PBDB Collection

Khara Khutul (PIN coll. 5268) [MPE/SMPE] (Cretaceous to of Mongolia)

Also known as Khara Khutul Ula; Khar Hötöl Uul; Kharakhutulia kalandadzei type

Where: Dornogov, Mongolia (44.5° N, 107.4° E: paleocoordinates 46.2° N, 97.5° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Baynshire Formation, Cenomanian to Cenomanian (100.5 - 83.6 Ma)

• = Bainshirenskaya, Bayn Shire, Bainshire Formation

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size class: macrofossils

Collected by N. N. Kalandadze, MPE, SMPE, SMGE in 1946, 1948–1949, 1967–1972, 1974, 1976; reposited in the PIN

• Soviet-Mongolian Paleontological Expedition

Primary reference: I. A. Efremov. 1954. Paleontologeskiye issledovaniya v Mongol'skoy Narodnoy Respulike (predvaritel'nyye rezul'taty ekspeditsiyy 1946, 1948 i 1949 gg.) [Paleontological investigations in the Mongolian People's Republic (preliminary results of the expeditions of 1946, 1947, and 1949)]. Trudy Mongolʹskoĭ Komissii 59:3-32 [M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 118167: authorized by Philip Mannion, entered by Philip Mannion on 07.10.2011, edited by Bethany Allen and Matthew Carrano

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Coniferales - Cupressaceae	Taxodium sp.⁶ Taxodium sp.⁶ Richard 1810 cypress
Reptilia
Dinosauria -	Dinosauria indet.² Dinosauria indet.² Owen 1842 dinosaur eggshells
Avetheropoda -	Carnosauria indet.⁶ Carnosauria indet.⁶ carnosaur
Theropoda - Therizinosauridae	Enigmosaurus mongoliensis n. sp.³, Segnosaurus galbinensis¹ Enigmosaurus mongoliensis n. sp.³ Barsbold and Perle 1983 enigmatic lizard "relatively large pelvis"; PST GIN AN MNP 100/84 Segnosaurus galbinensis¹ Perle 1979 coelurosaur
Saurischia -	Sauropoda indet.⁶ Sauropoda indet.⁶ sauropod
Testudinata -	Testudinata indet.⁵ Testudinata indet.⁵ Oppel 1811 turtle
Testudines - Sinemydidae	Sinemydidae indet.⁷ Sinemydidae indet.⁷ Yeh 1963 turtle
Testudines - Carettochelyidae	Kizylkumemys schultzi⁸ Kizylkumemys schultzi⁸ Nessov 1977 turtle Listed as Kizylkumemys schultzi mirabilis (Nessov 1981)
Testudines - Trionychidae	Trionychidae indet.⁸ Trionychidae indet.⁸ Gray 1825 softshell turtle
Testudines - Lindholmemydidae	Mongolemys sp.⁷ Mongolemys sp.⁷ Khosatzky and Mlynarski 1971 turtle
Testudines - Nanhsiungchelyidae	Nanhsiungchelyidae indet.⁸, Nanhsiungchelyidae indet.⁴, Kharakhutulia kalandadzei n. gen. n. sp.⁸, Hanbogdemys sp.⁸ Nanhsiungchelyidae indet.⁸ Yeh 1966 turtle Originally listed as ?Zangerlia sp. (Shuvalov and Chkhikvadze 1975) Nanhsiungchelyidae indet.⁴ Yeh 1966 turtle IP 11-2-2 (fragmentary shell and postcranial remains) Kharakhutulia kalandadzei n. gen. n. sp.⁸ Sukhanov et al. 2008 turtle PIN 5268-1 - holotype (a partial shell missing its right and posterior periphery); paratypes: PIN 5268-2 (partial shell including left lateral fragment of the carapace and left half of the plastron), PIN 5268-3 (fragment of a carapace, including pygal and sutured fragments of peripherals 11), PIN 5268-4 (left bridge peripheral). PIN 5268-3 and PIN 5268-4 may belong to the same individual as PIN 5268-1 or PIN 5268-2. There are a number of other shell fragments belonging either to PIN 5268-1 or/and PIN 5268-2. Hanbogdemys sp.⁸ Sukhanov and Narmandakh 2006 turtle Originally described as an individual of Hanbogdemys ("Basilemys") orientalis (Nessov 1987)
Testudines - Adocidae	Adocidae indet.⁸ Adocidae indet.⁸ Cope 1869 turtle

¹Barsbold 1983; ²Carpenter and Alf 1994; ³Currie 2000; ⁴Danilov and Syromyatnikova 2008; ⁵Joyce and Bourque 2016; ⁶Kalandadze and Kurzanov 1974; ⁷Shuvalov 1974; ⁸Sukhanov et al. 2008