Kubekovo village (PIN collection 1255) (Jurassic to of Russian Federation)

Where: Krasnoyarsk, Russian Federation (56.2° N, 93.2° E: paleocoordinates 59.6° N, 90.4° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Upper Member (Itat Formation), Bajocian to Bajocian (170.3 - 166.1 Ma)

Environment/lithology: lacustrine - large; claystone

• The Middle Jurassic locality of Kubekovo, Krasnoyarsk Region, is formed by deposits of a very large lake (Zherikhin, 1985). Late thermal subsidence phase of West Siberian Basin.
• Clays underlying coaly shales

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by V.G. Kovalev; reposited in the PIN

Primary reference: N. D. Sinichenkova. 1985. New Jurassic Stone Flies of the family Baleyopterygidae. Paleontological Journal 18(4):129-134 [M. Clapham/J. Karr/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 118010: authorized by Matthew Clapham, entered by Jered Karr on 30.09.2011, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Odonata - Campterophlebiidae
Campterophlebiidae indet.16 Handlirsch 1920 dragonfly
PIN 1255/2-4
 Ephemeroptera - Siphlonuridae
Mesobaetis sp.23 Brauer et al. 1889 mayfly
PIN 1255/8-10,19,20,39-41,43-45,47,51,54,55,59,60,64,69
 Blattodea - Blattulidae
"Chiloblattula sibirica n. sp." = Blattula sibirica28
"Chiloblattula sibirica n. sp." = Blattula sibirica28 Vishnyakova 1985 cockroach
PIN 1255/72
 Blattodea - Liberiblattinidae
Eublattula curvinervis n. sp.28 Vishnyakova 1985 cockroach
PIN 1255/78
 Plecoptera - Baleyopterygidae
Plutopteryx popovi n. sp. Sinichenkova 1985 stonefly
PIN 1255/480, 481, 482
Udopteryx kovalevi n. sp. Sinichenkova 1985 stonefly
PIN 1255/483, 484
 Plecoptera - Platyperlidae
Platyperla conferta n. sp.22 Sinitshenkova 1985 stonefly
PIN 1255/274,275
 Dermaptera - Protodiplatyidae
Microdiplatys perfectus n. sp.29 Vishnyakova 1985 earwig
PIN 1255/88
 Dicondylia -
Orthoptera indet.31 Olivier 1789 grasshopper
PIN 1255/92
 Orthoptera - Locustopsidae
? Locustopsis spp.31 Handlirsch 1906 grasshopper
PIN 1255/93.94
Locustopsis picta n. sp.31 Zherikhin 1985 grasshopper
PIN 1255/95
 Dicondylia -
Neuroptera indet.2 Linnaeus 1758 lacewing
PIN 1255/397, 1106
 Neuroptera - Saucrosmylidae
Thaumatomerobius mirabilis n. gen. n. sp.13
Thaumatomerobius mirabilis n. gen. n. sp.13 Ponomarenko 1985 elegant lance lacewing
PIN 1255/136; 1255/1090 in Khramov, 2017, who also claims 1255/138 to fragmentary to identify
 Neuroptera - Osmylidae
Osmylidae indet.2 Leach 1815 lance lacewing
PIN 1255/1096, 1097
Kubekius multiramosus n. gen. n. sp.2 Khramov 2017 lance lacewing
PIN 1255/1099
Mesosmylina sibirica n. sp.13 Ponomarenko 1985 lance lacewing
PIN 1255/139, 135
Epiosmylus longus n. sp.2 Khramov 2017 lance lacewing
PIN 1255/1100, 1092
 Neuroptera - Psychopsidae
? Psychopsidae indet.2 Handlirsch 1906 silky lacewing
PIN 1255/1094
 Coleoptera -
