Midco Locality 15 (Permian of the United States)

Where: Oklahoma (36.3° N, 97.3° W)

• Paleocoordinates: 0.5° N, 27.7° W (Wright 2013)

• coordinate stated in text

• small collection-level geographic resolution

When: Midco Member (Wellington Formation), Artinskian (290.1 - 279.3 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine; lithified, dolomitic, argillaceous lime mudstone

• Salty playa lake. The lower layer shows sedimentational and organic features which suggest that it was laid down close to the shore and that most of its biota was washed into the lake. The matrix and biota of the middle layer, which is highly irregular and characterized by pink partings, were probably transported to the water by violent dust storms. The upper layer is highly argillaceous, and was deposited under strongly saline conditions; insects are rare but they are very well preserved.
• Several layers of fine-grained, compact, argillaceous dolomitic lime stone, separated by fissile shale. Three of the limestone layers contain insects, and for convenience of reference they have been designated the lower, middle and upper insect layers.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Primary reference: F. M. Carpenter. 1947. Lower Permian Insects from Oklahoma. Part 1. Introduction and the Orders Megasecoptera, Protodonata, and Odonata. Proceedings of the American Academy of Arts and Sciences 76:25-54 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 104824: authorized by Matthew Clapham, entered by Matthew Clapham on 10.02.2011, edited by Jered Karr

Creative Commons license: CC0 (CC0)

Taxonomic list

Show authors, comments, and common names
Insecta
 Megasecoptera - Bardohymenidae
Calohymen permianus n. gen. n. sp.
Calohymen permianus n. gen. n. sp. Carpenter 1947 winged insect
 Megasecoptera - Protohymenidae
Protohymen curvatus, Protohymen largus n. sp., Protohymen venustus n. sp.
Protohymen curvatus Carpenter 1947 winged insect
Protohymen largus n. sp. Carpenter 1947 winged insect
Protohymen venustus n. sp. Carpenter 1947 winged insect
 Diaphanopterodea - Parelmoidae
Pseudelmoa ampla n. gen. n. sp., Parelmoa radialis n. sp., Parelmoa revelata n. gen. n. sp., Parelmoa obtusa n. sp.
Pseudelmoa ampla n. gen. n. sp. Carpenter 1947 winged insect
Parelmoa radialis n. sp. Carpenter 1947 winged insect
Parelmoa revelata n. gen. n. sp. Carpenter 1947 winged insect
Parelmoa obtusa n. sp. Carpenter 1947 winged insect
 Diaphanopterodea - Martynoviidae
Eumartynovia raaschi n. gen. n. sp., Phaneroneura martynovae n. gen. n. sp., Phaneroneura reducta n. sp.
Eumartynovia raaschi n. gen. n. sp. Carpenter 1947 winged insect
Phaneroneura martynovae n. gen. n. sp. Carpenter 1947 winged insect
Phaneroneura reducta n. sp. Carpenter 1947 winged insect
 Diaphanopterodea - Asthenohymenidae
Asthenohymen triangularis n. sp., Asthenohymen latus n. sp., Asthenohymen apicalis n. sp., Asthenohymen parvulus n. sp.
Asthenohymen triangularis n. sp. Carpenter 1947 winged insect
Asthenohymen latus n. sp. Carpenter 1947 winged insect
Asthenohymen apicalis n. sp. Carpenter 1947 winged insect
Asthenohymen parvulus n. sp. Carpenter 1947 winged insect
 Paoliida - Anthracoptilidae
Homocladus grandis1
Homocladus grandis1 Carpenter 1966 winged insect
MCZ 31155
 Dicondylia - Ditaxineuridae
Ditaxineura cellulosa
Ditaxineura cellulosa Carpenter 1933 winged insect
 Dicondylia - Progoneuridae
Progoneura nobilis
Progoneura nobilis Carpenter 1947 winged insect
 Meganisoptera - Meganeuridae
Meganeuropsis americana n. sp., Oligotypus tillyardi
Meganeuropsis americana n. sp. Carpenter 1947 griffinfly
Oligotypus tillyardi Carpenter 1931 griffinfly
 Ephemerida - Misthodotidae
Misthodotes edmundsi n. sp.2, Misthodotes ovalis2
Misthodotes edmundsi n. sp.2 Carpenter 1979 mayfly
MCZ 5184 ab, 5194, 313 ab
Misthodotes ovalis2 Tillyard 1932 mayfly
MCZ 5193
 Ephemerida - Protereismatidae
Kukalova americana n. gen. n. sp.3, "Protereisma directum n. sp." = Alexandrinia directa2
Kukalova americana n. gen. n. sp.3 Demoulin 1970 mayfly
"Protereisma directum n. sp." = Alexandrinia directa2 Carpenter 1979 mayfly
MCZ 5180 ab
Xiphosura
 Xiphosurida - Belinuridae
"Anacontium carpenteri n. gen. n. sp." = Anacontium carpenteri4, "Anacontium brevis n. sp." = Anacontium carpenteri4
"Anacontium carpenteri n. gen. n. sp." = Anacontium carpenteri4 Raymond 1944
"Anacontium brevis n. sp." = Anacontium carpenteri4 Raymond 1944
1Béthoux et al. 2010; 2Carpenter 1979; 3Demoulin 1970; 4Raymond 1944