Madygen, PIN collection 2785, Dzhailoucho area (Triassic of Kyrgyzstan)

Where: Osh, Kyrgyzstan (40.1° N, 70.2° E: paleocoordinates 36.5° N, 60.1° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Madygen Formation, Ladinian (242.0 - 237.0 Ma)

• Aristov (2020): The age of the Madygen locality is usually reported as the Ladinian or Carnian of the Middle or Upper Triassic (Shcherbakov, 2008). This concept emerged, because I.A. Dobruskina (1995) did not discriminate between Ladinian and Carnian floras. Our data show that only three of the six families found at Madygen are known from younger sediments (Fig. 1), and therefore we can confirm the assumption of the Ladinian age of the Madygen locality (A.G. Ponomarenko, personal communication, 2018).

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - small; lithified siliciclastic sediments

• Oxbow lake; Madygen basin; tectonically-induced intermontane basin. Complex facies pattern consistent with active tectonics in upland region.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Primary reference: D. S. Aristov, T. Wappler, and A. P. Rasnitsyn. 2009. New and Little-Known Grylloblattids of the Family Geinitziidae (Insecta: Grylloblattida) from the Triassic and Jurassic of Europe, Asia, and South Africa. Paleontological Journal 43:418-424 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 105007: authorized by Matthew Clapham, entered by Matthew Clapham on 13.02.2011, edited by Jered Karr, Lindy Edwards and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

• Shurabia serrata stated to come from Shurab, but collection number corresponds to Madygen
Insecta
 Dicondylia - Triadotypidae
"Triadotypus sogdianus n. sp." = Reisia sogdiana19
"Triadotypus sogdianus n. sp." = Reisia sogdiana19 Pritykina 1981 winged insect
PIN 2785/21
 Dicondylia - Zygophlebiidae
Cyrtophlebia sinuosa n. gen. n. sp.13, Cyrtophlebia sinuosa n. sp.19, Zygophlebia ramosa n. gen. n. sp.13, Zygophlebia ramosa n. gen. n. sp.19, Mixophlebia mixta19
Cyrtophlebia sinuosa n. gen. n. sp.13 Pritykina 1981 winged insect
PIN 2785/25
Cyrtophlebia sinuosa n. sp.19 Pritykina 1981 winged insect
PIN 2785/25
Zygophlebia ramosa n. gen. n. sp.13 Pritykina 1981 winged insect
PIN 2785/20
Zygophlebia ramosa n. gen. n. sp.19 Pritykina 1981 winged insect
PIN 2785/20, 4101
Mixophlebia mixta19 Pritykina 1981 winged insect
PIN 2785/8
 Dicondylia - Paurophlebiidae
Paurophlebia lepida n. sp.19 Pritykina 1981 winged insect
PIN 2785/7
Paurophlebia angusta n. sp.19 Pritykina 1981 winged insect
PIN 2785/4097
Neritophlebia longa n. sp.19 Pritykina 1981 winged insect
PIN 2785/24
Neritophlebia vicina n. sp.19 Pritykina 1981 winged insect
PIN 2785/1
Neritophlebia elegans n. gen. n. sp.19 Pritykina 1981 winged insect
PIN 2785/4, 3
 Dicondylia - Batkeniidae
Batkenia pusilla n. gen. n. sp.19
Batkenia pusilla n. gen. n. sp.19 Pritykina 1981 winged insect
PIN 2785/22
 Dicondylia - Kennedyidae
Kennedya gracilis n. sp.19 Pritykina 1981 winged insect
PIN 2785/23
Kennedya carpenteri n. sp.19 Pritykina 1981 winged insect
PIN 2785/2, 6
 Reculida - Geinitziidae
Shurabia serrata n. sp. Aristov et al. 2009 winged insect
PIN 2785/2161
 Hemiptera - Mesojabloniidae
Fulgobole evansi n. gen.22, Scytachile emeljanovi n. gen. n. sp.22
Fulgobole evansi n. gen.22 Shcherbakov 2011 true bug
PIN 2785/3238