"Strongulites stygicus" = Strongylites stygius12, Oxytoroptera punctata n. sp.12
"Strongulites stygicus" = Strongylites stygius12 Heer 1864 beetle
PIN 1255/100
Oxytoroptera punctata n. sp.12 Ponomarenko 1985 beetle
PIN 1255/122
 Coleoptera - Permosynidae
Hydrobiites minutissimus n. sp.12 Ponomarenko 1985 beetle
PIN 1255/121
Hydrobiites bellus12 Geinitz 1894 beetle
PIN 1255/111
Dzeregia lata12 Ponomarenko 1985 beetle
PIN 1255/117,126,130
Platycrossos punctatus n. sp.12 Ponomarenko 1985 beetle
PIN 1255/120
Platycrossos petalus n. sp.12 Ponomarenko 1985 beetle
PIN 1255/129
Platycrossos elongatus n. sp.12 Ponomarenko 1985 beetle
PIN 1255/105
 Coleoptera -
Prosthecogmus additus n. gen. n. sp.12
Prosthecogmus additus n. gen. n. sp.12 Ponomarenko 1985 beetle
PIN 1255/115
 Coleoptera - Staphylinidae
Morsum abdominale n. gen. n. sp.17
Morsum abdominale n. gen. n. sp.17 Ryvkin 1985 rove beetle
PIN 1255/134
 Coleoptera -
Polysitum elongatum n. sp.12 Ponomarenko 1985 beetle
PIN 1255/123
Polysitum latum n. sp.12 Ponomarenko 1985 beetle
PIN 1255/128
 Coleoptera - Trogossitidae
Thoracotes sibiricus n. sp.12 Ponomarenko 1985 bark gnawing beetle
PIN 1255/119, 127
 Coleoptera - Buprestidae
Agrillium minor n. sp.12
Agrillium minor n. sp.12 Ponomarenko 1985 jewel beetle
PIN 1255/132
 Coleoptera -
Plastelater glabratus n. sp.12 Ponomarenko 1985 beetle
PIN 1266/116
Adephaga indet.12 Clairville 1806 beetle
PIN 1255/104
Memptus redtenbacheri12 Handlirsch 1906 beetle
PIN 1255/110
Memptus handlirschi12 Ponomarenko 1985 beetle
PIN 1255/97,98,101,102,106,108,114
Memptus guttula n. sp.12 Ponomarenko 1985 beetle
PIN 1255/1,96,99,103,109a,g,112
 Dicondylia -
Mecoptera indet.24 Packard 1886 scorpionfly
PIN 1255/187
 Mecoptera - Mesopsychidae
Tarantogus sibiricus n. sp.10, Tarantogus obscurus n. sp.24, Tarantogus opiparus n. gen. n. sp.24
Tarantogus sibiricus n. sp.10 Novokshonov 1997 scorpionfly
PIN 1255/1546
Tarantogus obscurus n. sp.24 Sukatsheva 1985 scorpionfly
PIN 1255/185
Tarantogus opiparus n. gen. n. sp.24 Sukatsheva 1985 scorpionfly
PIN 1255/181
 Mecoptera - Nannochoristidae
Itaphlebia reducta n. sp.10, Itaphlebia jeniseica n. sp.10, Itaphlebia completa n. gen. n. sp.24, Itaphlebia multa n. sp.10
Itaphlebia reducta n. sp.10 Novokshonov 1997 scorpionfly
PIN 1255/1551
Itaphlebia jeniseica n. sp.10 Novokshonov 1997 scorpionfly
PIN 1255/1547
Itaphlebia completa n. gen. n. sp.24 Sukatsheva 1985 scorpionfly
PIN 1255/178
Itaphlebia multa n. sp.10 Novokshonov 1997 scorpionfly
PIN 1255/1555, 1548, 1552, 1554, and 10 others
 Mecoptera - Meropeidae
Boreomerope antiqua n. gen. n. sp.9
Boreomerope antiqua n. gen. n. sp.9 Novokshonov 1995 earwigfly
PIN 1255/184
 Mecoptera - Orthophlebiidae
"Orthophlebiinae indet." = Orthophlebiidae24, Orthophlebia sp.24, Orthophlebia obunca n. sp.24, Mesopanorpa sp.24
"Orthophlebiinae indet." = Orthophlebiidae24 Handlirsch 1906 scorpionfly
