Madygen, PIN collection 2785, Dzhailoucho area (Triassic of Kyrgyzstan)

Where: Osh, Kyrgyzstan (40.1° N, 70.2° E: paleocoordinates 36.5° N, 60.1° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Madygen Formation, Ladinian (242.0 - 237.0 Ma)

• Aristov (2020): The age of the Madygen locality is usually reported as the Ladinian or Carnian of the Middle or Upper Triassic (Shcherbakov, 2008). This concept emerged, because I.A. Dobruskina (1995) did not discriminate between Ladinian and Carnian floras. Our data show that only three of the six families found at Madygen are known from younger sediments (Fig. 1), and therefore we can confirm the assumption of the Ladinian age of the Madygen locality (A.G. Ponomarenko, personal communication, 2018).

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - small; lithified siliciclastic sediments

• Oxbow lake; Madygen basin; tectonically-induced intermontane basin. Complex facies pattern consistent with active tectonics in upland region.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Primary reference: D. S. Aristov, T. Wappler, and A. P. Rasnitsyn. 2009. New and Little-Known Grylloblattids of the Family Geinitziidae (Insecta: Grylloblattida) from the Triassic and Jurassic of Europe, Asia, and South Africa. Paleontological Journal 43:418-424 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 105007: authorized by Matthew Clapham, entered by Matthew Clapham on 13.02.2011, edited by Jered Karr, Lindy Edwards and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

• Shurabia serrata stated to come from Shurab, but collection number corresponds to Madygen
Insecta
 Eoblattida - Batkentakidae
Batkentak intactus n. gen. n. sp.1
Batkentak intactus n. gen. n. sp.1 Aristov 2015 insect
PIN 2785/2179
 Eoblattida - Mesorthopteridae
Parastenaropodites longiuscula n. sp.32 Storozhenko 1996 insect
Parastenaropodites fluxa n. gen.32 Storozhenko 1996 insect
Sharovites alexanderi4 Aristov and Storozhenko 2013 insect
PIN 2785/2199
Mesoidelia faceta32 Storozhenko 1996 insect
PIN 2785/2107,2167,2234
"Parastenaropodites nervosa" = Austroidelia nervosa32 Storozhenko 1996 insect
Austroidelia asiatica n. sp.32 Storozhenko 1996 insect
 Eoblattida - Blattogryllidae
Anoblattogryllus fundatus n. gen. n. sp.27, Baharellus madygensis29, Costatoviblatta aenigmatosa29, "Mesoblattogryllus conjunctus n. sp." = Costatoviblatta conjuncta29
Anoblattogryllus fundatus n. gen. n. sp.27 Storozhenko 1990 insect
PIN 2785/2166
Baharellus madygensis29 Storozhenko 1992 insect
PIN 2785/2245, 2266
Costatoviblatta aenigmatosa29 Storozhenko 1992 insect
PIN 2785/2126, 2127, 2160, 2201, 2240, 2243
"Mesoblattogryllus conjunctus n. sp." = Costatoviblatta conjuncta29 Storozhenko 1992 insect
PIN 2785/2242, 2264
 Eoblattida - Megakhosaridae
"Microblattogryllus variabilis" = Protoblattogryllus variabilis27, Megablattogryllus pinguis n. sp.27, Megablattogryllus austerus n. sp.27, Metakhosara sharovi n. gen. n. sp.30
"Microblattogryllus variabilis" = Protoblattogryllus variabilis27 Storozhenko 1990 insect
PIN 2785/2136, 2171
Megablattogryllus pinguis n. sp.27 Storozhenko 1990 insect
PIN 2785/2121
Megablattogryllus austerus n. sp.27 Storozhenko 1990 insect
PIN 2785/2178
Metakhosara sharovi n. gen. n. sp.30 Storozhenko 1993 insect