Scytachile emeljanovi n. gen. n. sp.22 Shcherbakov 2011 true bug
PIN 2785/3070
 Hemiptera - Curvicubitidae
Beaconiella pulchra n. sp.24 Shcherbakov 2021 true bug
PIN 2785/3022(3073), 3387(3893), 3405
Beaconiella cincta n. sp.24 Shcherbakov 2021 true bug
PIN 2785/3107(3381)
 Hemiptera - Maguviopseidae
Fasolinka beckermigdisovae n. gen. n. sp.22, Asiocula lima n. gen.22
Fasolinka beckermigdisovae n. gen. n. sp.22 Shcherbakov 2011 true bug
PIN 2785/3087&3088
Asiocula lima n. gen.22 Shcherbakov 2011 true bug
PIN 2785/3326
 Hemiptera - Naibiidae
Coccavus supercubitus21 Shcherbakov 2007 true bug
PIN 2785/3544
 Hemiptera - Ipsviciidae
Strivicia davidi25 Shcherbakov 2022 true bug
PIN 2785/2432
 Hemiptera - Serpentivenidae
Serpentivena tigrina n. gen. n. sp.20
Serpentivena tigrina n. gen. n. sp.20 Shcherbakov 1984 true bug
PIN 2785/3535(3635), 3102(3103)
 Mecoptera - Orthophlebiidae
Choristopanorpa temperata n. sp.16 Novokshonov 2001 scorpionfly
PIN 2785/4024
 Mecoptera - Mesopsychidae
Prochoristella longa15 Novokshonov 1997 scorpionfly
PIN 2785/3969
Mesopsyche justa17 Novokshonov and Sukacheva 2001 scorpionfly
PIN 2785/4021+3970
Mesopsyche tortiva n. sp.17 Novokshonov and Sukacheva 2001 scorpionfly
PIN 2785/4052, 3198
Mesopsyche ordinata n. sp.17 Novokshonov and Sukacheva 2001 scorpionfly
PIN 2785/4053
 Mecoptera - Thaumatomeropidae
Thaumatomerope sogdiana n. gen. n. sp.18 Rasnitsyn 1974 scorpionfly
PIN 2785/2496, 3753, 4072
Thaumatomerope oligoneura n. sp.18 Rasnitsyn 1974 scorpionfly
PIN 2785/2498, 3974
 Mecoptera - Parachoristidae
Parachorista immota n. sp.16 Novokshonov 2001 scorpionfly
PIN 2785/4135, 3228
Parachorista arguta n. sp.16 Novokshonov 2001 scorpionfly
PIN 2785/3769, 3887
Triassochorista kirgizica14 Novokshonov 1997 scorpionfly
PIN 2785/3793
 Mecoptera - Permochoristidae
Agetopanorpa deceptoria n. sp.15 Novokshonov 1997 scorpionfly
PIN 2785/4001
Liassochorista ? utilis15 Novokshonov 1997 scorpionfly
PIN 2785/3970
Mesageta rieki15 Novokshonov 1997 scorpionfly
PIN 2785/3841, 3896(3919)
 Coleoptera - Peltosynidae
Ofthalmopeltos synkritos n. gen. n. sp.36, Peltosyne varyvrosa n. sp.36
Ofthalmopeltos synkritos n. gen. n. sp.36 Yan et al. 2017 beetle
PIN 2785/2888
Peltosyne varyvrosa n. sp.36 Yan et al. 2017 beetle
PIN 2785/2655
 Coleoptera - Obrieniidae
Obrieniidae indet.37 Zherikhin and Gratshev 1993 weevil
PIN 2785/2360
Obrienia illaetabilis n. sp.37 Zherikhin and Gratshev 1993 weevil
PIN 2785/3725
 Neuroptera - Archeosmylidae
Madygoneura elongata n. gen. n. sp.12
Madygoneura elongata n. gen. n. sp.12 Khramov 2024 lacewing
PIN 2785/4075
 Neuroptera - Berothidae
Ferganoberotha minutissima n. gen. n. sp.11
Ferganoberotha minutissima n. gen. n. sp.11 Khramov 2023 beaded lacewing
PIN 2785/4061
 Dicondylia - Permithonidae
Relictovia pristina n. gen. n. sp.12
Relictovia pristina n. gen. n. sp.12 Khramov 2024 winged insect
PIN 2785/2489
 Dicondylia - Permosialidae
Permosialis triassica3 Novokshonov and Zhuzhgova 2004 winged insect
PIN 2785/3343
 Blattaria - Caloblattinidae
Thuringoblatta sogdianensis35 Vishnyakova 1998 cockroach
PIN 2785/1739-1742, 1746, 1748, 1750, 1753, 1761, 1765, 4103
Sogdoblatta porrecta n. sp.35 Vishnyakova 1998 cockroach
PIN 2785/1760(1764), 1744, 1745
Sogdoblatta nana35 Vishnyakova 1998 cockroach
PIN 2785/2471, 3724
 Plecoptera - Perlariopseidae