PIN 1255/184, 197
Orthophlebia sp.24 Westwood 1845 scorpionfly
PIN 1255/180
Orthophlebia obunca n. sp.24 Sukatsheva 1985 scorpionfly
PIN 1255/179
Mesopanorpa sp.24 Handlirsch 1906 scorpionfly
PIN 1255/177, 182, 186
 Mecoptera - Bittacidae
"Microbittacus kubekovensis n. gen. n. sp." = Mesobittacus kubekovensis8
"Microbittacus kubekovensis n. gen. n. sp." = Mesobittacus kubekovensis8 Novokshonov 1993 hangingfly
PIN 1255/1545
 Necrotrichoptera - Necrotauliidae
Necrotauliidae indet.25 Handlirsch 1906 winged insect
PIN 1255/191
Necrotaulius communis n. sp.26 Sukatsheva and Sinitshenkova 2023 winged insect
PIN 1255/670
Necrotaulius kubekovi n. sp.25 Sukatsheva 1985 winged insect
PIN 1255/193
 Trichoptera - Philopotamidae
Oncovena borealis n. gen. n. sp.11 Sukacheva and Novokshonov 1995 finger-net caddisfly
PIN 1255/990, 991
Archiphilopotamus kubekovensis n. sp.27 Sukatsheva and Vasilenko 2020 finger-net caddisfly
PIN 1255/1594, 1591, 1601
Archiphilopotamus absurdus n. sp.27 Sukatsheva and Vasilenko 2020 finger-net caddisfly
PIN 1255/696
Archiphilopotamus mancus n. sp.25 Sukatsheva 1985 finger-net caddisfly
PIN 1255/188; 1255/1607 in Sukatsheva & Vasilenko 2020
Archiphilopotamus luxus n. gen. n. sp.25 Sukatsheva 1985 finger-net caddisfly
PIN 1255/190; 1255/693, 1591 in Sukatsheva & Vasilenko 2020
Juraphilopotamus similis n. sp.26 Sukatsheva and Sinitshenkova 2023 finger-net caddisfly
PIN 1255/665
Juraphilopotamus funeralis n. sp.27 Sukatsheva and Vasilenko 2020 finger-net caddisfly
PIN 1255/1593
 Hemiptera - Karabasiidae
Karabasia kovalevi n. sp.15 Popov and Shcherbakov 1991 true bug
PIN 1255/449
Karabasia jeneiseica n. sp.14 Popov 1985 true bug
PIN 1255/170
 Hemiptera - Progonocimicidae
Progonocimicidae indet.14, "Olgamartynovia aff. admota" = Cicadocoris admotus14, "Olgamartynovia aff. frater" = Cicadocoris frater14, Olgamartynovia aff. nana14
Progonocimicidae indet.14 Handlirsch 1906 true bug
PIN 1255/167
"Olgamartynovia aff. admota" = Cicadocoris admotus14 Popov 1982 true bug
PIN 1255/169
"Olgamartynovia aff. frater" = Cicadocoris frater14 Popov 1982 true bug
PIN 1255/168
Olgamartynovia aff. nana14 Popov 1982 true bug
PIN 1255/166
 Hemiptera - Archegocimicidae
Kubecora ignamica n. gen. n. sp.18
Kubecora ignamica n. gen. n. sp.18 Ryzhkova 2013 shore bug
PIN 1255/447
 Hemiptera - Naucoridae
Naucoridae indet.14 Fallen 1814 creeping water bug
PIN 1255/131
 Hemiptera - Archijassidae
Kisa fasciata n. gen. n. sp.20, Kemobius lux n. gen. n. sp.20, Kubecola guttatus n. gen. n. sp.20
Kisa fasciata n. gen. n. sp.20 Shcherbakov 2012 true bug
PIN 1255/1562, 429, 1563-1567
Kemobius lux n. gen. n. sp.20 Shcherbakov 2012 true bug
PIN 1255/1561
Kubecola guttatus n. gen. n. sp.20 Shcherbakov 2012 true bug
PIN 1255/1570, 1571, 1572, 148, 1568, 1569, 1573
 Hemiptera - Dysmorphoptilidae
