Karatau-Mikhailovka (PIN collection 2784) (Jurassic to of Kazakhstan)

Where: Chimkent, Kazakhstan (42.9° N, 70.0° E: paleocoordinates 41.6° N, 73.7° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Karabastau Formation, Callovian to Callovian (166.1 - 157.3 Ma)

Environment/lithology: lacustrine - large; lithified siltstone

• "dark shales" Grimaldi & Engel, 2005

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Reposited in the PIN

Primary reference: A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [C. Beard/C. Beard/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 113439: authorized by Matthew Clapham, entered by Jered Karr on 31.07.2011, edited by Matthew Clapham, Andrew Spencer-Lee and Lindy Edwards

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hemiptera - Tettigarctidae
Sunotettigarcta kudryashevae n. sp.42 Shcherbakov 2008 hairy cicada
PIN 2784/2163
 Thysanoptera - Liassothripidae
Liassothrips crassipes43 Martynov 1927 thrips
PIN 2784/2207-2209, 2268, 2269
 Permopsocida - Archipsyllidae
Archipsylla similis n. sp.46 Vishnyakova 1976 winged insect
PIN 2784/2341, 2343
 Psocodea - Electrentomidae
Paramesopsocus adibi n. sp.2 Azar et al. 2008 book louse
PIN 2784/2096, 1998, 2001, 2340; 2339 ("Mesopsocidae in Rasnitsyn, 2002" according to Mockford et al., 2013)
 Blattaria - Skokidae
Skok svaba48 Vršanský 2007 jumping cockroach
PIN 2784/684
 Plecoptera - Baleyopterygidae
Udopteryx expansa44 Sinichenkova 1985 stonefly
PIN 2784/243
 Plecoptera - Perlariopseidae
Karanemoura abrupta45 Sinitshenkova 1987 stonefly
PIN 2784/208
 Blattodea - Liberiblattinidae
Hydrokhoohydra aquabella49 Vršanský 2019 cockroach
PIN 2784/789, 982
 Dermaptera - Semenoviolidae
Semenovioloides capitatus n. gen. n. sp.47
Semenovioloides capitatus n. gen. n. sp.47 Vishnyakova 1980 earwig
PIN 2784/2243
 Dermaptera - Dermapteridae
"Dermapteron incertae" = Dermapteron incerta47
"Dermapteron incertae" = Dermapteron incerta47 Martynov 1925 earwig
PIN 2784/2240
 Dermaptera - Protodiplatyidae
Protodiplatys fortis47 Martynov 1925 earwig
PIN 2784/2249
 Dermaptera - Turanoviidae
Turanovia incompleta n. gen. n. sp.47
Turanovia incompleta n. gen. n. sp.47 Vishnyakova 1980 earwig
PIN 2784/2250(2252)
 Orthoptera - Protogryllidae
"Protogryllus karatavicus" = Falsispeculum karatavicus
"Protogryllus karatavicus" = Falsispeculum karatavicus Sharov 1968 grasshopper
PIN 2784/3
 Orthoptera - Prophalangopsidae
? Aboilus amplus n. sp.7, Aboilus zebra n. sp.7, Karatailus micropterus n. gen. n. sp.7
? Aboilus amplus n. sp.7 Gorochov 1996 hump-winged cricket
PIN 2784/2360, 59
Aboilus zebra n. sp.7 Gorochov 1996 hump-winged cricket
PIN 2784/34
Karatailus micropterus n. gen. n. sp.7 Gorochov 1996 hump-winged cricket
PIN 2784/2275
 Orthoptera - Phasmomimidae
Jurophasmomima punctata n. gen. n. sp.6
Jurophasmomima punctata n. gen. n. sp.6 Gorochov 1988 grasshopper
PIN 2784/9
 Hymenoptera - Ephialtitidae
Leptephialtites stephanocephalus n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1209
Leptephialtites picturatus n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1200
Leptephialtites linearis n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1201