Fritaniopsis brevicaulis27 Sinitshenkova 1987 stonefly
PIN 2785/2740
 Orthoptera - Gryllavidae
Zagryllavus elongatus n. gen. n. sp.6, Paragryllavus curvatus n. gen. n. sp.6
Zagryllavus elongatus n. gen. n. sp.6 Gorochov 1986 grasshopper
PIN 2785/1941
Paragryllavus curvatus n. gen. n. sp.6 Gorochov 1986 grasshopper
PIN 2785/1945(1952), 1955
 Orthoptera - Hagloedischiidae
Hagloedischia primitiva n. gen. n. sp.6
Hagloedischia primitiva n. gen. n. sp.6 Gorochov 1986 grasshopper
PIN 2785/1951
 Orthoptera - Haglidae
Haglomorpha martynovi n. gen. n. sp.6, Lyrohagla uvarovi n. gen. n. sp.6, Zamaraga reticulata n. gen. n. sp.6, "Maraga angusta n. gen. n. sp." = Triassaga angusta6, Modihagla ovalis n. gen. n. sp.6, Tinnihagla zeuneri n. gen. n. sp.6, Paravoliopus dorsalis n. gen. n. sp.6, Voliopellus latus n. gen. n. sp.6, Macrovoliopus declivis n. gen. n. sp.6, Euvoliopus giganteus n. gen. n. sp.6, Voliopus ancestralis n. gen. n. sp.6, Dulcihagla mistshenkoi n. sp.6, Dolichohagla longa n. gen. n. sp.6, Sonohagla curta n. sp.6
Haglomorpha martynovi n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2378(1991)
Lyrohagla uvarovi n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1877
Zamaraga reticulata n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1992
"Maraga angusta n. gen. n. sp." = Triassaga angusta6 Gorochov 1986 hump-winged cricket
PIN 2785/2359
Modihagla ovalis n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2340
Tinnihagla zeuneri n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2343
Paravoliopus dorsalis n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1892(3)
Voliopellus latus n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1861
Macrovoliopus declivis n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1856
Euvoliopus giganteus n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2380
Voliopus ancestralis n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1947
Dulcihagla mistshenkoi n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/1961
Dolichohagla longa n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2368
Sonohagla curta n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2785/2335
 Orthoptera - Mesoedischiidae
Mesoedischia obliqua7 Gorochov 1987 grasshopper
PIN 2785/1940
 Orthoptera - Mesotitanidae
Prototitan sharovi n. sp.9 Gorochov 2003 grasshopper
PIN 2785/2042
 Orthoptera - Paratitanidae
Paratitan reductus n. sp.9 Gorochov 2003 grasshopper
PIN 2785/2021
 Orthoptera - Locustavidae
Miolocustavus reductus n. gen. n. sp.10 Gorochov 2005 grasshopper
PIN 2785/1949
Locustavus minutus n. sp.10 Gorochov 2005 grasshopper
PIN 2785/2349(2353)
Locustavus problematicus10 Gorochov 2005 grasshopper
PIN 2785/2339
Locustavus intermedius n. sp.10 Gorochov 2005 grasshopper
PIN 2785/2347
Brevilocustavus distinctus n. sp.10 Gorochov 2005 grasshopper
PIN 2785/1986
 Paoliida - Paoliidae
Ideliopsina stupenda n. sp.34 Storozhenko 1996 winged insect
PIN 2785/2122
Ideliopsina nana34 Storozhenko 1996 winged insect
PIN 2785/2204, 2120, 2180, 2194, 2233
Ideliopsina ornata34 Storozhenko 1996 winged insect
PIN 2785/2206
 Grylloblattodea -
Pseudoshurabia pallidula34 Storozhenko 1996 ice crawler
PIN 2785/2196
 Grylloblattodea - Gorochoviidae
Gorochovia individua n. gen.32 Storozhenko 1994 ice crawler
Gorochovia minuta32 Storozhenko 1994 ice crawler
Gorochovia bifurca n. sp.32 Storozhenko 1994 ice crawler