Eumorphoptila pritykinae n. gen. n. sp.21
Eumorphoptila pritykinae n. gen. n. sp.21 Shcherbakov 2022 true bug
PIN 1255/1608
 Hemiptera - Szelegiewicziidae
Tinaphis sibirica n. sp.30 Wegierek 1989 aphid
PIN 1255/990, 991
 Hemiptera - Naibiidae
Panirena tenella n. sp.19 Shcherbakov 2007 true bug
PIN 1255/992
 Diptera - Psychodidae
Protanyderus invalidus n. sp.7 Lukashevich and Krzemiński 2009 moth fly
PIN 1255/994
 Diptera - Canthyloscelidae
Prohyperoscelis jurassicus n. gen. n. sp.3 Kovalev 1985 fly
PIN 1255/262, 263
 Diptera - Chironomidae
? Podonomius undulatus n. sp.1 Kalugina 1985 non biting midge
PIN 1255/293
? Podonomius rotundatus n. sp.1 Kalugina 1985 non biting midge
PIN 1255/238, 216, 224, 236, 237, 241, 260, 295, 296
? Podonomius minimus n. sp.1 Kalugina 1985 non biting midge
PIN 1255/202, 201, 217, 241, 294, 296
? Podonomius simplex n. sp.1 Kalugina 1985 non biting midge
PIN 1255/227
 Diptera - Mesosciophilidae
Mesosciophilina irinae n. gen. n. sp.3 Kovalev 1985 fly
PIN 1255/252
Mesosciophilina bolshakovi n. sp.3 Kovalev 1985 fly
PIN 1255/259
 Diptera - Pediciidae
"Praearchitipula notabilis n. gen. n. sp." = Praearchitipula notabilis1, "? Praearchitipula lata n. sp." = Praearchitipula notabilis1, Praearchitipula trinervis n. sp.1
"Praearchitipula notabilis n. gen. n. sp." = Praearchitipula notabilis1 Kalugina 1985 hairy-eyed cranefly
PIN 1255/219
"? Praearchitipula lata n. sp." = Praearchitipula notabilis1 Kalugina 1985 hairy-eyed cranefly
PIN 1255/231
Praearchitipula trinervis n. sp.1 Kalugina 1985 hairy-eyed cranefly
PIN 1255/223
 Diptera - Limoniidae
Limoniidae indet.1 Speiser 1909 crane fly
PIN 1255/209
Eotipulina fumosa n. sp.1 Kalugina 1985 crane fly
PIN 1255/228
Architipula symmetrica n. sp.1 Kalugina 1985 crane fly
PIN 1255/204, 218
Architipula orientalis n. sp.1 Kalugina 1985 crane fly
PIN 1255/222
"Praearchitipula parvula n. sp." = Mesotipula (Irenatipula) parvula1 Kalugina 1985 crane fly
PIN 1255/232
 Diptera - Trichoceridae
"? Trichocera itatica n. sp." = Zherikhinina itatica1
"? Trichocera itatica n. sp." = Zherikhinina itatica1 Kalugina 1985 winter crane fly
PIN 1255/198
 Diptera - Ptychopteridae
? Crenoptychoptera sp.1, Crenoptychoptera antica n. sp.4, Crenoptychoptera defossa n. sp.4, "Eoptychopterina glaber n. sp." = Eoptychopterina glabra4, "Proptychoptera paramaculata n. gen. n. sp." = Eoptychoptera paramaculata1, "Proptychoptera modica n. sp." = Eoptychoptera modica4, "? Proptychopterina maxima n. sp." = Eoptychoptera maxima1, "Eoptychopteridae indet." = Ptychopteridae1, Proptychopterina handlirschi n. sp.1, Proptychopterina amota n. sp.4, Proptychopterina yeniseica n. sp.4
? Crenoptychoptera sp.1 Kalugina 1985 phantom crane fly
PIN 1255/199, 205
Crenoptychoptera antica n. sp.4 Kalugina 1985 phantom crane fly
PIN 1255/211, 777, 975, 1509, 1515
Crenoptychoptera defossa n. sp.4 Kalugina 1985 phantom crane fly