Leptephialtites caudatus n. gen. n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1206
Leptephialtites angustus n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1199
Parephialtites reductus n. gen. n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1177
Sessiliventer oppositus n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1207
Stephanogaster magna n. gen. n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1205
Stephanogaster longipalpa40 Rasnitsyn 1975 wasp
PIN 2784/1184, 1178
Symphytopterus minutus n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1211
Symphytopterus robustus40 Rasnitsyn 1975 wasp
PIN 2784/1183
Symphytopterus macrophthalmus n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1180
Symphyogaster cylindrica n. gen. n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1198
 Hymenoptera - Mesoserphidae
Auliserphus niger n. sp.41, Campturoserphus obscurus n. gen. n. sp.41
Auliserphus niger n. sp.41 Rasnitsyn 1986 wasp
PIN 2784/1227
Campturoserphus obscurus n. gen. n. sp.41 Rasnitsyn 1986 wasp
PIN 2784/1236
 Hymenoptera - Praeaulacidae
Praeaulacus obscurus n. sp.39 Rasnitsyn 1972 wasp
PIN 2784/1191
Praeaulacus leptogaster n. sp.39 Rasnitsyn 1972 wasp
PIN 2784/1203
Praeaulacites pachygaster n. gen. n. sp.39 Rasnitsyn 1972 wasp
PIN 2784/1185, 1202, 1208
Praeaulacon tenuis n. sp.39 Rasnitsyn 1972 wasp
PIN 2784/1182
 Hymenoptera - Megalyridae
Mesaulacinus sculpturatus n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1193
Leptocleistogaster alboscuta40 Rasnitsyn 1975 wasp
PIN 2784/1192
Leptocleistogaster pallida n. gen. n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1175
 Hymenoptera - Bethylonymidae
Bethylonymellus cervicalis n. gen. n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1189
Bethylonymellus pallens n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1195
Bethylonymellus bethyloides n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1195
 Hymenoptera - Brachycleistogastridae
Brachycleistogaster nigripes n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1176
Brachycleistogaster jurassica n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1174, 1194
 Hymenoptera - Kuafuidae
Leptogastrella leptogastra n. gen. n. sp.40
Leptogastrella leptogastra n. gen. n. sp.40 Rasnitsyn 1975 wasp
PIN 2784/1204
 Hymenoptera - Karatavitidae
Karatavites medius n. sp.38 Rasnitsyn 1969 wood wasp
PIN 2784/1169
 Hymenoptera - Anaxyelidae
Urosyntexis depressa n. sp.38 Rasnitsyn 1969 winged insect
PIN 2784/1160
Brachysyntexis nova n. sp.38 Rasnitsyn 1969 winged insect
PIN 2784/1159
Sphenosyntexis pallicornis n. sp.38 Rasnitsyn 1969 winged insect
PIN 2784/1162
 Hymenoptera - Protosiricidae
Protosirex xyelopterus n. gen. n. sp.38
Protosirex xyelopterus n. gen. n. sp.38 Rasnitsyn 1969 winged insect
PIN 2784/1164
 Hymenoptera - Sepulcidae
Xyelula hybrida n. gen. n. sp.38, Sirecomima xiphophora n. gen. n. sp.38
Xyelula hybrida n. gen. n. sp.38 Rasnitsyn 1969 stem sawfly
PIN 2784/1161
Sirecomima xiphophora n. gen. n. sp.38 Rasnitsyn 1969 stem sawfly
PIN 2784/1173
 Hymenoptera -
"Mesolyda depressa n. sp." = Prolyda depressa38
"Mesolyda depressa n. sp." = Prolyda depressa38 Rasnitsyn 1969 sawfly
