorthi³ ribbed mitre
Neogastropoda - Buccinidae	Phos (Phos) decussatus² Phos (Phos) decussatus² true whelk
Neogastropoda - Nassariidae	Nassarius bocholtensis², Nassarius tenuistriatus², Nassarius voorthuyseni², Nassarius karinae² Nassarius bocholtensis² snail Nassarius tenuistriatus² snail Nassarius voorthuyseni² snail Nassarius karinae² Wienrich 2001 snail
Neogastropoda - Columbellidae	Anachis (Costoanachis) hosiusi², Anachis (Costoanachis) terebralis², Mitrella (Macrurella) nassoides² Anachis (Costoanachis) hosiusi² dove shell Anachis (Costoanachis) terebralis² dove shell Mitrella (Macrurella) nassoides² dove shell
Neogastropoda - Fasciolariidae	Streptodictyon sexcostatus² Streptodictyon sexcostatus² spindle shell
Neogastropoda - Olividae	Oliva (Strephona) dufresnei² Oliva (Strephona) dufresnei² Basterot 1825 olive snail
Neogastropoda - Ancillariidae	"Ancilla (Baryspira) obsoleta" = Amalda obsoleta² "Ancilla (Baryspira) obsoleta" = Amalda obsoleta² Brocchi 1814 snail
Neogastropoda - Muricidae	"Murex (Haustellum) inornatus" = Haustellum², "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta², Typhis (Typhis) ? pungens², Lyrotyphis (Eotyphis) fistulosus² "Murex (Haustellum) inornatus" = Haustellum² murex snail "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta² Bronn 1831 murex snail Typhis (Typhis) ? pungens² Solander in Brander and Brander 1766 murex snail Lyrotyphis (Eotyphis) fistulosus² murex snail
Neogastropoda - Mangeliidae	sensu lato Mangelia andersoni n. sp.³, Sorgenfreispira "sp. 2"³, Sorgenfreispira tenella³, Benthomangelia aff. obtusangula³ sensu lato Mangelia andersoni n. sp.³ Wienrich and Janssen 2007 snail Sorgenfreispira "sp. 2"³ Moroni 1979 snail Sorgenfreispira tenella³ Conrad 1848 snail Benthomangelia aff. obtusangula³ snail
Neogastropoda - Conidae	"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ "Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ cone shell
Neogastropoda - Terebridae	Strioterebrum hoernesi³, Strioterebrum ronaldi³ Strioterebrum hoernesi³ auger snail Strioterebrum ronaldi³ auger snail
Neogastropoda - Borsoniidae	Asthenotoma pannoides³, Asthenotoma pannus³, Bathytoma jugleri³ Asthenotoma pannoides³ snail Asthenotoma pannus³ snail Bathytoma jugleri³ snail
Neogastropoda - Pseudomelatomidae	sensu lato Inquisitor borealis³ sensu lato Inquisitor borealis³ snail
Neogastropoda - Clavatulidae	Fusiturris aquensis³, Fusiturris duchastelii³, Clavatula boreointerrupta³ Fusiturris aquensis³ snail Fusiturris duchastelii³ snail Clavatula boreointerrupta³ snail
Neogastropoda - Horaiclavidae	? Ceritoturris hannoverana³ ? Ceritoturris hannoverana³ snail
Neogastropoda - Drilliidae	Splendrillia selenkae³ Splendrillia selenkae³ snail
Neogastropoda - Clathurellidae	Pleurotomoides luisae³, Pleurotomoides simplex³ Pleurotomoides luisae³ snail Pleurotomoides simplex³ snail
Neogastropoda - Turridae	Gemmula (Gemmula) denticula³, Gemmula (Gemmula) zimmermanni³, sensu lato Unedogemmula boreoturricula³, sensu lato Unedogemmula hanseata³, sensu lato Unedogemmula stoffelsii³, Boreodrillia hosiusi³, Nitidiclavus maitrejus³ Gemmula (Gemmula) denticula³ turrid G. denticula borealis Gemmula (Gemmula) zimmermanni³ turrid sensu lato Unedogemmula boreoturricula³ turrid sensu lato Unedogemmula hanseata³ turrid sensu lato Unedogemmula stoffelsii³ turrid Boreodrillia hosiusi³ turrid Nitidiclavus maitrejus³ turrid
Neogastropoda - Raphitomidae	Metuonella grippi³, Mioawateria andersoni³, Teretia anceps³, sensu lato Teretia "sp. 2"³ Metuonella grippi³ snail Mioawateria andersoni³ snail Teretia anceps³ snail sensu lato Teretia "sp. 2"³ Norman 1888 snail
Sorbeoconcha - Cassidae	"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata², "Semicassis (Echinophoria) bicoronata" = Echinophoria² "Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata² Sacco 1890 snail "Semicassis (Echinophoria) bicoronata" = Echinophoria² snail
Sorbeoconcha - Vitrinellidae	Circulus praecedens², Circulus hennei² Circulus praecedens² snail C. praecedens for. praecedens Circulus hennei² Glibert 1952 snail
Sorbeoconcha - Triphoridae	Norephora (Norephora) fritschi² Norephora (Norephora) fritschi² snail
Sorbeoconcha - Triforidae	Cerithiella (Cerithiella) genei² Cerithiella (Cerithiella) genei² snail