PIN 2785/2360
 Dicondylia - Kennedyidae
Kennedya gracilis n. sp.18 Pritykina 1981 winged insect
PIN 2785/23
Kennedya carpenteri n. sp.18 Pritykina 1981 winged insect
PIN 2785/2, 6
 Dicondylia - Batkeniidae
Batkenia pusilla n. gen. n. sp.18
Batkenia pusilla n. gen. n. sp.18 Pritykina 1981 winged insect
PIN 2785/22
 Dicondylia - Zygophlebiidae
Zygophlebia ramosa n. gen. n. sp.12, Zygophlebia ramosa n. gen. n. sp.18, Cyrtophlebia sinuosa n. gen. n. sp.12, Cyrtophlebia sinuosa n. sp.18, Mixophlebia mixta18
Zygophlebia ramosa n. gen. n. sp.12 Pritykina 1981 winged insect
PIN 2785/20
Zygophlebia ramosa n. gen. n. sp.18 Pritykina 1981 winged insect
PIN 2785/20, 4101
Cyrtophlebia sinuosa n. gen. n. sp.12 Pritykina 1981 winged insect
PIN 2785/25
Cyrtophlebia sinuosa n. sp.18 Pritykina 1981 winged insect
PIN 2785/25
Mixophlebia mixta18 Pritykina 1981 winged insect
PIN 2785/8
 Dicondylia - Paurophlebiidae
Paurophlebia lepida n. sp.18 Pritykina 1981 winged insect
PIN 2785/7
Paurophlebia angusta n. sp.18 Pritykina 1981 winged insect
PIN 2785/4097
Neritophlebia longa n. sp.18 Pritykina 1981 winged insect
PIN 2785/24
Neritophlebia vicina n. sp.18 Pritykina 1981 winged insect
PIN 2785/1
Neritophlebia elegans n. gen. n. sp.18 Pritykina 1981 winged insect
PIN 2785/4, 3
 Dicondylia - Triadotypidae
"Triadotypus sogdianus n. sp." = Reisia sogdiana18
"Triadotypus sogdianus n. sp." = Reisia sogdiana18 Pritykina 1981 winged insect
PIN 2785/21
 Mecoptera - Thaumatomeropidae
Thaumatomerope sogdiana n. gen. n. sp.17 Rasnitsyn 1974 scorpionfly
PIN 2785/2496, 3753, 4072
Thaumatomerope oligoneura n. sp.17 Rasnitsyn 1974 scorpionfly
PIN 2785/2498, 3974
 Mecoptera - Parachoristidae
Triassochorista kirgizica13 Novokshonov 1997 scorpionfly
PIN 2785/3793
Parachorista arguta n. sp.15 Novokshonov 2001 scorpionfly
PIN 2785/3769, 3887
Parachorista immota n. sp.15 Novokshonov 2001 scorpionfly
PIN 2785/4135, 3228
 Mecoptera - Permochoristidae
Agetopanorpa deceptoria n. sp.14 Novokshonov 1997 scorpionfly
PIN 2785/4001
Mesageta rieki14 Novokshonov 1997 scorpionfly
PIN 2785/3841, 3896(3919)
Liassochorista ? utilis14 Novokshonov 1997 scorpionfly
PIN 2785/3970
 Mecoptera - Orthophlebiidae
Choristopanorpa temperata n. sp.15 Novokshonov 2001 scorpionfly
PIN 2785/4024
 Mecoptera - Mesopsychidae
Prochoristella longa14 Novokshonov 1997 scorpionfly
PIN 2785/3969
Mesopsyche justa16 Novokshonov and Sukacheva 2001 scorpionfly
PIN 2785/4021+3970
Mesopsyche ordinata n. sp.16 Novokshonov and Sukacheva 2001 scorpionfly
PIN 2785/4053
Mesopsyche tortiva n. sp.16 Novokshonov and Sukacheva 2001 scorpionfly
PIN 2785/4052, 3198
 Coleoptera - Obrieniidae
Obrieniidae indet.36 Zherikhin and Gratshev 1993 weevil
PIN 2785/2360
Obrienia illaetabilis n. sp.36 Zherikhin and Gratshev 1993 weevil
PIN 2785/3725
 Coleoptera - Peltosynidae
Peltosyne varyvrosa n. sp.35, Ofthalmopeltos synkritos n. gen. n. sp.35
Peltosyne varyvrosa n. sp.35 Yan et al. 2017 beetle
PIN 2785/2655
Ofthalmopeltos synkritos n. gen. n. sp.35 Yan et al. 2017 beetle
PIN 2785/2888
 Neuroptera - Berothidae
Ferganoberotha minutissima n. gen. n. sp.11
Ferganoberotha minutissima n. gen. n. sp.11 Khramov 2023 beaded lacewing
PIN 2785/4061
 Hemiptera - Serpentivenidae
Serpentivena tigrina n. gen. n. sp.19