PIN 2784/1163
 Hymenoptera - Xyelidae
Microxyelecia brachycera n. gen. n. sp.38
Microxyelecia brachycera n. gen. n. sp.38 Rasnitsyn 1969 sawfly
PIN 2784/1172
 Coleoptera - Cerophytidae
"Idiomerus latissimus n. sp." = Necromera latissima4, "Idiomerus brevicornis n. sp." = Necromera brevicornis4
"Idiomerus latissimus n. sp." = Necromera latissima4 Dolin 1980 beetle
PIN 2784/1380
"Idiomerus brevicornis n. sp." = Necromera brevicornis4 Dolin 1980 beetle
PIN 2784/1374
 Coleoptera - Elateridae
Litholacon conicicollis n. sp.4 Dolin 1980 click beetle
PIN 2784/1371
Codemus micros4 Dolin 1980 click beetle
PIN 2784/1384
Codemus carinatus4 Dolin 1980 click beetle
PIN 2784/1405, 1372, 1399
Codemus alatus4 Dolin 1980 click beetle
PIN 2784/1368, 1391
Codemus zherichini4 Dolin 1980 click beetle
PIN 2784/1365, 1367
Codemus sharovi4 Dolin 1980 click beetle
PIN 2784/1378
Lapidostenus insignis n. sp.4 Dolin 1980 click beetle
PIN 2784/1379
Lapidostenus longicornis n. sp.4 Dolin 1980 click beetle
PIN 2784/1393
Hypnomorphus rasnitzyni n. sp.4 Dolin 1980 click beetle
PIN 2784/1362
Lithoptychus incertus4 Dolin 1980 click beetle
PIN 2784/1373, 1390, 1398
Negastrioides globicollis4 Dolin 1980 click beetle
PIN 2784/1375
Lithosomus longicollis n. sp.4 Dolin 1980 click beetle
PIN 2784/1363
Lapidiconides innatus n. sp.4 Dolin 1980 click beetle
PIN 2784/1376(1400)
Platyelater reflexicollis4 Dolin 1980 click beetle
PIN 2784/1387
Lithomerus cockerelli4 Dolin 1980 click beetle
PIN 2784/1366, 1389
Acheonus gracilis n. sp.4 Dolin 1980 click beetle
PIN 2784/1404
Desmatus lapidarius3 Dolin 1975 click beetle
PIN 2784/1377
Plesiorhaphes scaber n. gen. n. sp.4 Dolin 1980 click beetle
PIN 2784/1383
 Coleoptera - Anthribidae
Juranthribus thompsoni n. gen.19 Legalov 2011 fungus weevil
PIN 2784/1657
Protoscelis parvula21 Medvedev 1968 fungus weevil
PIN 2784/1434,1435,1437,1438,1442,1444,1445
Protoscelis nitidicornis21 Medvedev 1968 fungus weevil
PIN 2784/1430,1440
Protoscelis jurassica21 Medvedev 1968 fungus weevil
PIN 2784/1426,1441,1446
Protoscelis longicornis21 Medvedev 1968 fungus weevil
PIN 2784/1426,1428,1429,1436
Protoscelis medvedevi21 Legalov 2013 fungus weevil
PIN 2784/1433,1461
 Coleoptera - Nemonychidae
Astenorrhinus major9 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1458
"Pseudobrenthorrhinus tenuicornis n. sp." = Eccoptarthrus tenuicornis10 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1493(1517)
"Distenorrhinus (Distenorrhinus) pallidirostris" = Distenorrhinus (Distenorrhinus) pallidirostris11 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1681, 1582
"Distenorrhinus (Distenorrhinus) rotundicollis n. sp." = Distenorrhinus (Distenorrhinus) pallidirostris11 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1483(1502), 1493, 1471, 1774
Distenorrhinus (Distenorrhinus) antennatus11 Arnoldi 1977 pine flower snout beetle
PIN 2784/1488
"Brenthorrhinoides pubescens n. sp." = Paleocartus pubescens10 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2784/1464
"Ampliceps furcitibia" = Ampliceps furcitibia9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1460, 1462, 1467, 1468, 1472, 1487, 1488, 1491, 1495, 1501, 1503, 1506, 1508, 1515, 1516, 1524