Cerithioidea - Cerithiidae	"Bittium (Bittium) spina" = Bittium spina² "Bittium (Bittium) spina" = Bittium spina² Partsch 1856 cerith snail
Cerithioidea - Turritellidae	Turritella subangulata² Turritella subangulata² turret shell
Vanikoroidea - Eulimidae	Eulima (Eulima) bilineata², Eulima (Eulima) glabra², Balcis (Balcis) alba² Eulima (Eulima) bilineata² snail Eulima (Eulima) glabra² snail Balcis (Balcis) alba² snail
Epitonioidea - Epitoniidae	Opalia (Pliciscala) pertusa², Cirsotrema (Cirsotrema) crassicostatum², "Amaea (Scalina) subreticula" = Scalina² Opalia (Pliciscala) pertusa² wentletrap Cirsotrema (Cirsotrema) crassicostatum² Deshayes 1839 wentletrap "Amaea (Scalina) subreticula" = Scalina² wentletrap
Neotaenioglossa - Calyptraeidae	Calyptraea (Calyptraea) chinensis² Calyptraea (Calyptraea) chinensis² Linnaeus 1758 Chinese hat snail
Bivalvia
Nuculida - Nuculidae	Nuculoma haesendoncki¹ Nuculoma haesendoncki¹ nut clam N. haesendoncki hanseata
Nuculanida - Nuculanidae	"Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ "Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ Nyst and Westendorp 1839 pointed nut clam
Nuculanida - Yoldiidae	"Portlandia (Yoldiella) pygmaea" = Yoldiella¹, Yoldia (Yoldia) glaberrima¹ "Portlandia (Yoldiella) pygmaea" = Yoldiella¹ clam Yoldia (Yoldia) glaberrima¹ von Münster 1837 clam
Ostreida - Pinnidae	Atrina pectinata¹ Atrina pectinata¹ Linnaeus 1758 oyster
Pectinida - Pectinoidae	Korobkovia woodi¹, Aequipecten seniensis¹, Pseudamussium lilli¹ Korobkovia woodi¹ scallop Aequipecten seniensis¹ scallop Pseudamussium lilli¹ Pusch 1837 scallop
Arcida - Limopsidae	Limopsis (Limopsis) aurita¹, "Limopsis (Pectunculina) anomala" = Limopsis¹, Limopsis (Pectunculina) lamellata¹ Limopsis (Limopsis) aurita¹ Brocchi 1814 clam "Limopsis (Pectunculina) anomala" = Limopsis¹ clam Limopsis (Pectunculina) lamellata¹ Lehmann 1885 clam
Arcida - Arcidae	Scapharca (Scapharca) diluvii¹ Scapharca (Scapharca) diluvii¹ Lamarck 1805 clam
Carditida - Carditidae	Cyclocardia sp.¹ Cyclocardia sp.¹ Conrad 1867 clam
Carditida - Astartidae	Astarte (Astarte) radiata¹, Goodallia (Goodallia) angulata¹ Astarte (Astarte) radiata¹ Nyst and Westendorp 1839 clam Astarte radiata fa. radiata 28 specimens, Astarte radiata fa. convexior 33 specimens, Astarte radiata fa. goldfussi 2 specimens Goodallia (Goodallia) angulata¹ Lehmann 1885 clam
Pholadida - Corbulidae	Corbula (Varicorbula) gibba¹ Corbula (Varicorbula) gibba¹ Oliva 1792 clam C. (V.) gibba gibba
Cardiida - Veneridae	"Gouldia minima" = Gouldia (Gouldia) minima¹, Venus (Ventricoloidea) multilamella¹ "Gouldia minima" = Gouldia (Gouldia) minima¹ Montagu 1803 venus clam Venus (Ventricoloidea) multilamella¹ Lamarck 1818 venus clam V. (V.) multilamella multilamella
Cardiida - Glossidae	Glossus (Glossus) lunulatus¹ Glossus (Glossus) lunulatus¹ clam
Cardiida - Mactridae	Spisula elliptica¹ Spisula elliptica¹ Brown 1827 clam
Cardiida - Tellinidae	"Macoma elliptica" = Laciolina magna¹ "Macoma elliptica" = Laciolina magna¹ Spengler 1798 tellin clam
Cardiida - Semelidae	sensu lato Abra lehmanni¹ sensu lato Abra lehmanni¹ abra
Cardiida - Cardiidae	Parvicardium straeleni¹, Parvicardium scabrum¹, Habecardium dingdense¹ Parvicardium straeleni¹ cockle Parvicardium scabrum¹ Philippi 1844 cockle Habecardium dingdense¹ Lehmann 1885 cockle
Pandorida - Lyonsiidae	Lyonsia (Lyonsia) aff. norwegica¹ Lyonsia (Lyonsia) aff. norwegica¹ clam
Lucinida - Lucinidae	"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti¹ "Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti¹ Nyst 1861 clam C. droueti droueti
Sedentaria
Sabellida - Serpulidae	Serpulidae indet.¹ Serpulidae indet.¹ Rafinesque 1835 Serpulidae indet. D
Anthozoa
Scleractinia - Flabellidae	Flabellum sp., Flabellum tuberculatum, Flabellum cristatum, Flabellum pompeckji Flabellum sp. Lesson 1831 stony coral Flabellum tuberculatum Keferstein 1859 stony coral Flabellum cristatum Milne-Edwards and Haime 1851 stony coral Flabellum pompeckji Krejci 1926 stony coral
Scleractinia - Caryophylliidae	Deltocyathus reussi Deltocyathus reussi stony coral
Scleractinia - Micrabaciidae	Stephanophyllia imperialis Stephanophyllia imperialis Michelin 1841 stony coral

¹Wienrich 1999; ²Wienrich 2001; ³Wienrich and Janssen 2007