Verdigris Coulee (UA-MR-4): Late Santonian, Canada
collected by R. Fox

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Osteichthyes - Temnospondyli - Albanerpetontidae
Gardner 2000 11 specimens
Gardner 2000 4 specimens
Gardner 2000 2 specimens
Osteichthyes - Anguidae
Anguidae indet. Gray 1825
Gao and Fox 1996 3 specimens
UALVP 33378 (left dentary) plus many unnumbered fragments
Osteichthyes
Gao and Fox 1996 1 specimen
UALVP 29830 (partial right maxilla)
Osteichthyes - Tyrannosauridae
Larson 2008 5 specimens
Osteichthyes
Larson 2008 18 specimens
morphs II and V
Osteichthyes - Dromaeosauridae
Larson 2008 1 specimen
UALVP 50213
Larson 2008 85 specimens
Williamson and Brusatte 2014 39 specimens
"Milk River Saurornitholestinae gen. et sp."
Larson 2008 5 specimens
incl. "Milk River Dromaeosaurinae gen. et sp."
Larson 2008 2 specimens
Larson 2008 51 specimens
Williamson and Brusatte 2014 29 specimens
"Aquilan cf. Richardoestesia gilmorei"
Larson 2008 44 specimens
Williamson and Brusatte 2014 7 specimens
"?Aquilan cf. Richardoestesia isosceles"
Osteichthyes - Eutriconodonta - Triconodontidae
Osteichthyes - Multituberculata - Cimolomyidae
Fox 1971
Fox 1971
Osteichthyes - Multituberculata
Fox 1971
Fox 1971
Fox 1971
Osteichthyes - Multituberculata - Neoplagiaulacidae
Fox 1971
Osteichthyes - Multituberculata - Cimolodontidae
Fox 1971
Fox 1971
Osteichthyes - Alphadontidae
Fox 1971
Fox 1971
Osteichthyes - Aquiladelphidae
Fox 1971
Fox 1971
Osteichthyes
Fox 1971
Chondrichthyes - Hybodontiformes - Hybodontidae
Brinkman et al. 2017 2 specimens
see common names

Geography
Country:Canada State/province:Alberta
Coordinates: 49.1° North, 111.8° West (view map)
Paleocoordinates:57.8° North, 67.1° West (Wright 2013)
Basis of coordinate:based on nearby landmark
Geographic resolution:small collection
Time
Period: Cretaceous Epoch: Late Cretaceous
Stage: Santonian 10 m.y. bin: Cretaceous 6
*Epoch:Senonian
*International age/stage:Campanian
Key time interval: Late Santonian
Age range of interval: 85.7 - 83.6 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Milk River Member:Deadhorse Coulee
Stratigraphic resolution:bed
Stratigraphy comments: upper part of formation
Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:mesofossils
Collection methods and comments
Collection excludes:some genera
Collection methods:bulk,sieve,field collection
Reason for describing collection:taxonomic analysis
Collectors:R. Fox
Metadata
Also known as:UALVP MR-4, MR-04
Database number:14423
Authorizer:J. Alroy, R. Benson, M. Carrano, P. Holroyd Enterer:J. Alroy, M. Carrano, P. Holroyd, R. Benson
Modifier:M. Carrano Research group:vertebrate
Created:1994-04-30 00:00:00 Last modified:2026-02-12 10:23:43
Access level:the public Released:1994-04-30 00:00:00
Creative Commons license:CC0
Reference information

Primary reference:
1508. R. C. Fox. 1969. Studies of Late Cretaceous vertebrates. III. A triconodont mammal from Alberta. Canadian Journal of Zoology 47:1253-1256 [J. Alroy/J. Alroy/R. Benson]

Secondary references:

6294 J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
64013 D. B. Brinkman, A. G. Neuman, and J. D. Divay. 2017. Non-marine fishes of the late Santonian Milk River Formation of Alberta, Canada – evidence from vertebrate microfossil localities. Vertebrate Anatomy Morphology Paleontology 3:7-46 [P. Holroyd/P. Holroyd/M. Carrano]
1510 R. C. Fox. 1971. Early Campanian multituberculates (Mammalia: Allotheria) from the upper Milk River Formation, Alberta. Canadian Journal of Earth Sciences 8:916-938 [J. Alroy/J. Alroy/M. Carrano]
1511 R. C. Fox. 1971. Marsupial mammals from the early Campanian Milk River Formation, Alberta, Canada. Zoological Journal of the Linnean Society 50(suppl. 1):145-164 [J. Alroy/J. Alroy/M. Carrano]
37933 K. -Q. Gao and R. C. Fox. 1996. Taxonomy and evolution of Late Cretaceous lizards (Reptilia: Squamata) from western Canada. Bulletin of the Carnegie Museum of natural History 33:1-107 [R. Benson/R. Benson]
42675 J. D. Gardner. 2000. Albanerpetontid amphibians from the Upper Cretaceous (Campanian and Maastrichtian) of North America. Geodiversitas 22(3):349-388 [M. Carrano/M. Carrano/M. Carrano]
94184 D. W. Larson. 2008. Diversity and variation of theropod dinosaur teeth from the uppermost Santonian Milk River Formation (Upper Cretaceous), Alberta: a quantitative method supporting identification of the oldest dinosaur tooth assemblage in Canada. Canadian Journal of Earth Sciences 45(12):1455-1468 [M. Carrano/M. Carrano]
50790 T. E. Williamson and S. L. Brusatte. 2014. Small theropod teeth from the Late Cretaceous of the San Juan Basin, northwestern New Mexico and their implications for understanding latest Cretaceous dinosaur evolution. PLoS ONE 9(4):e93190:1-23 [M. Carrano/M. Carrano]