"Medmetrioxenoides prodromus n. gen. n. sp." = Ampliceps furcitibia8 Arnoldi 1977 pine flower snout beetle
PIN 2784/1486
Oxycorynoides (Oxycorynoides) similis9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1465, 1466, 1469, 1470, 1473, 1475-1479, 1481, 1485, 1490, 1492, 1494, 1497, 1498, 1500, 1504, 1507, 1509-1514, 1519-1523, 1913
Archaeorrhynchus carpenteri n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1453, 1456
Probelus (Probelus) scudderi n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1457
Probelus (Probelus) longitarsus9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1505
Probelus (Probelus) curvispinus9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1448
Probelus (Probelus) handlirschi n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1451, 1455
Probelus (Probelus) nikolaevi n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1449
Belonotaris (Arnoldibelus) gratshevi9 Legalov 2010 pine flower snout beetle
PIN 2784/1450
Belonotaris (Arnoldibelus) karatavicus9 Arnoldi 1977 pine flower snout beetle
PIN 2784/1454, 1459, 1499
Belonotaris (Arnoldibelus) rohdendorfi n. sp.9 Legalov 2014 pine flower snout beetle
PIN 2784/1452
Belonotaris (Arnoldibelus) heeri9 Legalov 2014 pine flower snout beetle
PIN 2784/1518
 Coleoptera - Brentidae
Nanophydes ovatus22 Arnoldi 1977 weevil
PIN 2784/1496
 Coleoptera - Hybosoridae
Protohybosorus karatavicus n. gen. n. sp.34
Protohybosorus karatavicus n. gen. n. sp.34 Nikolajev 2010 scavenger scarab beetle
PIN 2784/1656
 Coleoptera - Coptoclavidae
Charonoscapha ovata1 Ponomarenko 1977 beetle
PIN 2784/1925
 Coleoptera - Trachypachidae
Karatoma agilis n. gen. n. sp.1
Karatoma agilis n. gen. n. sp.1 Ponomarenko 1977 false ground beetle
PIN 2784/1528
 Coleoptera - Carabidae
Ovrabites ovalis1 Ponomarenko 1977 ground beetle
PIN 2784/1326
Protorabus planus1 Ponomarenko 1977 ground beetle
PIN 2784/1299
Cordorabus notatus1 Ponomarenko 1977 ground beetle
PIN 2784/1320
Mesorabus elongatus n. gen. n. sp.1 Ponomarenko 1977 ground beetle
PIN 2784/1525
 Coleoptera - Obrieniidae
Kararhynchus jurassicus n. sp.20 Legalov 2012 beetle
PIN 2784/1602
 Mecoptera - Bittacidae
Prohylobittacus sorosi n. gen. n. sp.35
Prohylobittacus sorosi n. gen. n. sp.35 Novokshonov 1993 hangingfly
PIN 2784/2219
 Mecoptera - Nannochoristidae
Itaphlebia sharovi36 Novokshonov 1997 scorpionfly
PIN 2784/2223
 Diptera -
"? Xylomya shcherbakovi n. sp." = Xylomyia shcherbakovi31
"? Xylomya shcherbakovi n. sp." = Xylomyia shcherbakovi31 Mostovski 1999 fly
PIN 2784/131
 Diptera - Archisargidae
Archisargidae indet.28, Calosargus (Calosargus) niger n. sp.29, ? Sharasargus spiniger n. sp.28, Parvisargus malus n. sp.28, "? Prosolva karataviensis n. sp." = Mesosolva karataviensis28
Archisargidae indet.28 Rohdendorf 1962 fly
PIN 2784/357
Calosargus (Calosargus) niger n. sp.29 Mostovski 1997 fly
PIN 2784/77
? Sharasargus spiniger n. sp.28 Mostovski 1996 fly
PIN 2784/344
Parvisargus malus n. sp.28 Mostovski 1996 fly
PIN 2784/430
"? Prosolva karataviensis n. sp." = Mesosolva karataviensis28 Mostovski 1996 fly
PIN 2784/292
 Diptera - Rhagionempididae