PIN 2785/2262,2183,2184,2237
Pseudoliomopterites obscurus n. gen.32 Storozhenko 1994 ice crawler
PIN 2785/2111
Gorochoviella conjuncta n. gen.32 Storozhenko 1994 ice crawler
PIN 2785/2183,2237
 Grylloblattodea - Madygenophlebiidae
Madygenophlebia nana29 Storozhenko 1992 ice crawler
PIN 2785/2169
Madygenophlebia bella29 Storozhenko 1992 ice crawler
PIN 2785/2155,2256
 Cnemidolestodea - Sylvabestiidae
Sharovala triassica n. gen. n. sp.2
Sharovala triassica n. gen. n. sp.2 Aristov 2020 winged insect
PIN 2785/3021
 Embioptera - Alexarasniidae
Madygembia rasnitsyni n. gen. n. sp.5 Aristov and Storozhenko 2021 webspinners
PIN 2785/2162
Nestorembia novojilovi23 Shcherbakov 2015 webspinners
PIN 2785/2252
 Dicondylia - Xiphopteridae
Xiphopterum sharovi n. sp.8 Gorochov 1994 winged insect
PIN 2785/2089, 2085, 2012, 2089a
 Dicondylia - Aeroplanidae
"Paraplana parallelica n. sp." = Sharovoplana parallelica8
"Paraplana parallelica n. sp." = Sharovoplana parallelica8 Gorochov 1994 winged insect
PIN 2785/2090
 Dicondylia - Prochresmodidae
Triassophasma intermedium8 Gorochov 1994 winged insect
PIN 2785/2275
Triassophasma pusillum n. sp.8 Gorochov 1994 winged insect
PIN 2785/2084
Prochresmoda parva n. sp.8 Gorochov 1994 winged insect
PIN 2785/2097, 2098, 2074
 Eoblattida - Batkentakidae
Batkentak intactus n. gen. n. sp.1
Batkentak intactus n. gen. n. sp.1 Aristov 2015 insect
PIN 2785/2179
 Eoblattida - Megakhosaridae
Metakhosara sharovi n. gen. n. sp.31, Megablattogryllus pinguis n. sp.28, Megablattogryllus austerus n. sp.28, "Microblattogryllus variabilis" = Protoblattogryllus variabilis28
Metakhosara sharovi n. gen. n. sp.31 Storozhenko 1993 insect
PIN 2785/2360
Megablattogryllus pinguis n. sp.28 Storozhenko 1990 insect
PIN 2785/2121
Megablattogryllus austerus n. sp.28 Storozhenko 1990 insect
PIN 2785/2178
"Microblattogryllus variabilis" = Protoblattogryllus variabilis28 Storozhenko 1990 insect
PIN 2785/2136, 2171
 Eoblattida - Mesorthopteridae
Sharovites alexanderi4 Aristov and Storozhenko 2013 insect
PIN 2785/2199
Mesoidelia faceta33 Storozhenko 1996 insect
PIN 2785/2107,2167,2234
Parastenaropodites fluxa n. gen.33 Storozhenko 1996 insect
Parastenaropodites longiuscula n. sp.33 Storozhenko 1996 insect
Austroidelia asiatica n. sp.33 Storozhenko 1996 insect
"Parastenaropodites nervosa" = Austroidelia nervosa33 Storozhenko 1996 insect
 Eoblattida - Blattogryllidae
Costatoviblatta aenigmatosa30, "Mesoblattogryllus conjunctus n. sp." = Costatoviblatta conjuncta30, Anoblattogryllus fundatus n. gen. n. sp.28, Baharellus madygensis30
Costatoviblatta aenigmatosa30 Storozhenko 1992 insect
PIN 2785/2126, 2127, 2160, 2201, 2240, 2243
"Mesoblattogryllus conjunctus n. sp." = Costatoviblatta conjuncta30 Storozhenko 1992 insect
PIN 2785/2242, 2264
Anoblattogryllus fundatus n. gen. n. sp.28 Storozhenko 1990 insect
PIN 2785/2166
Baharellus madygensis30 Storozhenko 1992 insect
PIN 2785/2245, 2266
 Diptera - Protorhyphidae
Vymrhyphus tuomikoskii n. gen. n. sp.26
Vymrhyphus tuomikoskii n. gen. n. sp.26 Blagoderov 1995 fly
PIN 2785/3052
 Diptera - Gnomuscidae
Gnomusca molecula n. gen. n. sp.26
Gnomusca molecula n. gen. n. sp.26 Shcherbakov 1995 fly
PIN 2785/4096
 Diptera - Psychotipidae
Psychotipa predicta n. gen. n. sp.26
Psychotipa predicta n. gen. n. sp.26 Shcherbakov 1995 fly
PIN 2785/2490