Serpentivena tigrina n. gen. n. sp.19 Shcherbakov 1984 true bug
PIN 2785/3535(3635), 3102(3103)
 Hemiptera - Ipsviciidae
Strivicia davidi24 Shcherbakov 2022 true bug
PIN 2785/2432
 Hemiptera - Naibiidae
Coccavus supercubitus20 Shcherbakov 2007 true bug
PIN 2785/3544
 Hemiptera - Mesojabloniidae
Scytachile emeljanovi n. gen. n. sp.21, Fulgobole evansi n. gen.21
Scytachile emeljanovi n. gen. n. sp.21 Shcherbakov 2011 true bug
PIN 2785/3070
Fulgobole evansi n. gen.21 Shcherbakov 2011 true bug
PIN 2785/3238
 Hemiptera - Curvicubitidae
Beaconiella pulchra n. sp.23 Shcherbakov 2021 true bug
PIN 2785/3022(3073), 3387(3893), 3405
Beaconiella cincta n. sp.23 Shcherbakov 2021 true bug
PIN 2785/3107(3381)
 Hemiptera - Maguviopseidae
Fasolinka beckermigdisovae n. gen. n. sp.21, Asiocula lima n. gen.21
Fasolinka beckermigdisovae n. gen. n. sp.21 Shcherbakov 2011 true bug
PIN 2785/3087&3088
Asiocula lima n. gen.21 Shcherbakov 2011 true bug
PIN 2785/3326
 Dicondylia - Permosialidae
Permosialis triassica3 Novokshonov and Zhuzhgova 2004 winged insect
PIN 2785/3343
 Blattaria - Caloblattinidae
Thuringoblatta sogdianensis34 Vishnyakova 1998 cockroach
PIN 2785/1739-1742, 1746, 1748, 1750, 1753, 1761, 1765, 4103
Sogdoblatta nana34 Vishnyakova 1998 cockroach
PIN 2785/2471, 3724
Sogdoblatta porrecta n. sp.34 Vishnyakova 1998 cockroach
PIN 2785/1760(1764), 1744, 1745
 Plecoptera - Perlariopseidae
Fritaniopsis brevicaulis26 Sinitshenkova 1987 stonefly
PIN 2785/2740
 Embioptera - Alexarasniidae
Madygembia rasnitsyni n. gen. n. sp.5 Aristov and Storozhenko 2021 webspinners
PIN 2785/2162
Nestorembia novojilovi22 Shcherbakov 2015 webspinners
PIN 2785/2252
 Dicondylia - Prochresmodidae
Prochresmoda parva n. sp.8 Gorochov 1994 winged insect
PIN 2785/2097, 2098, 2074
Triassophasma pusillum n. sp.8 Gorochov 1994 winged insect
PIN 2785/2084
Triassophasma intermedium8 Gorochov 1994 winged insect
PIN 2785/2275
 Dicondylia - Aeroplanidae
"Paraplana parallelica n. sp." = Sharovoplana parallelica8
"Paraplana parallelica n. sp." = Sharovoplana parallelica8 Gorochov 1994 winged insect
PIN 2785/2090
 Dicondylia - Xiphopteridae
Xiphopterum sharovi n. sp.8 Gorochov 1994 winged insect
PIN 2785/2089, 2085, 2012, 2089a
 Orthoptera - Locustavidae
Locustavus problematicus10 Gorochov 2005 grasshopper
PIN 2785/2339
Locustavus minutus n. sp.10 Gorochov 2005 grasshopper
PIN 2785/2349(2353)
Locustavus intermedius n. sp.10 Gorochov 2005 grasshopper
PIN 2785/2347
Brevilocustavus distinctus n. sp.10 Gorochov 2005 grasshopper
PIN 2785/1986
Miolocustavus reductus n. gen. n. sp.10 Gorochov 2005 grasshopper
PIN 2785/1949
 Orthoptera - Paratitanidae
Paratitan reductus n. sp.9 Gorochov 2003 grasshopper
PIN 2785/2021
 Orthoptera - Mesotitanidae
Prototitan sharovi n. sp.9 Gorochov 2003 grasshopper
PIN 2785/2042
 Orthoptera - Mesoedischiidae
Mesoedischia obliqua7 Gorochov 1987 grasshopper
PIN 2785/1940
 Orthoptera - Gryllavidae
Paragryllavus curvatus n. gen. n. sp.6, Zagryllavus elongatus n. gen. n. sp.6
Paragryllavus curvatus n. gen. n. sp.6 Gorochov 1986 grasshopper
PIN 2785/1945(1952), 1955
Zagryllavus elongatus n. gen. n. sp.6 Gorochov 1986 grasshopper
PIN 2785/1941
 Orthoptera - Hagloedischiidae
Hagloedischia primitiva n. gen. n. sp.6
Hagloedischia primitiva n. gen. n. sp.6 Gorochov 1986 grasshopper