Rhagionempis antennata15 Rohdendorf 1938 fly
PIN 2784/88
 Diptera - Nemestrinidae
Protonemestrius rohdendorfi n. sp.30 Mostovski 1998 tangle-veined fly
PIN 2784/73
Archinemestrius karatavicus30 Rohdendorf 1968 tangle-veined fly
PIN 2784/621
Archinemestrius litigiosus n. sp.30 Mostovski 1998 tangle-veined fly
PIN 2784/79
 Diptera - Protorhyphidae
Archirhyphus asiatcus25 Rohdendorf 1964 fly
PIN 2784/211, 598
 Diptera - Elliidae
Polyanka minuta16 Krzemińska et al. 1993 fly
PIN 2784/388, 407, 414
 Diptera - Trichoceridae
Tanyochoreta (Trichokara) composita17 Krzemińska et al. 2009 winter crane fly
PIN 2784/280, 285
Tanyochoreta (Trichokara) minuta17 Krzemińska et al. 2009 winter crane fly
PIN 2784/475
Zherikhinina zherikhini17 Krzemińska et al. 2009 winter crane fly
PIN 2784/346
 Diptera - Tanyderidae
Praemacrochile ansorgei27 Lukashevich and Krzemiński 2009 primitive crane fly
PIN 2784/151
 Diptera - Ptychopteridae
Nedoptychoptera karatavica n. gen. n. sp.26 Lukashevich 1998 phantom crane fly
PIN 2784/261
Eoptychopterina angularis n. sp.24 Lukashevich 1993 phantom crane fly
PIN 2784/74
 Diptera - Ansorgiidae
"Ansorgia praedicta n. gen. n. sp." = Ansorgius praedicta18
"Ansorgia praedicta n. gen. n. sp." = Ansorgius praedicta18 Krzemiński and Lukashevich 1993 fly
PIN 2784/114, 410, 428
 Neuroptera -
"Nymphoides latus n. gen. n. sp." = Osmylites latus37, Chrysoleonites intactus n. sp.37
"Nymphoides latus n. gen. n. sp." = Osmylites latus37 Panfilov 1980 lacewing
PIN 2784/1046
Chrysoleonites intactus n. sp.37 Panfilov 1980 lacewing
PIN 2784/1048
 Neuroptera - Psychopsidae
Propsychops karatavicus n. gen. n. sp.37
Propsychops karatavicus n. gen. n. sp.37 Panfilov 1980 silky lacewing
PIN 2784/1060
 Neuroptera - Kalligrammatidae
Meioneurites (Parameioneurites) spectabilis n. sp.5 Engel 2005 lacewing
PIN 2784/1069
Kalligrammula karatensis n. sp.23 Liu et al. 2015 lacewing
PIN 2784/1032
 Neuroptera - Chrysopidae
Mesypochrysa reducta n. sp.37 Panfilov 1980 green lacewing
PIN 2784/1059
 Neuroptera - Osmylidae
Ensiosmylus acutus n. gen. n. sp.13
Ensiosmylus acutus n. gen. n. sp.13 Khramov 2014 lance lacewing
PIN 2784/1020
 Neuroptera - Mantispidae
"Longicollum benmaddoxi n. gen. n. sp." = Longipronotum benmaddoxi12, Promantispa similis n. gen. n. sp.37
"Longicollum benmaddoxi n. gen. n. sp." = Longipronotum benmaddoxi12 Jepson et al. 2018 mantidfly
PIN 2784/1056
Promantispa similis n. gen. n. sp.37 Panfilov 1980 mantidfly
PIN 2784/1080
 Neuroptera - Berothidae
Berothone protea14, Mesithone magna n. sp.37, Mesithone maculata n. gen. n. sp.37
Berothone protea14 Panfilov 1980 beaded lacewing
PIN 2784/1061
Mesithone magna n. sp.37 Panfilov 1980 beaded lacewing
PIN 2784/1025
Mesithone maculata n. gen. n. sp.37 Panfilov 1980 beaded lacewing
PIN 2784/1058
 Dicondylia - Protomyrmeleontidae
Protomyrmeleon sp.33 Geinitz 1888 winged insect
PIN 2784/69
Protomyrmeleon kazakhstanensis n. sp.33 Nel et al. 2005 winged insect
PIN 2784/70-71(pc)
Protomyrmeleon grandis n. sp.33 Nel et al. 2005 winged insect
PIN 2784/67
 Odonata - Paragonophlebiidae
Paragonophlebia inexpectata n. gen. n. sp.32
Paragonophlebia inexpectata n. gen. n. sp.32 Nel 2009 dragonfly
PIN 2784/08