PIN 1255/215, 779, 780, 1516-1518
"Eoptychopterina glaber n. sp." = Eoptychopterina glabra4 Lukashevich 1993 phantom crane fly
PIN 1255/1501-1503
"Proptychoptera paramaculata n. gen. n. sp." = Eoptychoptera paramaculata1 Kalugina 1985 phantom crane fly
PIN 1255/200
"Proptychoptera modica n. sp." = Eoptychoptera modica4 Lukashevich 1993 phantom crane fly
PIN 1255/1504
"? Proptychopterina maxima n. sp." = Eoptychoptera maxima1 Kalugina 1985 phantom crane fly
PIN 1255/230, 1505
"Eoptychopteridae indet." = Ptychopteridae1 Osten-Sacken 1862 phantom crane fly
PIN 1255/276-279, 284, 288, 297-299, 271, 272, 195, 235, 270
Proptychopterina handlirschi n. sp.1 Kalugina 1985 phantom crane fly
PIN 1255/229, 210
Proptychopterina amota n. sp.4 Lukashevich 1993 phantom crane fly
PIN 1255/1508
Proptychopterina yeniseica n. sp.4 Lukashevich 1993 phantom crane fly
PIN 1255/1506, 1507, 1510-1514, 1519-1522
 Diptera - Rhagionidae
Kubekovia accessoria n. gen. n. sp.3
Kubekovia accessoria n. gen. n. sp.3 Kovalev 1985 snipe fly
PIN 1255/269
 Diptera - Mythicomyiidae
Palaeoplatypygus zaitzevi n. gen. n. sp.3
Palaeoplatypygus zaitzevi n. gen. n. sp.3 Kovalev 1985 fly
PIN 1255/268
 Diptera - Pleciofungivoridae
Pleciofungivora yeniseyica n. sp.3 Kovalev 1985 fungus gnat
PIN 1255/242
Bryanka elegantissima n. sp.3 Kovalev 1985 fungus gnat
PIN 1255/256
Krasnoyarskia subcosta n. gen. n. sp.3 Kovalev 1985 fungus gnat
PIN 1255/247, 250
Willihennigia variabilis n. sp.3 Kovalev 1985 fungus gnat
PIN 1255/246, 234, 240, 244, 245, 251
Willihennigia latipennis n. sp.3 Kovalev 1985 fungus gnat
PIN 1255/249
Eohesperinus sibiricus n. sp.3 Kovalev 1985 fungus gnat
PIN 1255/314
 Diptera - Archizelmiridae
Archizelmira jarzembowskii n. sp.6 Lukashevich 2022 fungus gnat
PIN 1255/2218
 Diptera - Protorhyphidae
Protorhyphus sibiricus n. sp.3 Kovalev 1985 fly
PIN 1255/265
 Diptera - Siberhyphidae
Siberhyphus lebedevi n. gen. n. sp.3
Siberhyphus lebedevi n. gen. n. sp.3 Kovalev 1985 fly
PIN 1255/264
 Diptera - Perissommatidae
Palaeoperissomma collessi n. gen. n. sp.3
Palaeoperissomma collessi n. gen. n. sp.3 Kovalev 1985 fly
PIN 1255/261; 1255/1341 in Lukashevich & Blagoderov 2020
 Diptera - Dixidae
Eucorethrina flexa n. gen. n. sp.5, Syndixa mollis n. sp.5, Syndixa sibirica n. gen. n. sp.5
Eucorethrina flexa n. gen. n. sp.5 Kalugina 1985 meniscus midge
PIN 1255/1546, 1547 (3 wings), 1545, 1553, 1554, 234 (4 small wings)
Syndixa mollis n. sp.5 Lukashevich 1996 meniscus midge
PIN 1255/1548
Syndixa sibirica n. gen. n. sp.5 Lukashevich 1996 meniscus midge
PIN 1255/1549-1552, 226 (formerly paratype of E. flexa)
 Diptera - Chaoboridae
Helokrenia nana n. gen. n. sp.1, Chaoborites argillaceus n. sp.1
Helokrenia nana n. gen. n. sp.1 Kalugina 1985 phantom midge
PIN 1255/207, 206, 208
Chaoborites argillaceus n. sp.1 Kalugina 1985 phantom midge
PIN 1255/220