PIN 2785/1951
 Orthoptera - Haglidae
Dulcihagla mistshenkoi n. sp.6, Zamaraga reticulata n. gen. n. sp.6, "Maraga angusta n. gen. n. sp." = Triassaga angusta6, Sonohagla curta n. sp.6, Lyrohagla uvarovi n. gen. n. sp.6, Voliopellus latus n. gen. n. sp.6, Voliopus ancestralis n. gen. n. sp.6, Paravoliopus dorsalis n. gen. n. sp.6, Macrovoliopus declivis n. gen. n. sp.6, Euvoliopus giganteus n. gen. n. sp.6, Haglomorpha martynovi n. gen. n. sp.6, Dolichohagla longa n. gen. n. sp.6, Tinnihagla zeuneri n. gen. n. sp.6, Modihagla ovalis n. gen. n. sp.6
Dulcihagla mistshenkoi n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1961
Zamaraga reticulata n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1992
"Maraga angusta n. gen. n. sp." = Triassaga angusta6 Gorochov 1986 hump-winged cricket
PIN 2785/2359
Sonohagla curta n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2335
Lyrohagla uvarovi n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1877
Voliopellus latus n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1861
Voliopus ancestralis n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1947
Paravoliopus dorsalis n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1892(3)
Macrovoliopus declivis n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1856
Euvoliopus giganteus n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2380
Haglomorpha martynovi n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2378(1991)
Dolichohagla longa n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2368
Tinnihagla zeuneri n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2343
Modihagla ovalis n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2340
 Cnemidolestodea - Sylvabestiidae
Sharovala triassica n. gen. n. sp.2
Sharovala triassica n. gen. n. sp.2 Aristov 2020 winged insect
PIN 2785/3021
 Paoliida - Paoliidae
Ideliopsina stupenda n. sp.33 Storozhenko 1996 winged insect
PIN 2785/2122
Ideliopsina ornata33 Storozhenko 1996 winged insect
PIN 2785/2206
Ideliopsina nana33 Storozhenko 1996 winged insect
PIN 2785/2204, 2120, 2180, 2194, 2233
 Grylloblattodea - Madygenophlebiidae
Madygenophlebia bella28 Storozhenko 1992 ice crawler
PIN 2785/2155,2256
Madygenophlebia nana28 Storozhenko 1992 ice crawler
PIN 2785/2169
 Grylloblattodea - Gorochoviidae
Gorochoviella conjuncta n. gen.31 Storozhenko 1994 ice crawler
PIN 2785/2183,2237
Gorochovia minuta31 Storozhenko 1994 ice crawler
Gorochovia bifurca n. sp.31 Storozhenko 1994 ice crawler
PIN 2785/2262,2183,2184,2237
Gorochovia individua n. gen.31 Storozhenko 1994 ice crawler
Pseudoliomopterites obscurus n. gen.31 Storozhenko 1994 ice crawler
PIN 2785/2111
 Grylloblattodea -
Pseudoshurabia pallidula33 Storozhenko 1996 ice crawler
PIN 2785/2196
 Reculida - Geinitziidae
Shurabia serrata n. sp. Aristov et al. 2009 winged insect
PIN 2785/2161
 Diptera - Psychotipidae
Psychotipa predicta n. gen. n. sp.25
Psychotipa predicta n. gen. n. sp.25 Shcherbakov 1995 fly
PIN 2785/2490
 Diptera - Gnomuscidae
Gnomusca molecula n. gen. n. sp.25
Gnomusca molecula n. gen. n. sp.25 Shcherbakov 1995 fly
PIN 2785/4096
 Diptera - Protorhyphidae
Vymrhyphus tuomikoskii n. gen. n. sp.25
Vymrhyphus tuomikoskii n. gen. n. sp.25 Blagoderov 1995 fly
PIN 2785/3052